21,449 research outputs found
Tricorythodes biluhue Emmerich & Molineri 2020, sp. nov.
No full textTricorythodes biluhue sp. nov. (Figures 1–28) Type material. Holotype (MUR) female mature nymph from Uruguay, Tacuarembó, A° Tranqueras, S 31° 47’ 29” – W 55° 58’ 51”, 22/II/2008, Emmerich & Molineri cols. Paratypes (IBN): 10 nymphs, same data as holotype. One nymph, Uruguay, Salto, Arroyo (abbreviation = A°) Itapebí, finca Iribarne, S 31º 22’ 3.4” – W 57º 32’ 24.5”, 102 m, 17/II/2008, Emmerich & Molineri cols. 16 nymphs, Uruguay, Artigas, Aº de los Chanchos, S 30º 34’ 48.3” – W 56º 37’ 10.8”, 130 m, 18/II/2008, Emmerich & Molineri cols. Three nymphs, Uruguay, Artigas, Finca Mataojo Grande, Aº Mataojo Grande, 274 m, S 31º 17’ 29.7” – W 56º 13’ 10”, 18/II/2008, Emmerich & Molineri cols. Two nymphs, Uruguay, Rivera, Tranqueras, río Tacuarembó, 147 m, S 31º 10’ 40” – W 55º 45’ 44”, 19/II/2008, Emmerich & Molineri cols. Two nymphs, Uruguay, Rivera, Santa Ernestina, A° sin nombre, 146 m, S 31º 32’ 23.8” – W 55º 33’ 42”, 19/II/2008, Emmerich & Molineri cols. Two nymphs, Uruguay, Rivera, Mina de Corrales, Aº Corrales, 140 m, S 31º 34’ 34” – W 55º 27’ 58”, 19/II/2008, Emmerich & Molineri cols. Three nymphs, Uruguay, Rivera, Aº Zapucay, 140 m, S 31º 47’ 01” – W 55º 20’ 02”, 19/II/2008, Emmerich & Molineri cols. Two nymphs, Uruguay, Tacuarembó, A° Carpintería, 121 m, S 31º 48’ 28” – W 55º 13’ 09”, 20/II/2008, Emmerich & Molineri cols. Six nymphs, Uruguay, Cerro Largo, Aº Tacuarí, 92 m, S 32º 31’ 51” – W 54º 07’ 48”, 20/II/2008, Emmerich & Molineri cols. One nymph, Uruguay, Durazno, Sarandí del Yí, río Yí, 132 m, S 33° 20’ 41” – W 55° 37’ 07”, 22/II/2008, Emmerich & Molineri cols. Two nymphs, Uruguay, Tacuarembó, Valle Edén, A° Tambores, S 31° 48’ 48” – W 56° 10’ 24”, 160 m, 23/ II/2008, Emmerich & Molineri cols. Six nymphs, Uruguay, Paysandú, A° Soto, 43m, S 32° 03’ 10” – W57° 40’ 23”, 23/II/2008, Emmerich & Molineri cols. One nymph, Uruguay, Paysandú, A° Guayabos Grande, 83 m, S 32° 22’ 45” – W 57° 20’ 11”, 1/III/08, Emmerich & Pérez, cols. 15 nymphs, Uruguay, Río Negro, A° Don Esteban, 34 m, S 32° 50’ 07” – W 57° 30’ 59”, 2/III/2008, Emmerich & Pérez cols. Seven nymphs, Uruguay, Flores, Camino vecinal, A° sin nombre, 115 m, S 33° 42’ 52” – W 56° 39’ 40”, 2/III/2008, Emmerich & Pérez cols. Three nymphs, Uruguay, Lavalleja, Minas, Cañada Solís, 123 m, S 34° 24’ 08” – W 55° 21’ 11”, 3/III/2008, Emmerich & Pérez cols. Six male imagoes, one female imago, Uruguay, Maldonado, A° El Rodeo, 75 m, S 34° 40’ 20” – W 55° 14’ 15”, 7–8/XI/2008, Emmerich col. Two nymphs, Uruguay, Paysandú, A° Sacra, 10 m, S 32° 21’ 02” – W 58° 03’ 04”, 7/III/2008, Emmerich & Pérez cols. Two nymphs, Uruguay, San José, Aº Pavón, 86 m, S 34° 18’ 53.2” – W 56º 52’ 24.7”, 6/XI/2008, Emmerich col. Two nymphs, Uruguay, Rivera, Cañada sin nombre, 289 m, S 31º 05’ 36.7” – W 55º 57’ 57.9”, 11/ XI/2008, Emmerich col. Three nymphs, Uruguay, Tacuarembó, A° Potrero, 157 m, S 31º 54’ 13.9” – W 56º 46’ 34.9”, 26/III/2010, Emmerich & Molineri cols. One nymph. Uruguay, Río Negro, ”El Chicharrón”, A° Grande, S 32º 31’ 45” – W 57º 08’ 11.9”, 9/8/07. E. Morelli & A. Verdi cols. Diagnosis. The nymphs of Tricorythodes biluhue sp. nov. can be distinguished from all other species of the genus by the following combination of characters: 1) maxillary palp 2 to 3–segmented (Figs. 6–7); 2) hypopharynx with spine on inner margin of superlinguae (Fig. 9); 3) pronotum with pointed anterolateral projection and small blunt tubercle medially; 4) legs relatively long and slender, covered with long setae (Figs. 11, 13 and 14); 5) fore tarsal claws with 10–13 marginal denticles and with 0–1 + 1 submarginal denticles (Fig. 12); 6) abdominal segments with lateral flanges on III–VI and small posterolateral spines on VI–IX (Fig. 17); 7) operculate gills subtriangular, shaded widely except on paler area close to insertion (Fig. 18); 8) gills III–V with relatively wide ventralmost lamellae. The adults are characterized by: 1) vein CuP present and complete; 2) the ratio length of forceps segment I/forceps segment II 0.8; 3) penes subpyramidal, with a small apical furrow and with sclerotized yellowish lateral margins (Fig. 24). The eggs are characterized by: 1) pear–shaped with one polar cap present (Fig. 28). Descriptions. Mature nymph (Figs. 1–23). Length (mm): body, 4.2–4.8, caudal filament, 3.0–3.7, cerci, 2.2–2.6. General coloration yellowish gray with black markings (Fig. 2). Head shaded with gray mainly on occiput. Mouthparts: labrum with relatively shallow anteromedian emargination (Fig. 3) with marginal row of stout bifid setae ventrally, and with pectinate setae dorsally as in figures 4 and 5; mandibles with setae on outer margin; hypopharynx with medially concave lingua, superlingua with a spine on inner margin at proximal third (Fig. 9); maxillary palp small but elongated, 2 or 3 segmented with apical seta (Figs. 6–8), distal brush of maxilla with 25–35 long and curved setae, with stipes 1.0 the length of galea–lacinia, inner proximal margin of the galea–lacinia (opposite to palpus) with row of 3–4 simple setae and one large pectinate seta (Figs. 6 and 7); labium with paraglossae and glossae with the same length, lateral margin of prementum with 19–32 long and thin setae (Fig. 10). Thorax. Pronotum yellowish shaded with gray, with small blunt tubercle medially (Fig. 15) and pointed anterolateral projection (Fig. 16). Mesonotum yellowish shaded extensively with gray, darker on anterolateral corners; wingbuds yellowish with black veins; small anteromedian blunt tubercle. Metanotum and thoracic sterna paler, not shaded. Legs yellowish, shaded gray on femora and with a proximal and a distal macula on tibiae. Foreleg: transverse row of long setae (Fig. 11) located at 0.4–0.5 of total femoral length from base; total length (tl, same figure) / maximum width (mw) of femur = 2.4–3.1; fore tarsal claw with 10–13 marginal denticles and with 0–1 + 1 submarginal denticles (Fig. 12). Middle and hind legs (Figs. 13 and 14): femur covered with many long weak setae, some forming a proximal transversal row, other scattered over surface and margins; total length / maximum width of hind femur = 2.8–3.6; tarsal claw with 11–13 marginal denticles and 0–1 + 1 subapical sumarginal denticles. Abdomen. Yellowish shaded with gray widely,except on intersegmental membranes, darker on medial longitudinal line; small posterolateral spines present on segments VI–IX (smaller on IX), lateral flanges present on segments III–VI (Fig. 17). Gill formula 3/3/3/3/2. Operculate gills subtriangular, shaded widely with irregular gray markings except on a paler area near insertion (Fig. 18); gills III–VI with a pair of basal black marks, gills III–V with relatively wide ventral most lamellae (Figs. 20–23). Caudal filaments yellowish white shaded with gray on basal segments. Variations. The coloration is variable; some nymphs are pale with few dark markings, others are blackish or reddish (Figs. 1 and 2). Operculate gills also vary in coloration: some nymphs present a basal black band and the rest homogeneously shaded with gray, others nymphs show blackish operculum. Male imago (Figs. 24–27) Length (mm): Body, 4.8–5.0, Fore wing, 5.0–5.5, Caudal filaments, 13.0, Terminal filament, 14.0–18.5. General coloration gray brown. Head. Yellowish with gray shading on occiput. Pronotum. Medially whitish with longitudinal black stripe, meso– and metanotum brownish, mesopleura and meso- and metasternum brownish with lighter membranes (Figs. 25 and 26). Legs. Whitish, coxae, trochanters and femora yellowish brown; tibiae and tarsi much paler; blackish shading medially and subapically on femora; tibiae with a basal and distal black macula; inner face of tibiae II and III with a medial black stripe along patellotibial suture (Fig. 27). Fore wing. Translucent, veins shaded with gray, vein CuP complete. Abdomen. Brownish yellow with whitish intersegmental membranes black maculae present medially and also irregular gray shading on all terga. Sterna similar to terga with a medial black macula on each intersegmental membrane (Figs 25 and 26). Genitalia. Styliger plate whitish with brownish hind margin. Forceps translucent. Penes subpyramidal, whitish yellow shaded with brown as in figure 24. Caudal filaments whitish translucent shaded slightly with gray. Female imago. Length (mm): Body, 5.5, Forewing, 6.8. Coloration as in male except abdomen plenty of orangish eggs. Eggs. Pear–shaped with disc-like polar cap on thinner pole (Fig. 28). Distribution. T. biluhue sp. nov. has a wide distribution in Uruguay as shown in figure 29. Etymology. The epithet biluhue comes from the charrua language ”bilú hue” and means ”beautiful water”. We want to dedicate this species to the Uruguayan people and the Franciscan Family of this country who are demanding the repeal of the recently irrigation law that violates the human and biotic right to water.Published as part of Emmerich, Daniel & Molineri, Carlos, 2020, A new species of Tricorythodes Ulmer (Ephemeroptera: Leptohyphidae) from Uruguay, pp. 125-132 in Zootaxa 4885 (1) on pages 126-132, DOI: 10.11646/zootaxa.4885.1.8, http://zenodo.org/record/429644
Cloeodes guenoa Nieto & Emmerich, 2011, sp. nov.
No full textCloeodes guenoa sp. nov. (Figs. 2, 15– 25) Nymph (Fig. 2). Length: body, 4.9–5.1 mm; cerci, 1.4–1.5 mm; terminal filament, 1.2–1.3 mm. Antennae broken. Head yellowish brown, longer than wide, compound eyes yellowish brown. Antennae pale yellow. Mouthparts (Figs. 15–20): labrum (Fig. 15 a) with one subapical seta centrally and one seta near lateral margin, anterior margin with bifid setae (Fig. 15 b). Left mandible (Fig. 16) without setae between prostheca and mola, thumb of molar area transverse to anterior margin. Right mandible (Fig. 17) without setae between prostheca and mola. Hypopharynx (Fig. 18): lingua with rounded projection and longer than superlinguae. Maxillae (Fig. 19), palpi as long as galealacinia, two segmented, segment II with a constriction. Labium (Fig. 20) with segment III of palpi rounded. Thorax pale yellow. Pleura yellowish brown, sterna pale yellow. Legs yellowish brown. Dorsal margin of femora with a row of pointed spines, apically with a subquadrangular projection and two blunt spines (Fig. 21). Tarsal claws 0.3 times the length of tarsi. Hind wing pads present. Abdomen yellowish brown (Fig. 2), segments III-VII with two whitish spots near midline, segments IV and VIII pale brown. Abdominal terga with covers, posterior margin of terga with spines as in Fig. 22. Paraprocts with 10–11 spines apically (Fig. 23). Gills translucent white, main and secondary trachea pigmented (Fig. 24), 2 times the length of each tergum. Caudal filament yellowish, with flattened setae basally sclerotized (similar to Fig. 14 b). Cerci with long spines toward external margin every two segments, terminal filament with long spines toward midline every two segments (Fig. 25). Adults. Unknown. Etymology. “ Guenoa ” is the name of the native people who inhabits the current districts of Treinta y Tres, Cerro Largo and Tacuarembó. Diagnosis. Cloeodes guenoa sp. nov. can be distinguished from the other species of the genus by the following combination of characters: 1) hind wing pads present; 2) lingua with rounded projection and longer than superlinguae (Fig. 18); 3) labium with segment III of palpi rounded (Fig. 20); 4) dorsal margin of femora with a row of pointed spines, apically with a subquadrangular projection and two blunt spines (Fig. 21); 5) abdominal color pattern as in Fig. 2, with segments III-VII with two whitish spots near midline, segments IV and VIII pale brown. Material. Holotype female mature nymph, URUGUAY, Tacuarembó, R 5, km 379, Arroyo Tranqueras, S 31 º 47´29 ´´, W 55 º 58´51 ´´, 124m, 22 / 2 / 2008, Emmerich & Molineri colls. Paratypes six nymphs (one dissected) same data as holotype. Three nymphs: Lavalleja, R 8, km 190, Arroyo Tapes Chico, S 33 º 59´18 ´´, W 54 º 46´54 ´´, 130m, 4 / 3 / 2008, Emmerich & Pérez colls. One nymph: Treinta y Tres, Quebrada de los Cuervos, Arroyo Los Helechos, S 32 º 55 ’ 27.3 ’’, W 54 º 27 ’ 34.4 ’’, 125 m. 8 / 11 / 2004, D. Emmerich & E. Morelli colls. Six nymphs: Maldonado, R 109, Cerro Cordillera, Arroyo sin nombre, S 34 º 20 ’ 29.4 ’’, W 54 º 37 ’ 40.5 ’’, 301 m, 23 / 3 / 10, D. Emmerich & C. Molineri colls.Published as part of Nieto, Carolina & Emmerich, Daniel, 2011, Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay, pp. 57-65 in Zootaxa 2996 on pages 61-63, DOI: 10.5281/zenodo.20543
A new species of Asthenopus (Ephemeroptera: Polymitarcyidae: Asthenopodinae) from Brazil and Colombia
No full textMolineri, Carlos, Cruz, Paulo Vilela, Emmerich, Daniel (2011): A new species of Asthenopus (Ephemeroptera: Polymitarcyidae: Asthenopodinae) from Brazil and Colombia. Zootaxa 2750: 33-38, DOI: 10.5281/zenodo.27667
Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay
No full textNieto, Carolina, Emmerich, Daniel (2011): Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay. Zootaxa 2996: 57-65, DOI: 10.5281/zenodo.20543
Cloeodes vaimaca Nieto & Emmerich, 2011, sp. nov.
No full text<i>Cloeodes vaimaca</i> sp. nov. <p>(Figs. 3, 26–35)</p> <p>Nymph (Fig. 3). Length: body, 4.1–4.2 mm; cerci, 1.4–1.5 mm; terminal filament, 1.3–1.4 mm. Antennae partially broken. Head pale yellow, longer than wide, compound eyes pale yellow. Antennae pale yellow. Mouthparts (Figs. 26–31): labrum (Fig. 26 a) with one subapical seta centrally and two-three setae near lateral margin, anterior margin with bifid setae (Fig. 26 b). Left mandible (Fig. 27) without setae between prostheca and mola, thumb of molar area transverse to anterior margin. Right mandible (Fig. 28) without setae between prostheca and mola. Hypopharynx (Fig. 29): lingua with rounded projection and longer than superlinguae. Maxillae (Fig. 30), palpi scarcely shorter than galea-lacinia, two segmented, segment II with a constriction. Labium (Fig. 31) with segment III of palpi rounded.</p> <p>Thorax pale yellow. Pleura pale yellow, sterna whitish. Legs pale yellow. Dorsal edge of femora with a row of pointed spines, apically with a rounded projection and two blunt spines (Fig. 32). Tarsal claws 0.4 times the length of tarsi. Hind wing pads present.</p> <p>Abdomen pale yellow, segments I–V with a reddish spot in the midline. Abdominal terga with covers, posterior margin of terga with spines as in Fig. 33. Sterna pale yellow with trachea pigmented, sterna II-VI with a tuft of long fine setae laterally on each side. Paraprocts with spines apically (Fig. 34). Gills (Fig. 35) translucent, trachea pigmented, 2.5 times the length of each tergum. Caudal filaments pale yellow, with flattened setae basally sclerotized (similar to Fig. 14 b). Cerci with long spines toward external margin every two segments, terminal filament with long spines toward midline every two segments (similar to Fig. 35).</p> <p> <b>Etymology.</b> Vaimaca was one of the four charrúas taken to Paris in 1833 to be exhibited in the "Museum of the Man ". This fact crowned the humiliation and destruction of this people for the colonial Spanish model. We dedicate respectfully this species to his memory.</p> <p> <b>Diagnosis.</b> <i>Cloeodes vaimaca</i> <b>sp. nov.</b> can be distinguished from the other species of the genus by the following combination of characters: 1) hind wing pads present; 2) lingua with rounded projection and longer than superlinguae (Fig. 29); 3) labium (Fig. 31) with segment III of palpi rounded; 3) dorsal margin of femora with a row of pointed spines, apically with a rounded projection and two blunt spines (Fig. 32); 4) abdominal color pattern as in Fig. 3, with segments I-V with a reddish spot in the midline</p> <p> <b>Material.</b> Holotype male nymph: URUGUAY, Flores, R. 57, km 30, Arroyo Guardia Vieja, S 33º 42´12´´, W 57º 06´17´´, 120m, 6/ 3/ 2008, Emmerich & Pérez colls. Paratypes: three nymphs (one dissected) same data as holotype. One nymph: Paysandú, R. 3, km 366, Arroyo Sacra, S 32º 21´02´´, W 58º 03´04´´, 10m, 7/ 3/ 2008, Emmerich & Pérez colls. One nymph: Río Negro, R 3, Km 291, Camping. Arroyo San Esteban, S 32° 50’ 07’’, W 57º 30’ 59’’, 34 m, 2/ 3/ 0 8, D. Emmerich & S. Pérez colls.</p>Published as part of <i>Nieto, Carolina & Emmerich, Daniel, 2011, Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay, pp. 57-65 in Zootaxa 2996</i> on pages 63-65, DOI: <a href="http://zenodo.org/record/205436">10.5281/zenodo.205436</a>
Tricorythodes arequita Traver 1959
No full textT. arequita Traver Previously reported in Uruguay from Lavalleja, we here provide additional records. Uruguay: 4 nymphs from Rivera, Cañada sin nombre, 289 m, S 31° 5’ 36.7’’ – W 55° 57’ 57.9’’, 11/XI/2008, Emmerich col.; and 5 nymphs from Treinta y Tres, Arroyo Yerbal Chico, 78 m, S 32° 55’ 35.2’’ – W 54° 27’ 39.6’’, 25/III/2010, Emmerich & Molineri cols.Published as part of Emmerich, Daniel & Molineri, Carlos, 2020, A new species of Tricorythodes Ulmer (Ephemeroptera: Leptohyphidae) from Uruguay, pp. 125-132 in Zootaxa 4885 (1) on page 132, DOI: 10.11646/zootaxa.4885.1.8, http://zenodo.org/record/429644
Paracloeodes morellii Emmerich & Nieto, 2009, sp. nov.
No full textParacloeodes morellii sp. nov. (Figs. 13–25) Nymph. Length: body: 2.8–3.3 mm; cerci 1.0– 1.3 mm. Antennae 0.7–0.8 mm, 1.5 times the head capsule. General color brownish orange. Head orange brown, with small brownish spots as in Figs. 13–14. Frontal keel present, ocelli brownish. Antennae: scape and pedicel pale yellow, flagellum pale. Mouthparts (Figs. 15–20): labrum (Figs. 15 a–b) dorsally with 2–3 subapical setae near midline, anterior margin with bifid setae. Mandibles (Figs. 16–17) without setae between prostheca and mola, incisors cleft in two sets, inner set at right angle with outer set. Left mandible (Fig. 16) with molars with constrictions, prostheca broad with 1–2 denticles. Right mandible (Fig. 17) with prostheca bifid. Hypopharynx (Fig. 18) with lingua subequal to superlinguae and rounded apically. Maxillae (Fig. 19) with palpi two segmented, 1.5 times the length of galealacinia. Labium (Fig. 20) with segment II of palpi with rounded projection, 2 times width of segment III, segment III conical. Thorax (Fig. 13) orange brown. Pronotum with a pair of brownish spots in the midline. Mesonotum shadowed with brown and with small brownish spots. Metanotum brownish. Pleura reddish brown, sterna whitish. Legs (Fig. 21) pale yellow. Tarsal claws 0.4 times the length of tarsi, with 2 rows of minute denticles (Fig. 22). Hind wing pads absent. Female (Fig. 14) paler than male and with a brown band in metanotum and anterior portion of fore wing pads. Abdomen orange brown. Color pattern: segment II–VII with medial elongate spot as in Fig. 13, segment IX brownish. Posterior margin of terga with spines as long as wide (Fig. 23). Sterna pale yellow, sternum IX brownish except at lateral sides. Gills whitish, translucent, oblong, 1.5 times the length of each tergum. Paraprocts (Fig. 24) with 8–10 spines apically. Terminal filaments pale yellow, with crown of small spines on each segment (Fig. 25). Adults. Unknown. Etymology. This species is dedicated to Dr. Enrique Morelli, entomologist of Universidad de la República (Uruguay) who continually stimulates the study of the aquatic insects of Uruguay. Discussion. This species can be distinguished from the other species of the genus by the following combination of characters: 1) frontal keel present; 2) antennae short, 1.5 times the head capsule; 3) segment II of labial palpi with rounded projection, 2 times width of segment III (Fig. 20); 4) tarsal claws with two rows of minute denticles (Fig. 22); 5) hind wing pads absent; 6) general color pattern with head and thorax with small spots, abdomen with segment II–VII with medial elongate spot as in Fig. 13; 7) posterior margin of terga with spines as long as wide (Fig. 23); 8) paraprocts with 8–10 spines apically (Fig. 24); 9) terminal filament with crown of small spines on each segment (Fig. 25). Material. Holotype, male nymph: URUGUAY, Deptos. Rivera-Tacuarembó, R 44, km 26.5, A° Carpintería, S 31°48´28´´, W 55°13´09´´, 121 m, 20/02/2008, Emmerich & Molineri col. Paratypes: 5 nymphs same data (mouthparts, legs and paraprocts of 1 nymph mounted on slide); 1 nymph (parts on slide Depto. Durazno, Sarandí del Yi, Río Yi, S 33°20´41´´, W 55°37´07´´, 132 m, 21/02/2008, same collectors; 2 nymphs (1 on slide): Depto. Cerro Largo, ruta 8, km 335, A° Otazo, S 32°50´00´´, W 54°14´55´´, 123 m, 20/02/2008, Same collectors. The holotype and 1 paratype female are deposited in the Museo de la Universidad de la República, Montevideo, Uruguay (MUR); remaining paratypes are housed in Instituto Miguel Lillo, Tucumán, Argentina (IML).Published as part of Emmerich, Daniel & Nieto, Carolina, 2009, First record of the genus Paracloeodes Day (Ephemeroptera: Baetidae) in Uruguay and description of two new species, pp. 57-62 in Zootaxa 2225 (1) on pages 60-62, DOI: 10.11646/zootaxa.2225.1.3, http://zenodo.org/record/532027
Paracloeodes charrua Emmerich & Nieto, 2009, sp. nov.
No full textParacloeodes charrua sp. nov. (Figs. 1–12) Nymph. Length: body: 3.8–4.3 mm; cerci 1.8 mm. Antennae long, 2.1 mm, 3.5 times width of the head capsule. General color yellowish brown. Head yellowish brown with a characteristic pattern (Fig. 1). Frontal keel present, ocelli black. Antennae pale yellow. Mouthparts (Figs. 2–7): labrum (Figs. 2 a–b) dorsally with a pair of subapical setae near midline, anterior margin with bifid setae. Mandibles (Figs. 3–4) without setae between prostheca and mola, incisors cleft in two sets, inner set at right angle with outer set. Left mandible (Fig. 3) with molars with constrictions, prostheca with two denticles. Right mandible (Fig. 4) with prostheca slender and bifid. Hypopharynx with lingua shorter than superlinguae and rounded apically. Maxillae (Fig. 6) with palpi two-segmented, twice the length of galea-lacinia. Labium (Fig. 7) with segment II of palpi with rounded projection, 2 times width of segment III, segment III conical. Thorax yellowish brown, shadowed with brown (Fig. 1). Pleura brownish, sterna pale yellow. Legs (Fig. 8) pale yellow, femora with brownish transverse band apically (more conspicuous in female). Tarsal claws (Fig. 9) 0.5 times the length of tarsi, with 2 rows of small denticles basally and 4–5 denticles increasing in size apically. Hind wing pads present. Abdomen yellowish brown (Fig. 1). Color pattern: segment II–IX with 3 brownish spots forming a triangle, segment V and IX darker than others. Posterior margin of terga with spines 2 times longer than wide (Fig. 10). Sterna yellowish brown, sternum IX darker than others. Gills whitish, elongate, less than twice the length of each tergum, main branch of trachea pigmented. Paraprocts (Fig. 11) with 10–12 spines apically. Terminal filaments with crown of spines on each segment, cerci with long spines toward the external margin every four segments (Fig. 12). Adults. Unknown. Etymology. Charrua: ethnic group that inhabited Uruguay and was exterminated in the middle of the 19th Century. We dedicate this species to their memory. Discussion. This species can be distinguished from the other species of the genus by the following combination of characters: 1) frontal keel present; 2) antennae long, 3.5 times width of head capsule; 3) maxillae with palpi twice the length of galea-lacinia (Fig. 6); 4) segment II of labial palpi with rounded projection, 2 times width of segment III (Fig. 7); 5) tarsal claws 0.5 times the length of tarsi with two rows of small denticles basally and 4–5 denticles increasing in size apically (Figs. 8–9); 6) hind wing pads present; 7) abdomen with a characteristic color pattern: segment II–IX with 3 brownish spots forming a triangle (Fig. 1); 8) posterior margin of terga with spines twice longer than wide; 9) paraprocts with 10–12 spines apically; 10) cerci with long spines toward the external margin every four segments (Fig. 12). Material. Holotype, female nymph: URUGUAY, Depto. Cerro Largo, Arroyo Otazo, ruta 8, km 335, S 32°50´00´´, W 54°14´55´´, 123 m. 20/02/2008, Emmerich & Molineri cols. Paratypes: 4 nymphs same data (mouthparts, legs and paraprocts of one nymph mounted on slides); 4 nymphs: Depto. Rivera, Tranqueras, ruta 30, km 251, Río Tacuarembó, S 31°10´40´´, W 55°45´44´´, 147 m, 19/02/2008, same collectors; 5 nymphs: Depto. Tacuarembó, ruta 5, km 379, A° Tranqueras, S 31°47´29´´, W 55°58´51´´, 124 m, 22/02/ 2008, same collectors; 3 nymphs: Dpto Soriano, ruta 21, km 339, Aº Las Maulas, S 33º 23’ 32’’, W 58º 07’ 02’’, 16 m, 6/03/08, Emmerich & Pérez cols.; 11 nymphs: Dpto Paysandú, “Predio”, Aº Grande, S 32º 39’ 34.2’’, W 57º 14’ 56.7’’, 9/08/07, Morelli col. The holotype is deposited in the Museo de la Universidad de la República, Montevideo Uruguay (MUR), paratypes are deposited in Instituto Miguel Lillo, Tucumán, Argentina (IML).Published as part of Emmerich, Daniel & Nieto, Carolina, 2009, First record of the genus Paracloeodes Day (Ephemeroptera: Baetidae) in Uruguay and description of two new species, pp. 57-62 in Zootaxa 2225 (1) on pages 58-60, DOI: 10.11646/zootaxa.2225.1.3, http://zenodo.org/record/532027
Campsurus vulturorum Emmerich & Molineri, 2011, sp. nov.
No full text<i>Campsurus vulturorum</i> sp. nov. <p> <b>Material.</b> Holotype male Imago from Uruguay, Maldonado, Ruta 60, Km 23, Arroyo El Rodeo, S 34° 40' 20" W 55° 14' 15", 75 m, 4/III/08, at dusk, D. Emmerich & S. Pérez cols.; allotype female imago, same data as holotype. Paratypes: nineteen male imagos and 9 female imagos, same data as holotype. Five male imagos and 5 female imagos, same data as holotype except: 7/XI/08, D. Emmerich col.; one male and 1 female imagos, Uruguay, Treinta y Tres, Quebrada de los Cuervos, Arroyo Los Helechos, S 32° 55' 27.3" W 54° 27' 34.4", 125 m. 8/XI/04, at dusk, D. Emmerich & E. Morelli cols. One male imago and 1 female imago from Uruguay, Rivera, Santa Ernestina, Ruta 29, Arroyo sin nombre, near Mina Corrales, S 31º 32' 23.8" W 55º 33' 42", 146 m, 10/XI/2008, at dusk, D. Emmerich col. Two male subimagos Argentina, Misiones, Dpto Alem, 5 km W Cerro Azul, RN 14, Arroyo Mártires, 17/ XI/1998, E. Domínguez, C. Molineri, C. Nieto & M. Orce cols. Seven mature nymphs, Uruguay, Canelones, Carrasco, "lagunas areneras Minetti", 17/VI/51.</p> <p> <b>Depositories.</b> Holotype and allotype, six male imagos, six female imagos, two male subimagos and three nymphs housed in IML (Instituto Miguel Lillo, Tucumán, Argentina); remaining material in MUR (Museo de la Facultad de Ciencias de la Universidad de la República, Uruguay).</p> <p>Male imago. Length (mm): body, 16.0–21.5; fore wing, 15.9–20.5; hind wing, 7.0–9.5; cerci, 43.0–50.0; fore leg, 8.0–9.0. General coloration whitish with well-defined black markings. Head (Fig. 1) whitish, completely shaded black dorsally; shading slightly more diffuse on occipute, except medial pale mark near hind margin. Antennae whitish, shaded slightly with purplish. Thorax. Pronotum whitish, except pronotal hump translucent; large medial area shaded with black except on posterior ¼; medial line on distal ½, hind margin and posterior half of lateral margin shaded black (lateral marks wider); prosternum translucent. Meso- and metanotum cream, shaded with black on carinae; transverse dash anterior to wing insertion and on a posterior V-shaped mark shaded diffusely with gray on mesoscutellum and around parapsidal sutures; rest of meso-and metathorax pale cream without marks (Fig. 1); inner margins of furcasternal plates diverging on posterior ½. Legs: fore legs whitish, widely shaded with grayish purple (darker on tibiae), except on a pale dorsal mark on proximal third of femur; apices of tarsites paler (Fig. 2); middle and hind legs whitish translucent. Wings. Membrane of fore wing (Fig. 9) hyaline; C, Sc and R1 purplish, turning hyaline on distal half; remaining longitudinal and cross veins translucent hyaline; hind wing (Fig. 10) veins translucent hyaline. Abdomen translucent whitish shaded with brownish gray on terga except medial line; terga II–VII with pale lateral dashes, terga VIII–X shaded widely except on a pair of submedian pale spots (Figs 3– 4); abdominal sterna pale without shading, except a pair of small black dashes on sternum X. Genitalia (Figs 11– 16): sternum IX whitish, slightly projected medially (arrow in Fig.12), pedestals whitish except posterolateral projection yellowish (Figs 12, 16), forceps whitish, penes whitish except membranous ventral lobe hyaline and dorsal sclerotized portion (ventrally curved) yellowish (Figs 13–15). Caudal filaments translucent whitish.</p> <p>Female imago. Length (mm): body, 19.0–25.0; fore wing, 21.0–28.0; hind wing, 8.2–12.0; cerci, 6.7–7.2. General coloration similar to male (Figs 5–8). Legs whitish shaded with purplish in fore femur and grayish on middle coxa (Fig. 6). Mesopleura with small blackish mark ventrally to wing insertion. Wings membrane translucent slightly smoky yellow, costal area shaded with purplish almost entirely. Abdomen shaded darker and more widely with gray than in male; sternum X shaded black on anterolateral corners (Figs 7–8). Sternum VIII on anterior margin with a pair of posteriorly divergent sockets (Fig. 17).</p> <p>Eggs. Maximum length, 330–355 µm; maximum width, 255–290 µm. Yellowish white, bowl shaped, oval, with a relatively large polar cap situated at the apex of the main axis (Fig. 56 B).</p> <p>Nymph. Length (mm). Male: body, 17.0; cerci, 17.0; caudal filament, 8.0. Female: body, 24,5–30.0; cerci, 9.0; caudal filament, 7,5–8.0. General coloration yellowish with brownish marks (Fig.18). Head shaded with purplish brown between lateral ocelli and netted pattern on occipute (Fig. 19). Mandibular tusks slightly curved dorsally at the apex, with numerous setae dorsally; inner margin with a prominent subbasal tubercle and 6 smaller more distal tubercles; outer margin of mandibles with 3–4 rows of stout spines and numerous setae (Figs 20–22). Maxillae with a short ventral gill (Fig. 23). Thorax. Pronotum whitish-yellow shaded with brownish as in male; prosternum translucent with a medial gray mark anterior to coxae; mesonotum extensively shaded with purplish brown, darker on posterior margin; wing buds translucent, shaded slightly with purplish on base, costal margin and main veins (Fig. 19). Legs. Fore legs yellowish white; fore tibia-tarsus strongly flattened with a large dorso-distal projection (Fig. 24), ventral surface with 4 rows of filtering setae: a basal transversal row, and three longitudinal rows (Fig. 25), dorsal surface almost completely covered with short spines and simple setae; femur with anterobasal U-shaped row of filtering setae, and contiguously with a posterior elongated tuft of simple setae; dorsal margin of femur with posterobasal and distal tufts of setae (Fig. 25). Middle and hind legs whitish yellow almost completely covered with yellowish setae. Abdomen whitish, shaded with purplish gray dorsally except on pale medial line, terga shaded mainly on submedian area and near posterior margin; terga III–VIII with pale semicircular sublateral marks, terga VIII–X shaded more extensively except on 1 or 2 pairs of sublateral pale marks (Fig. 18). Gills. Abdominal gill I bilobed and translucent, gills II–VII shaded with purplish darker on trachea and posterior margin except articulation between basal and distal portions of dorsal lamellae pale; fringes whitish except on distal half of posterior margin light purplish at base. Caudal filament whitish.</p> <p> <b>Etymology.</b> From the Latin word <i>vulturus</i>, plural genitive: <i>vulturorum</i> (= of the vultures). From the name of one of the localities “Quebrada de los Cuervos” where the species was collected.</p> <p> <b>Diagnosis and discussion.</b> <i>Campsurus vulturorum</i> can be distinguished from all other species of the genus by the following combination of characters, in the adults: 1) posterior margin of male abdominal sternum IX convex, rounded (Figs 11–12); 2) pedestals subrectangular, with a short projection on outer-posterior margin (Figs 12, 16); 3) penes widely separated distally, each arm formed by two lobes, a larger sclerotized dorsal lobe, and a ventral smaller and membranous lobe (Figs 13–15); 4) male gonopore large, clearly visible before an apical rounded ventral projection on the dorsal penean lobe (Fig.14); 5) very large size (male fore wings 15.9–20.5 mm, female fore wings 21.0–28.0 mm); 6) blackish pigments strongly marked on head and pronotum, mainly on medial line and posterolateral margins of pronotum (Figs 1, 5); 7) abdominal color pattern as in figures 3–4 and 7–8; 8) female sternum VIII with paired and posteriorly divergent sockets on anterior margin as in figure 17. In mature nymphs: 1) mandibular tusks robust with a large submedian tubercle and 6 subapical rounded denticles (Figs 20–22); 2) maxilla with a short flat and triangular membranous gill on cardo (Fig. 23). The association between adults and nymphs is tentative, based on shared color pattern, wing venation and genital rudiments.</p> <p> <b>Distribution.</b> Uruguay and Northeastern Argentina (Misiones)</p>Published as part of <i>Emmerich, Daniel & Molineri, Carlos, 2011, A new species of Campsurus (Ephemeroptera: Polymitarcyidae: Campsurinae) from Argentina and Uruguay and redescription of C. evanidus and C. jorgenseni with new synonymies, pp. 51-60 in Zootaxa 2965</i> on pages 52-53, DOI: <a href="http://zenodo.org/record/205154">10.5281/zenodo.205154</a>
FIGURES 1–3. Nymphs, dorsal habitus. 1 in Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay
No full textFIGURES 1–3. Nymphs, dorsal habitus. 1, Cloeodes dialutoi sp. nov. 2, Cloeodes guenoa sp. nov. 3, Cloeodes vaimaca sp. nov.Published as part of Nieto, Carolina & Emmerich, Daniel, 2011, Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay, pp. 57-65 in Zootaxa 2996 on page 59, DOI: 10.5281/zenodo.20543
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