421 research outputs found
FIGURE 2. Sarcoglottis matogrossensis Engels & E.C.Smidt.—A in A new Sarcoglottis (Orchidaceae: Spiranthinae) from the southern Brazilian Amazon
No full textFIGURE 2. Sarcoglottis matogrossensis Engels & E.C.Smidt.—A. Habit of three leafless flowering individuals. B. Inflorescence. C. Flower in frontal view. D. Flower in oblique view. E. Flower in lateral view. F. Habit of leafy individuals, before flowering. G. Leaf adaxial and abaxial surfaces. Photos by M.E. Engels based on the type.Published as part of Engels, Mathias Erich & Smidt, Eric De Camargo, 2021, A new Sarcoglottis (Orchidaceae: Spiranthinae) from the southern Brazilian Amazon, pp. 171-176 in Phytotaxa 487 (2) on page 174, DOI: 10.11646/phytotaxa.487.2.8, http://zenodo.org/record/575755
Microchilus dasilvae: a new species of Goodyerinae (Orchidaceae) from Southeastern Brazil
No full textEngels, Mathias E., Smidt, Eric C. (2022): Microchilus dasilvae: a new species of Goodyerinae (Orchidaceae) from Southeastern Brazil. Phytotaxa 555 (3): 252-258, DOI: 10.11646/phytotaxa.555.3.4, URL: http://dx.doi.org/10.11646/phytotaxa.555.3.
FIGURE 1. Microchilus dasilvae.—A. Habit. B. Flower bract. C in Microchilus dasilvae: a new species of Goodyerinae (Orchidaceae) from Southeastern Brazil
No full textFIGURE 1. Microchilus dasilvae.—A. Habit. B. Flower bract. C. Trichome of inflorescence. D-E. Flowers. D. Lateral view. E. Frontal view. F. Dorsal sepal. G. Petal. H. Lateral sepal. I-J. Lip. I. Flattened. J. Lateral view. K. Detail of the apex of the lateral lobe. L-N. Column. L. Dorsal view. M. Ventral view. N. Detail of the rostellum remain. O. Pollinarium. Drawing by Engels from M.E.Engels 9306.Published as part of Engels, Mathias E. & Smidt, Eric C., 2022, Microchilus dasilvae: a new species of Goodyerinae (Orchidaceae) from Southeastern Brazil, pp. 252-258 in Phytotaxa 555 (3) on page 254, DOI: 10.11646/phytotaxa.555.3.4, http://zenodo.org/record/691110
FIGURE 1 in Taxonomic and nomenclatural notes in Microchilus longicornu (Goodyerinae Orchidaceae)
No full textFIGURE 1. Lectotype of Microchilus longicornu (Riedel 77 at LE [barcode LE00001368]). Courtesy of the Herbarium LE curators.Published as part of Engels, Mathias E. & Smidt, Eric C., 2022, Taxonomic and nomenclatural notes in Microchilus longicornu (Goodyerinae Orchidaceae), pp. 296-300 in Phytotaxa 568 (3) on page 298, DOI: 10.11646/phytotaxa.568.3.7, http://zenodo.org/record/722365
Microchilus dasilvae Engels & E. C. Smidt
No full textMicrochilus dasilvae Engels & E.C.Smidt sp nov. (Figures 1–2) Type: — BRAZIL. São Paulo: Ibiúna. Vegetative material received from Marcos Aurélio da Silva and cultivated until flowering, fl., 24 September 2021, M.E.Engels 9306 (holotype MBM!; isotypes HCF!; RB!; SP!; UPCB!). Similar to Microchilus decorus and Microchilus metallescens, differs in lip with trilobed mesochile, yellow, with acute lateral lobes; and by the broad-ovate epichile, about the same width as the mesochile when flattened. Description:— Humicolous herb, up to 30-35 cm tall. Roots 6–16 × 1–2 mm, cylindrical, hairy, white. Rootbearing portion of stem 5–8 × 2–4 mm, cylindrical, green; internodes 1.5–2.1 cm long, one root per internodes; foliate portion of stem 11.5–13.2 × 0.3–0.5 cm, cylindrical, 6–9 foliate, green; internodes (0.7)– 1–2.4 cm long. Leaves petiolate, lanceolate to narrow-lanceolate; petiole + sheath 1.1–1.7 × 0.8–1.2 cm amplective, pale-green; blade 2.0-4.2 × 0.7-1.9 cm, dark-green, discolourous, base decurrent, margin entire and smooth, acute. Inflorescence simple, erect, racemose, (3)–19–27–flora; peduncle 11.1–12.7 × 0.1–0.2 cm, pubescent, greenish-brown; peduncle bracts 0.9–1.3 × 0.2–0.6 cm, lanceolate, base acute or amplective, margin entire to ciliate, acute, brown, in the basal portion of peduncle amplective and the distal portion free; rachis 3.5–5.8 × 0.1–0.15 mm, pubescent, green; floral bracts 5–9 × 1–3 mm, lanceolate, acute, margin entire and ciliate, acute, light-brown. Flowers with ovary + pedicel ca. 7 × 1.5 mm, sub-oblanceoloid, hairy, green; dorsal sepal ca. 4 × 2 mm, oval, base cuneate to acute, margin entire, apex rounded, abaxial face hairy, white with central longitudinal bilinear brown macula; lateral sepals ca 4 × 2 mm, elliptical, slightly asymmetric, acute base, entire margin, obtuse-rounded apex, hairy abaxial face, white with bilinear central longitudinal macula; petals ca. 4 × 1 mm, narrow-elliptical, sub-falcate, slightly asymmetric, base acute, margin entire, apex obtuse, glabrous, white; lip 13–13.5 × 2 mm, subdivided; hypochile ca. 7 × 1 mm, narrow-ellipsoid, cylindrical, apex rounded, glabrous, calcariform, white; mesochile ca. 4 × 2 mm, obovate, margin entire, yellow, lateral lobes acute; epichile ca. 2–2.5 × 2 mm, broad-ovate, base acute, margin entire, apex obtuse and apiculate, white. Column ca. 4 × 1.25–1.5mm; stigma ca. 0.3 × 1 mm, obsemilunate, bilobed; rostellum ca. 2 mm long, deltoid and long narrow apiculate, acute; rostellar remnant with truncated scar; anther ca 1 × 0.5 mm, elliptical-ovate, brown, apiculate; pollinia 2, ca. 1 mm long, obclaviform, slightly flattened, longitudinally grooved, white. Fruits and seeds not seen. Distribution and ecology:— Occurs as humicolous in the forest understory, in an ecotonal region between Ombrophilous Dense Forest and Semideciduous Seasonal Forest, in the Atlantic Forest Domain. It flowered in cultivation between September and October. Etymology:— The specific epithet is in honour of the discoverer of a new species in the field, a Brazilian Orchidologist Marcos Aurélio da Silva. Additional specimen examined (paratype):— BRAZIL. São Paulo: Ibiúna. Material cultivated until flowering and kindly provided by Marcos Aurélio da Silva, fl., 05 October 2020, M.E.Engels 8231 (MBM!; RB!, UPCB [spirit]!). Conservation status:— According to IUCN criteria (2017), M. dasilvae falls under Data Deficient (DD). More studies are necessary to locate more populations and estimate the number of mature individuals. The area where the species was discovered suffers from anthropological disturbance, including real estate pressure, making the species naturally threatened due to habitat loss. Taxonomic discussion:— Microchilus dasilvae is similar to M. decorus (Reichenbach 1873: 177) Smidt et al. (2021: 339) (Table 1) due to the habit (small plants, up to 35 cm tall, and leaves up to about 6–7.5 cm in length) and floral display with stained sepals and mesochile with acute lateral lobes. It can be distinguished by the yellow (vs. white) mesochile and the broad-oval epichile about the same width as mesochile (vs. oblanceolate to narrow elliptical, proportionately narrower than mesochile). The complete illustration of M. decorus can be consulted in Sprunger et al. (1996: t. 136), as Physurus bicolor Barbosa Rodrigues (1882: 290). Additionally, M. dasilvae is similar to M. metallescens (Barbosa Rodrigues 1882: 291) Smidt et al. (2021: 342) (Table 1) due to the habit and floral display with stained sepals and white lip with yellow mesochile. It can be distinguished from this one by the narrow-elliptical and sub-falcated petals, slightly asymmetrical (vs. ovate and symmetrical); lip with mesochile with acute lateral lobes (vs. rounded); broad-ovate epichile with apiculate and obtuse apex (vs. oblong, non-apiculate obtuse apex); sepals with bilinear central longitudinal macula (vs. longitudinally trilinear macula). The complete illustration of M. metallescens can be consulted in Sprunger et al. (1996: 135) and Engels et al. (2016: 940).Published as part of Engels, Mathias E. & Smidt, Eric C., 2022, Microchilus dasilvae: a new species of Goodyerinae (Orchidaceae) from Southeastern Brazil, pp. 252-258 in Phytotaxa 555 (3) on pages 253-256, DOI: 10.11646/phytotaxa.555.3.4, http://zenodo.org/record/691110
Sarcoglottis matogrossensis Engels & E. C. Smidt 2021, sp. nov.
No full textSarcoglottis matogrossensis Engels & E.C.Smidt sp. nov. (Figs. 1–2) Type:— BRAZIL. Mato Grosso: Nova Canaã do Norte, Teles Pires River, UHE Colíder, 18 August 2018, [fl.], M . E. Engels, T. A. Silva & W. Monteiro 6617 (holotype MBM, isotype RB). Diagnosis:— The new species is morphologically similar to S. metallica but can be distinguished by the discolored-green leaves, that may or may not be variegated with white spots, flowering that occurs when the leaves have become senescent, and petals with apices conspicuously free, acute, and reflexed. Description:— Humicolous and saxicolous, acaulescent herbs. Roots 2–5 mm long, appearing fasciculate, thick, cylindrical, hairy. Leaves 13.6–22.2 × 2.4–5.2 cm, narrowly elliptical to oblanceolate, discoloured-green, that may or may not be variegated with white spots, attenuating towards the base, margin entire, apex obtuse to acute. Inflorescence up to ca. 50 cm long, racemose, erect, 1–8-flowered; peduncle 19.2–40.9 × 0.3–0.5 cm, cylindrical, pubescent, greenish-brown; rachis 4.5–10.2 × 0.2–0.3 cm, cylindrical, pubescent, greenish-brown to brown; peduncle bracts 3.2–4.4 × 0.2–0.6 cm, narrowly triangular to narrowly elliptical, glabrous, greenish-brown, entire margin, acute apex; floral bracts 1.5–2.6 × 0.2–0.3 cm, narrowly lanceolate, glabrous, greenish-brown, entire margin, acute apex. Flowers resupinated; pedicellate ovary 2.8–3.2 × 0.4–0.6 cm, pubescent, green; dorsal sepal 1.4–1.7 × 0.3–0.4 cm, elliptical, concave, cuneate at base, margin entire, acute and recurved apex, pubescent adaxial face, light green with three darkergreen veins; lateral sepals 3.2–3.8 × 0.4–0.5 cm, falcate, decurrent in the ventral portion of the ovary, approximately 1/3 basal-connate, light green, pubescent adaxial face, portion free lanceolate, entire margin, acute apex; petals 1.2–1.4 × 0.2 cm, sub-sigmoid, elliptical in the general view, proximal margin internally adnate to the dorsal sepal in ca. 2/3 of the length, free proximal margin ciliated in ca. 2/3 of the length, acute apex, curved in a natural position, light green with three darker-green veins; lip 3.0–3.4 × 0.5–0.7 cm, spatulate in overview; narrowly oblanceolate, greenish-white; nectar glands ca. 0.5 cm long., narrowly ellipsoids, light green; apical lobe 4–5 × 5–7 mm, broadly oval, rounded base, entire margin, obtuse and apiculate apex, light green to yellowish-green with darker-green veins. Column 1–1.2 × 0.3 cm, flattened ventrally, glabrous, whitish; rostellum ca. 2 × 2 mm, sub-deltoid, truncated. Anther ca. 4 × 2 mm, elliptical, brownish-white. Stigma bilobed; elliptical, and confluent lobes; ca. 2 × 1 mm, whitish. Pollinarium with two elliptical-oblanceolate, whitish pollinia, ca. 4–5 × 1 mm; viscidium ca. 1 × 1 mm, transversely ellipsoid, dark brown. Fruits not seen. Additional material examined (Paratypes):— BRAZIL. Mato Grosso: Colíder, 17 August 2017, [fl.], M . E. Engels 5666 (MBM). Nova Canaã do Norte, Teles Pires River, UHE Colíder, 03 August 2019, [fl.], M . E. Engels 7338 (MBM). Discussion: — The architecture of the inflorescence, shape of the leaves and floral morphology of S. matogrossensis are reminiscent of S. maroaënsis (Romero-González et al. 2010: 514), S. amazonica and S. metallica. However, it can be distinguished by the light green, discoloured leaves that may or may not be variegated with white spots (vs. reddish, brownish, or chocolate-purplish with irregular, bright green spots on the adaxial surface and homogeneously purplishred on the abaxial surface); the flowering which occurs after the senescence of the leaves (vs. fresh leaves present at flowering); and by the petals with apices conspicuously free, acute and reflexed (vs. adhering to the dorsal sepal nearly to the apex, obtuse, and straight). From S. maroaënsis specifically, it can also be distinguished by the glabrous lip (vs. rugose thickening near the apex of the apical lobe and extending to the apex of the central portion of the lip) and the apical lobe with an entire margin (vs. crenate and plicate). With the recently described S. neillii (Salazar et al. 2019), S. matogrossensis shares the floral structure and predominantly green leaves. The latter can be easily distinguished by the narrowly elliptical to oblanceolate leaves (vs. elliptical to ovate) that are proportionally larger (13.6–22.2 vs. 5−13 cm long.), by the petals with apices conspicuously free, acute, and reflexed (vs. adherent internally to the dorsal sepal except at their little free recurved apices, obtuse and sometimes obscurely apiculate); and flowering occurs after the senescence of the leaves (vs. fresh leaves present at flowering). Distribution, habitat, and phenology:— Sarcoglottis matogrossensis occurs as a saxicolous or humicolous plant in the stony soil and between the rocky outcrops of the understory of seasonal evergreen forest, in the Amazon Biome of the north-central region of the Mato Grosso state, at around 200–300 m of elevation. Flowering occurs during the dry season, in August and September. Etymology:—The specific epithet refers to the Mato Grosso state, where the species was discovered. Conservation status:—According to the IUCN (2017) criteria, Sarcoglottis matogrossensis qualifies as Data Deficient (DD). The two localities, where the new species was found, are separated by only ca. 30 km. The habitat is highly fragmented, and the two populations consist of less than ten individuals each. Deforestation and climate change are a major threat to Amazonian Biome (Gomes et al. 2019). This is especially problematic in the Mato Grosso state, where the rate of illegal deforestation is very high. Recent reports suggest that between August 2016 and July 2017, illegal logging affected more than 60.000 hectares of forests (Silgueiro et al. 2019). There are no studies of the impact on the herbaceous diversity, but without a forest canopy, the understory plants are likely threatened. Due to the low botanical coverage in the Mato Grosso state, further fieldwork is required to verify the presence of other populations of this species.Published as part of Engels, Mathias Erich & Smidt, Eric De Camargo, 2021, A new Sarcoglottis (Orchidaceae: Spiranthinae) from the southern Brazilian Amazon, pp. 171-176 in Phytotaxa 487 (2) on pages 172-175, DOI: 10.11646/phytotaxa.487.2.8, http://zenodo.org/record/575755
FIGURE 1. Thismia mantiqueirensis Engels & E.C in Thismia mantiqueirensis (Thismiaceae), a new species of Brazilian Atlantic Rainforest
No full textFIGURE 1. Thismia mantiqueirensis Engels & E.C.Smidt sp. nov. A. Habit. B. Detail of stem in transversal section. C. Leaf. D. Flower in sagittal section. E. Detail of stigma. F. Frontal view of distended tepals and upper disc of the floral tube. G. Detail of tepal lamellae. H. Detail of trichomes from abaxial surface of tepals. I. Detail of trichomes of the adaxial surface of tepals. J. Anther in frontal view. K. Anther in lateral view. L. Mature fruit and leaves in lateral view. M. Mature fruit in dorsal view. Drawn by M.E. Engels based on the type.Published as part of Engels, Mathias Erich, Muscat, Edelcio, Moroti, Matheus De Toledo & Smidt, Eric De Camargo, 2022, Thismia mantiqueirensis (Thismiaceae), a new species of Brazilian Atlantic Rainforest, pp. 223-228 in Phytotaxa 570 (2) on page 225, DOI: 10.11646/phytotaxa.570.2.8, http://zenodo.org/record/725668
FIGURE 2. A–C in Taxonomic and nomenclatural notes in Microchilus longicornu (Goodyerinae Orchidaceae)
No full textFIGURE 2. A–C. Microchilus foliosus (M.E.Engels 8218 at MBM). A. Habit in the natural environment. B. Inflorescence. C. Detail of flowers in an inflorescence. D–F. Microchilus longicornu (M.E.Engels 6618 et al. at MBM). D. Habit. E. Inflorescence, arrow indicates the long hypochile. F. Detail of flowers in an inflorescence. Photos by M.E.Engels.Published as part of Engels, Mathias E. & Smidt, Eric C., 2022, Taxonomic and nomenclatural notes in Microchilus longicornu (Goodyerinae Orchidaceae), pp. 296-300 in Phytotaxa 568 (3) on page 299, DOI: 10.11646/phytotaxa.568.3.7, http://zenodo.org/record/722365
FIGURE 3. Arrojadoa flava. A–E. Habit and environments. F. Flowers detail. G in Arrojadoa flava (Cactaceae, Cactoideae), a new yellow flowered species from the Brazilian flora
No full textFIGURE 3. Arrojadoa flava. A–E. Habit and environments. F. Flowers detail. G. Flowers, immature and mature fruits. Photos by M.E.Engels from type population.Published as part of Gonzaga, Diego Rafael, Liebsch, Dieter & Engels, Mathias Erich, 2023, Arrojadoa flava (Cactaceae, Cactoideae), a new yellow flowered species from the Brazilian flora, pp. 73-79 in Phytotaxa 597 (1) on page 77, DOI: 10.11646/phytotaxa.597.1.8, http://zenodo.org/record/791883
Eltroplectris paranaensis Engels & E. C. Smidt 2023, sp. nov.
No full textEltroplectris paranaënsis Engels & E.C.Smidt, sp. nov. (Figures 2–3) Type: — BRAZIL. Paraná: Campina Grande do Sul, Serra do Mar, fl., 27 April 2022, M. E . Engels, L. Passos, T. F. Santos & M. Klingelfus 9669 (holotype UPCB!). Similar to E. janeirensis (Porto & Brade 1940: 33) Pabst (1974: 469) but differs from this by the narrow elliptical-lanceolate leaves; pauciflorous inflorescence; externally glabrous flowers; oblanceolate petals and central lobe of the lip with entire margins. Description:— Humicolous acaulescent herbs, 26–27 cm tall. Roots 2.7–4.8 × 0.3–0.4 cm, appearing fasciculate, cylindrical, pilose. Leaves pseudo-petiolate, rosulate; pseudopetiole 3.4–4.5 × 0.2–0.3 cm, conduplicate; blade 8.5– 10.2 × 1.4– 1.3 cm narrowly elliptical-lanceolate, dark green, discolored, base acute and decurrent, margin entire, apex acute. Inflorescence racemose, erect, 1–2-flowered; peduncle 23.9–24 × 0.15–0.2 cm, cylindrical, pubescent in apical portion, green; rachis ca. 1.6 × 0.1 cm, cylindrical, pubescent, green; peduncle bracts 2.1–2.7 × 0.4–0.5 cm, amplective, narrowly triangular to narrowly lanceolate, glabrous, green, margin entire, apex acute; floral bracts 1.5–1.6 × 0.2– 0.4 cm, elliptical-lanceolate, glabrous, green, base acute, margin entire, apex acute. Flowers resupinate; pedicellate ovary 2.3–2.5 × 0.3 cm, glabrous, light-green; dorsal sepal ca. 1.7 × 0.7 cm, lanceolate, concave, base acute, margin entire, apex acute, glabrous, light green; lateral sepals ca. 2.0 × 0.5 cm (free portion, excluding the calcar), narrowly lanceolate, slightly asymmetrical, base long decurrent on the ventral portion of the ovary and forming a calcar, light-green, glabrous, margin entire, apex acute; calcar 1.5–1.7 × 0.2 cm, cylindrical, free, white, formed from the free part of the column foot, base of the lateral sepals and base of the lip; petals ca. 1.5 × 0.6 cm, asymmetrical, oblanceolate, base acute, margin entire, apex obtuse, white with light green apex; lip ca. 2.5 × 0.75 cm, unguiculate, white with light green apex; unguicule ca. 9 × 3.5 mm, oblong; blade trilobed; medial portion ca. 9 × 7.5 mm, oblanceolate; lateral lobes rounded; apical lobe ca. 7 × 4 mm, oval-lanceolate, base rounded-obtuse, margin entire, apex acute. Column 1 × 0.2 cm (excluding column foot), flattened ventrally, above rostrate and cucullate, whitish; column foot 1.5–1.7 cm long, flattened, with oblanceoloid unicellular glandular trichomes on the ventral side; rostellum ca. 4 mm long, long narrowly apiculate, acute. Anther ca. 4 × 2 mm, narrowly lanceolate, base cordate, margin sinuate, apex acute, brownish. Stigma bilobed, lanceolate, confluent at the apex, ca. 2 × 1 mm, white. Pollinarium with two narrowly oblanceolate, whitish pollinia, ca. 5 mm long; viscidium ca. 0.7 mm long, deltoid, apex sub-rounded, whitish hyaline. Fruits not seen. Distribution and ecology: —Known only by the population in the type locality, where it occurs as a humicolous in the understory litter of the Upper Montane Dense Ombrophilous Forest at an elevation of 1200 m. Found flowering in April, during autumn in the Southern Hemisphere. Etymology: —The specific epithet refers to the Paraná State, where the species was discovered. Taxonomic discussion: —Among the congeners, E. paranaënsis is morphologically related to E. janeirensis and E. schlechteriana (Porto & Brade 1940: 32) Pabst (1974: 469), due to the presence of leaves at anthesis; light-green and white flowers; floral display with wide, open petals; trilobed lip and the shape of the central lobe (i.e. deltoid, ovate to lanceolate and not fimbriate). Eltroplectris paranaënsis can be distinguished from E. janeirensis by its smaller habit (26–27 cm total height vs. 75–80 cm); narrow, smaller leaves (narrowly elliptical-lanceolate, base acute, 8.5–10.2 × 1.4– 1.3 cm vs. ovate-subrounded, base rounded, ca. 15 × 10.5 cm), ovate petals (vs. oblanceolate); lip with apical lobe proportionally shorter than the median portion (1/3 vs. 1/1) and lip with smooth apical lobe margin (vs. little denticulation). It differs from E. schlechteriana in its smaller habit (26–27 cm total height vs. 50–70 cm); narrow, smaller leaves (narrowly elliptical-lanceolate, base acute, 8.5–10.2 × 1.4– 1.3 cm vs. ovate to ovate-lanceolate, base rounded, ca. 13–20 × 6–9 cm); lip with apical lobe proportionally shorter than the median portion (1/3 vs. 1/1) and lip with smooth apical lobe margin (vs. little denticulation). Additionally, E. paranaënsis is similar to E. assumpcaoana Campacci & Kautsky (1999: 108) and E. kuhlmanniana (Hoehne 1944: 133) Szlachetko & Rutkowski (2008: 164) due to the presence of leaves at anthesis and the habit with relatively narrow leaves (i.e., narrowly elliptical-lanceolate, lanceolate or oblong-lanceolate). However, E. paranaënsis can be distinguished from E. assumpcaoana by the short pseudopetiole (3.4–4.5 cm long vs. 12–16 cm long), number of flowers per inflorescence (1–2 flowers vs. ca. 9 flowers), flower coloration (light green and white vs. brown and white), lip with apical lobe proportionally shorter than the median portion (1/3 vs. ca. 1/1) and by the shape of the apical lobe (oval-lanceolate vs. large-linear). Eltroplectris paranaënsis differs from E. kuhlmanniana by the short pseudopetiole (3.4–4.5 cm long vs. 10.5–17 cm long), reduced number of flowers (1–2 vs. ca. 9), by the length of the calcar (proportionately shorter than the ovary + pedicel vs. the same length to longer) and by the shape of the apical lobe of the lip (oval-lanceolate vs. subovate-obcordate).Published as part of Engels, Mathias Erich, Silva-Pereira, Viviane Da & Smidt, Eric De Camargo, 2023, Eltroplectris paranaënsis: a new species of Spiranthinae (Orchidaceae) from the nebular Atlantic Rain Forest, pp. 115-123 in Phytotaxa 609 (2) on pages 117-120, DOI: 10.11646/phytotaxa.609.2.3, http://zenodo.org/record/827229
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