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    Euchondrus adwani Eike Neubert & Zuhair Amr 2016, n. sp.

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    Euchondrus adwani n. sp. (Figure 1) Material: Holotype NMBE 539263; paratype NMBE 539264 /1. Type locality: Syria, surrounding of the monastery of Deir Moussa, 34.0219°N 36.8423°E, 1300 m a.s.l., 11.iii. 2010, leg. Adwan Shehab. *Corresponding author. Email: [email protected] © 2016 Taylor & Francis Measurements (holotype): Height = 11.04 mm; diameter = 4.13 mm; peristome height = 4.03 mm; peristome diameter = 2.97 mm; number of whorls = 8. Diagnosis. Euchondrus adwani n. sp. differs from the widespread E. septemdentatus by its conical shell (broadly oval in E. septemdentatus), its flat suture and teleoconch whorls (suture deeper, whorls much more rounded in E. septemdentatus), the heavy palatal labial callus (weaker in E. septemdentatus), the bar-like subangularis (weaker in E. septemdentatus), and the keeled last whorl (rounded in E. septemdentatus). Description: Shell solid, dextral, cylindrical, upper part cylindro-conical in outline; shell pale brownish to horny yellow coloured; 8 rather flat-sided teleoconch whorls, suture flat with a distinct white sutural thread; teleoconch smooth, glossy, last whorl with fine, straight and irregularly spaced striae; aperture subtriangular, peristome strongly thickened by a labial callus, moderately reflected, with a rich dentition (description clockwise): palatum with a small suturalis followed by a conical palatalis superior and a broad infrapalatalis with the latter two denticles placed on a thick callus; columellar side with a basalis and a straight columellaris; parietum with a strong and long parietalis, bordered by a small spiralis, subangularis large, bar-like, left side of the parietum with another small denticle at the attachment site of the peristome; last teleoconch whorl dorsally compressed forming a distinct blunt ridge (arrows); umbilicus slit-like open, periomphalum large, dish-like. Remarks: This species shows some superficial similarities with E. desertorum Rochanaburananda in Forcart, 1981 (Figure 2), which is endemic to the Negev Desert (Heller, 2009). Both species have a straight conical shell, but E. desertorum is considerably larger than E. adwani n. sp. and its aperture is rounded and not subtriangular. It also differs in the formation of the dentition: in E. desertorum, the infrapalatalis is bifid (simple in E. adwani), the spiralis is large and connected to the parietalis (small and disconnected in E. adwani), and the subangularis is weaker (very strong in E. adwani). The last whorl of E. adwani displays a distinct keel with an enlarged periomphalum, while in E. desertorum the dorsum is rounded, and the periomphalum is much smaller. Etymology: This species is named in honour of Dr. Adwan Shawabi, who was a keen collector of molluscs from Syria, and a personal friend, and who was killed in February 2015 in the Syrian civil war (Amr, 2015).Published as part of Eike Neubert & Zuhair Amr, 2016, On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae), pp. 58-60 in Zoology in the Middle East 62 on pages 58-60, DOI: 10.1080/09397140.2015.1132564, http://zenodo.org/record/88703

    On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae)

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    Eike Neubert, Zuhair Amr (2016): On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae). Zoology in the Middle East 62: 58-60, DOI: 10.1080/09397140.2015.113256

    Figure 1–2. 1 in On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae)

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    Figure 1–2. 1- Euchondrus adwani, n. sp., holotype NMBE 539263, Syria, monastery of Deir Moussa. – 2- Euchondrus desertorum, Negev, 1 km W Sedé Boqer, 06.03.1962, leg. H. Sandler, coll. Bank. — Photograph: E. Bochud.Published as part of Eike Neubert & Zuhair Amr, 2016, On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae), pp. 58-60 in Zoology in the Middle East 62 on page 59, DOI: 10.1080/09397140.2015.1132564, http://zenodo.org/record/88703

    Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)

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    Ruud, A. Bank, Henk P.M.G Menkhorst, Eike Neubert (2016): Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae). Basteria 80 (1): 5-30, DOI: 10.5281/zenodo.43974

    Clausilioides Lindholm 1925

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    Clausilioides Lindholm, 1925 Clausilioides Lindholm, 1925: 29, 38. Type species (by monotypy): Buliminus (Brephulus) biplicatus Retowski, 1889. So far, only two species have been assigned to the genus Clausilioides, namely C. biplicatus (Retowski, 1889) (synonym: eplicatus Lindholm, 1913) and C. filifer (Lindholm, 1913). We here describe another two species. The genus seems to be endemic for the northeastern part of Turkey, although both biplicatus and filifer have originally been described from Georgia (but only found in deposition of the Çoruh river). A characteristic feature of the genus is the presence of a well developed palatal fold in the last whorl. This is a rare character within the family Enidae. It has been reported in Pseudonapaeus entoptyx (Lindholm, 1925) (the type species of Siraphorus Lindholm, 1925), Siraphoroides moltschanovi (Likharev & Rammelmeier, 1952) (a monotypic genus), and various species of the genus Pupopsis Gredler, 1898. The first two species are known from Kirgizstan, whereas Puposis is distributed in the northwestern part of China (Xinjiang at the border of Kirgistan and Kazachstan) as well as in central China (SE Gansu and N Sichuan). For a review on Pupopsis see Wu & Gao (2010). Furthermore, a palatal fold is seen in Buliminus glabratus (Mousson, 1861) from the central part of Israel.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on page 17, DOI: 10.5281/zenodo.43974

    Imparietula Lindholm 1925

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    Imparietula Lindholm, 1925 Imparietula Lindholm, 1925: 30, 39. Type species (by monotypy): Bulimus leucodon L. Pfeiffer, 1846. Forcart (1940: 206) considered Pseudochondrula a synonym of Imparietula, which was followed by Gittenberger (1967: 130-137), Akramowski (1976: 154-158) and Hausdorf (1999: 153). Schileyko (1984: 288, 306) separated Pseudochondrula from Imparietula, which was followed by Bank & Neubert (1998: 81). In his latest edition of the Turkish land snails, Schütt (2010: 97-99) also separated Pseudochondrula from Imparietula, and included the following taxa into the latter genus: altenai (Gittenberger, 1967), pelidne (Biggs, 1946), leucodon (L. Pfeiffer, 1846) (with lasistanicus as a synonym), bre- vior (Mousson, 1876) and ridvani Schütt, 1995. Following a taxonomic revision, Páll-Gergely & Bank (in press) raised lasistanica (Lindholm, 1914) to species level, and excluded brevior as a representative of Imparietula. By doing so, Imparietula has become a genus that is endemic for the northeastern part of Turkey. We here introduce another species to the genus.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on pages 13-14, DOI: 10.5281/zenodo.43974

    Multidentula Lindholm 1925

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    Multidentula Lindholm, 1925 Multidentula Lindholm, 1925: 30, 39. Type species (by monotypy): Bulimus ovularis Olivier, 1801. Bollingeria Forcart, 1940: 194. Type species (by original designation): Chondrus pupoides Krynicki, 1833. Tokatia Hudec, 1972: 217. Type species (by original designation): Bulimus lamelliferus Rossmässler, 1858. Improvisa Schileyko, 1978: 846. Type species (by monotypy): Chondrus pupoides Krynicki, 1833. Senaridenta Schileyko, 1978: 846. Type species (by monotypy): Chondrula (Chondrula) nachicevanjensis Hudec, 1972. The genus Multidentula is known from the Caucasus, Armenia, Georgia, Azerbaijan, northwestern Iran and the northern half of Turkey; its presence in Cyprus is questionable (Bank & Neubert, 2015: 73). The following species belong to the genus: lamellifera (Ross- mässler, 1858), nachicevanjensis (Hudec, 1972), ovularis (Olivier, 1802), pupoides (Krynicki, 1833), ridens (Nägele, 1906) and squalina (L. Pfeiffer, 1848). The latter species is polytypic and is almost endemic for Turkey. Forcart (1940: 197-199, pl. 3 figs 64-65) mentioned M. s. squalina and M. s. eudoxina (Nägele,1894) for Turkey. Based on a single shell, Lindholm (1922b: 358-359) described Chondrula acutior from Georgia; this shell (holotype) has been figured by Sysoev & Schileyko (2009: fig. 38E) under the name Euchondrus acutior. It has never been found again after the de- scription from Lindholm. Our revision revealed that this taxon is also distributed in the northeastern part of Turkey, and that it is a subspecies of M. squalina. We here characterize this virtually unknown subspecies, and also describe a new species from Turkey.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on page 22, DOI: 10.5281/zenodo.43974

    Multidentula reducta Ruud, Menkhorst & Neubert, 2016, spec. nov.

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    Multidentula reducta spec. nov. (Fig. 19) Type locality & type specimens. – Turkey, Vilayet Sivas, Zara, 16 km towards Imranlı, 1400 m (39.8606°N 37.8765°E), H.P.M.G. Menkhorst leg., 15.vii.1988. Holotype NMBE 544681, paratypes NMBE 544682/1, HMK/12, RBA/1, RMNH/1. Diagnosis. – A middle-sized, oval Multidentula species with fine oblique striae, a missing suprapalatalis and a weakly developed or missing basalis, an oblique infrapalatalis, a non-fused subangularis, and a parietalis that is only slightly curved. Description. – Shell dextral, oval in outline, with a rather wide, open, slit-like umbilicus. The 6.0-7.2 whorls are convex with a rather deep suture. Teleoconch with irregular, sparse, oblique striae which are mostly rather fine; there are no spiral striae. Shell rather solid, slightly translucent, yellowish to horny yellowish, with a whitish band behind the peristome. The last whorl has no or only an indistinct impression and only the infrapalatalis is visible outside as a blurred white stripe. Peristome well reflected, thickened by a prominent labial callus, the columellar and palatal insertion connected by a clearly visible callus (but the callus is not extra thickened near the insertions). The subangularis is vertically pointing downwards, and is connected (but not fused) with the palatal peristome by a callus. The subangularis and parietalis are connected by a thin callus. Parietalis prominent, slightly curved and deeply recessed. A spiralis is missing. Columellaris ± oblique to the columellar peristome, deeply recessed. The basalis is absent, or present as an indistinct thickening only. Infrapalatalis well developed, oblique, and always stronger developed than the non-oblique palatalis superior. There is no suprapalatalis; a small suturalis is present. Measurements (n = 6). – H = 5.3-6.9 (mean 6.1); LWH = 3.2-3.8 (mean 3.5); MH = 2.1-2.4 (mean 2.2); LWD = 2.7-2.8 (mean 2.8); LWM = 2.7-3.0 (mean 2.8); MD = 1.7-1.9 (mean 1.8); NW = 6.0-7.2 (mean 6.6). Localities. – Known from the locus typicus only (see above). Derivatio nominis. – The name refers to the reduced apertural armature. Differentiation. – The armature of M. reducta is generally reduced: a basalis is absent or only weakly developed, a suprapalatalis is missing, and the parietalis is not as prominent and less curved as in M. squalina or M. pupoides. The palatal folds are, compared to M. squalina, also less deep in the aperture, but are placed closer to the peristome.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on page 23, DOI: 10.5281/zenodo.43974

    Amphiscopus sturmii subsp. marmoratus Ruud, Menkhorst & Neubert, 2016, subspec. nov.

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    Amphiscopus sturmii marmoratus subspec. nov. (Fig. 15) Type locality & type material. – Turkey, Vilayet Kütahya, Örencik, western part, 950 m (39.4426°N 29.0662°E), B. Hausdorf leg., 21.ix.1987. Holotype NMBE 544670/1, paratypes NMBE 54467/5, HMK/2, RBA/*2, ZMH/33. Diagnosis. – A subspecies of A. sturmii characterized by the whitish/ marbled colour of the shell, the less well-developed parietal callus and the absence of an impression at the last whorl. Description. – Shell dextral, turreted, with a relative wide, open, slit-like umbilicus. The 8.3-10.1 whorls are rather convex with a moderately deep suture. Teleoconch with densely packed, rather pronounced, oblique striae; there are no spiral striae. Shell solid, not translucent, glossy, upper part of the teleoconch horny yellow coloured, the remaining part of the teleoconch whitish/ marbled coloured, but below the periphery (i.e. the part of the whorl below the insertion of the palatal peristome) again more horny yellow; there is a prominent white band behind the peristome. The last whorl does not have an impression near the palatalis superior. Peristome not or hardly reflected (with the exception of the columellar part), robustly thickened by a labial callus, the columellar and palatal insertion connected by a clearly visible callus which is often more thickened near the columellar peristome. The subangularis is tear-like and vertically pointing downwards; it is connected with the palatal peristome by a thin callus. The subangularis and parietalis are not connected. The parietalis is well developed, and only moderately deeply recessed. Palatalis superior well thickened, but not deeply recessed. There is no spiralis, infrapalatalis, basalis or columellaris. The columellar ledge reaches halfway or above the middle of the columellar side of the aperture. Measurements (n = 11). – H = 6.7-9.0 (mean 7.8); LWH = 3.0-3.5 (mean 3.3); MH = 1.9-2.2 (mean 2.1); LWD = 2.3-2.7 (mean 2.4); LWM = 2.3-2.8 (mean 2.6); MD = 1.5-1.8 (mean 1.6); NW = 8.3-10.1 (mean 9.2). Localities. – Vilayet Kütahya: type locality (see above); 4 km E. Köprüören, 1020 m (39.5115°N 29.8137°E) (ZMH/>50). Derivatio nominis. – Named after the marble-like colour of the shell. Differentiation. – Amphiscopus sturmii sturmii has a uniform dark-brown coloured shell, more convex whorls, a more prominent parietal callus, and the last whorl has a more or less faint impression at the position of the palatalis superior. Amphiscopus substurmii is sinistral and has the same colour pattern as A. sturmii sturmii.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on page 20, DOI: 10.5281/zenodo.43974

    Amphiscopus moolenbeeki Ruud, Menkhorst & Neubert, 2016, spec. nov.

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    Amphiscopus moolenbeeki spec. nov. (Fig. 14) Type locality & type material. – Turkey, Vilayet Tokat, Reşadiye, 8 km towards Çekerek, 770 m (40.1523°N 35.6320°E), H.P.M.G. Menkhorst leg., 19.vii.1990. Holotype NMBE 544668, paratypes NMBE 544669/2, HMK/>50, RBA/3, RMNH/3, ZMH/3. Diagnosis. – A large, highly turreted Amphiscopus species without a parietalis or palatalis superior. Description. – Shell dextral, highly turreted, with an open, slit-like umbilicus. The 10.4-13.2 whorls are convex with a moderately deep suture. Teleoconch with widely spaced, fine, oblique striae; there are no spiral striae. Shell solid, not translucent, glossy, uniformly horny yellow coloured, with a prominent whitish band behind the peristome. Peristome not or hardly reflected (with the exception of the columellar part), thickened by a labial callus, the columellar and palatal insertion connected by a clearly visible callus which is quite thickened near the columellar peristome. The edge of the palatal insertion forms a wide arc. The dot-like to tear-like subangularis is connected with the palatal peristome by a thin callus. There is no parietalis, spiralis, palatalis superior, infrapalatalis, basalis or columellaris. The columellar ledge reaches halfway or above the middle of the columellar side of the aperture. Measurements (n = 8). – H = 10.0-14.4 (mean 12.3); LWH = 4.0-4.8 (mean 4.3); MH = 2.6-2.9 (mean 2.7); LWD = 3.0-3.3 (mean 3.1); LWM = 2.9-3.3 (mean 3.1); MD = 1.9-2.2 (mean 2.1); NW = 10.4-13.2 (mean 11.9). Localities. – Vilayet Tokat: type locality (see above); Reşadiye, 7.5 km towards Çekerek (HMK/1). Derivatio nominis. - Named after our friend Rob(ert) Moolenbeek, who made numerous important malacological contributions and who curated the molluscan collection of the formerly Zoologisch Museum Amsterdam for decades (now integrated in the Naturalis Biodiversity Center, Leiden). Remarks. – This taxon has been assigned to Amphiscopus because of the overall shape of the shell, the small aperture and the thickened peristome. So far, only the species sturmii and substurmii has been assigned Amphiscopus; therefore moolenbeeki is now the third known Amphiscopus species. It can be easily recognized from A. sturmii and A. substurmii by the characters mentioned in the diagnosis.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on pages 20-22, DOI: 10.5281/zenodo.43974
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