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Pheidole magna Eguchi
Pheidole magna Eguchi Figs. 14a-g Pheidole magna Eguchi, 2006: 125-127. Holotype: major, "Bang Khoang (Site-A: ca. 1700-1800 m alt.), Sa Pa, Lao Cai, Vietnam, Eg02-VN-137", IEBR, examined; paratypes: 34 majors & 35 minors, same data as holotype, BMNH, FSKU, IEBR, MHNG, MCZC & ACEG, examined. Pheidole sp. eg-162. Bui & Eguchi 2003: 9 (checklist), Eguchi, Bui et al. 2005: 91 (checklist). Other material examined: Vietnam: Lao Cai: Sa Pa town [Eg02-VN-087], Bang Khoang (Site-A: a stream-side secondary forest), 1700-1800 m alt., Sa Pa [Eg02-VN-116, -124, -129], Bang Khoang (Site-B: a well-developed forest), ca. 1700 m alt. [Eg02-VN-165, -169, -175], Sa Seng (small fragment of limestone forest), Sa Pa [Eg02-VN-280]; Ha Tay: Ba Vi N.P. [T.V. Bui]. Eguchi's informal species code " Pheidole sp. eg- 162" has been applied to these specimens. Worker measurements & indices: Major (data from the original description). - HL 2.21-2.39 mm; HW 2.13-2.32 mm; CI 92-99; SL 1.04-1.14 mm; SI 45-51; FL 1.59-1.66 mm; FI 70-75. Minor (data from the original description). - HL 0.87-0.94 mm; HW 0.79-0.90 mm; CI 91-96; SL 0.94- 1.03 mm; SI 113-122; FL 1.12-1.22 mm; FI 134-143. Worker description Major. - Head in lateral view not or very weakly impressed on vertex; frons and vertex longitudinallyobliquely rugose; vertexal lobe largely smooth and shining; frontal carina and antennal scrobe absent; clypeus with a median longitudinal carina which is sometimes reduced into a weak rugula; hypostoma with a low or inconspicuous median and moderately to strongly developed submedian processes; lateral processes present but reduced, much smaller than submedian processes; antenna with a 3-segmented club; maximal diameter of eye a little longer than antennal segment X. Promesonotal dome with a conspicuous prominence on its posterior slope; humerus not or very weakly produced laterad; the dome at the humeri narrower than at the bottom; propodeal spine small. Petiole almost as long as postpetiole (excluding helcium); postpetiole relatively massive. First gastral tergite smooth and shining, often with a weakly punctured area just around its articulation with postpetiole. Minor. - Head smooth and shining; preoccipital carina complete but weak dorsally; median part of clypeus smooth and shining, with a median longitudinal carina in its anterior half; antenna with a 3-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye shorter than antennal segment X. Promesonotal dome smooth and shining, in lateral view raised relatively highly in front of a conspicuous prominence/mound on its posterior slope; humerus in dorso-oblique view not produced, or very weakly produced; mesopleuron, metapleuron and lateral face of propodeum largely punctured weakly. Petiole almost as long as or a little shorter than postpetiole (excluding helcium); postpetiole massive. Recognition: This species is characterized among Indo-Chinese species by the combination of the following features: in the major vertexal lobe largely smooth and shining; in the major and minor promesonotal dome having a conspicuous prominence on its posterior slope; in the major and minor postpetiole relatively massive. This large-bodied species is similar to Pheidole dugasi Forel, but well distinguished from the latter which has the following characteristics in the major: dorsum of vertexal lobes distinctly rugose; first gastral tergite entirely rugoso-punctured (see also Eguchi 2006). Distribution & bionomics: Known from N. Vietnam. This species occurs from relatively open habitats to forests at relatively high altitude (1000 m alt. or higher), and nests in the soil and rotting logs (Eguchi 2006). Majors serve as repletes (e.g., the type series, Eg02-VN-280).Published as part of Eguchi, K., 2008, A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)., pp. 1-118 in Zootaxa 1902 on pages 52-5
Pheidole magna Eguchi
Pheidole magna Eguchi Figs. 14a-g Pheidole magna Eguchi, 2006: 125-127. Holotype: major, "Bang Khoang (Site-A: ca. 1700-1800 m alt.), Sa Pa, Lao Cai, Vietnam, Eg02-VN-137", IEBR, examined; paratypes: 34 majors & 35 minors, same data as holotype, BMNH, FSKU, IEBR, MHNG, MCZC & ACEG, examined. Pheidole sp. eg-162. Bui & Eguchi 2003: 9 (checklist), Eguchi, Bui et al. 2005: 91 (checklist). Other material examined: Vietnam: Lao Cai: Sa Pa town [Eg02-VN-087], Bang Khoang (Site-A: a stream-side secondary forest), 1700-1800 m alt., Sa Pa [Eg02-VN-116, -124, -129], Bang Khoang (Site-B: a well-developed forest), ca. 1700 m alt. [Eg02-VN-165, -169, -175], Sa Seng (small fragment of limestone forest), Sa Pa [Eg02-VN-280]; Ha Tay: Ba Vi N.P. [T.V. Bui]. Eguchi's informal species code " Pheidole sp. eg- 162" has been applied to these specimens. Worker measurements & indices: Major (data from the original description). - HL 2.21-2.39 mm; HW 2.13-2.32 mm; CI 92-99; SL 1.04-1.14 mm; SI 45-51; FL 1.59-1.66 mm; FI 70-75. Minor (data from the original description). - HL 0.87-0.94 mm; HW 0.79-0.90 mm; CI 91-96; SL 0.94- 1.03 mm; SI 113-122; FL 1.12-1.22 mm; FI 134-143. Worker description Major. - Head in lateral view not or very weakly impressed on vertex; frons and vertex longitudinally-obliquely rugose; vertexal lobe largely smooth and shining; frontal carina and antennal scrobe absent; clypeus with a median longitudinal carina which is sometimes reduced into a weak rugula; hypostoma with a low or inconspicuous median and moderately to strongly developed submedian processes; lateral processes present but reduced, much smaller than submedian processes; antenna with a 3-segmented club; maximal diameter of eye a little longer than antennal segment X. Promesonotal dome with a conspicuous prominence on its posterior slope; humerus not or very weakly produced laterad; the dome at the humeri narrower than at the bottom; propodeal spine small. Petiole almost as long as postpetiole (excluding helcium); postpetiole relatively massive. First gastral tergite smooth and shining, often with a weakly punctured area just around its articulation with postpetiole. Minor. - Head smooth and shining; preoccipital carina complete but weak dorsally; median part of clypeus smooth and shining, with a median longitudinal carina in its anterior half; antenna with a 3-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye shorter than antennal segment X. Promesonotal dome smooth and shining, in lateral view raised relatively highly in front of a conspicuous prominence/mound on its posterior slope; humerus in dorso-oblique view not produced, or very weakly produced; mesopleuron, metapleuron and lateral face of propodeum largely punctured weakly. Petiole almost as long as or a little shorter than postpetiole (excluding helcium); postpetiole massive. Recognition: This species is characterized among Indo-Chinese species by the combination of the following features: in the major vertexal lobe largely smooth and shining; in the major and minor promesonotal dome having a conspicuous prominence on its posterior slope; in the major and minor postpetiole relatively massive. This large-bodied species is similar to Pheidole dugasi Forel, but well distinguished from the latter which has the following characteristics in the major: dorsum of vertexal lobes distinctly rugose; first gastral tergite entirely rugoso-punctured (see also Eguchi 2006). Distribution & bionomics: Known from N. Vietnam. This species occurs from relatively open habitats to forests at relatively high altitude (1000 m alt. or higher), and nests in the soil and rotting logs (Eguchi 2006). Majors serve as repletes (e.g., the type series, Eg02-VN-280).Published as part of Eguchi, K., 2008, A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)., pp. 1-118 in Zootaxa 1902 on pages 52-5
Pheidole fortis Eguchi
Pheidole fortis Eguchi Figs. 7a-7g Pheidole fortis Eguchi, 2006: 118-120. Holotype: major, "Cat Cat (a trail to Mt. Phansipan), ca. 1300-1400 m alt., Sa Pa, Lao Cai, Vietnam, Eg02-VN-264", IEBR, examined; paratypes: 1 major & 19 minors, same data as holotype, IEBR, MHNG, MCZC, BMNH, FSKU & ACEG, examined. Pheidole sp. eg-160. Bui & Eguchi 2003: 9 (checklist), Eguchi, Bui et al. 2005: 91 (checklist). Other material examined: N. Vietnam: Vinh Phuc: Tam Dao N.P., 21°27'N, 105°38'E, 950 m alt. [Sk. Yamane]. Thailand: Chiang Mai: Doi Pui, ca. 1200 m alt., Doi Suthep-Pui N.P. [Eg01-TH-113]. Eguchi's informal species code " Pheidole sp. eg-160" has been applied to these specimens. Worker measurements & indices: Major (data from the original description). - HL 2.12-2.23 mm, HW 1.79-2.05 mm, CI 84-92, SL 0.90-0.91 mm, SI 44-51, FL 1.33-1.39 mm, FI 67-78. Minor (data from the original description). - HL 0.71-0.75 mm, HW 0.64-0.71 mm, CI 90-95, SL 0.79- 0.87 mm, SI 118-124, FL 0.92-0.97 mm, FI 134-144. Worker description Major. - Head in lateral view not or hardly impressed on vertex; head densely covered with short decumbent to subdecumbent hairs entirely; frons with longitudinal rugulae which reach posterolateral corner of vertexal lobes; frontal carina and antennal scrobe absent; clypeus with a median longitudinal carina; hypostoma with an inconspicuous median process and low or relatively developed submedian processes in addition to small lateral processes; antenna with a 3-segmented club; maximal diameter of eye almost as long as or longer than antennal segment X. Promesonotal dome with a much reduced prominence on its posterior slope; humerus not or very weakly produced laterad; the dome at the humeri as broad as or narrower than at the bottom. Petiole as long as postpetiole (excluding helcium); postpetiole massive. First gastral tergite rugoso-punctured at least in its anterior 1/3. Minor. - Dorsum of head largely smooth and shining; preoccipital carina complete but weak dorsally; median part of clypeus almost smooth, usually with a conspicuous to weak median longitudinal carina; antenna with a 3-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye shorter than antennal segment X. Promesonotal dome smooth, in lateral view lacking a conspicuous prominence/mound on its posterior slope; humerus of the dome in dorso-oblique view hardly to weakly produced laterad; propodeal spine small, elongate-triangular. Petiole shorter than postpetiole (excluding helcium); postpetiole massive. Recognition: This species is characterized among Indo-Chinese species by the following characteristics: in the major head densely covered with short decumbent to subdecumbent hairs entirely; in the major frons with longitudinal-oblique rugulae which reach posterolateral corner of vertexal lobes; in the minor promesonotal dome lacking a conspicuous prominence on its posterior slope; in the major and minor postpetiole massive. Pheidole fortis is similar to P. wroughtoni Forel (the type material housed in MHNG was examined), but well distinguished from the latter in which vertex and dorsum of vertexal lobe in lateral view forms an obtuse angle in the major, maximal diameter of eye longer than antennal segment X in the minor, and the posterior slope of promesonotal dome has a conspicuous prominence/mound in the minor. The minor of this species is similar to that of P. magna, but the minor of P. magna has a conspicuous prominence/mound on the posterior slope of promesonotal dome (see also Eguchi 2006). Distribution & bionomics: Known from N. Vietnam and Thailand. This species inhabits open forests and forest edges, and nests in the soil (Eguchi 2006).Published as part of Eguchi, K., 2008, A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)., pp. 1-118 in Zootaxa 1902 on pages 31-3
Pheidole spinicornis Eguchi 2001
Pheidole spinicornis Eguchi, 2001 Pheidole spinicornis Eguchi, 2001: 116-117. Type material examined: BMNH, MBD, MCZC, MSNG, NHMW, UMS; Malaysia, Sabah, Sepilok Forest; K. Eguchi leg. [Eg 98 -BOR- 880 (Accession No. LC 020539: voucher specimen No. Ext 20131029 - 1), type images: CASENT0901620 and CASENT0901621 of AntWeb (http://www.antweb.org/)]. Nontype material examined: ACEG; Malaysia, Sabah, Poring, 500 m alt., Kinabalu Park, H. Hirosawa leg., 20 / vii/ 1997, Poring, 600 m alt., T. Kikuta leg. [6 x 2906 - 5 - Bd], Sayap Kinabalu, ca. 1000 m alt., K. Eguchi leg., 15 /vii/ 1996. Distinction from P. leloi: In the major, promesonotal dome with a pair of spines which slightly or weakly curve backward, with a distinct transverse ridge on its posterior slope; first gastral tergite finely punctate over the surface. In the minor, propodeal spine horn-like, weakly downcurved; petiolar node in lateral view blunt at apex, and in posterior view not emarginate at apex.Published as part of Katsuyuki Eguchi, Bui Tuan Viet, Emiko Oguri & Seiki Yamane, 2016, The first discovery of the " Pheidole quadricuspis group " in the Indo-Chinese Peninsula (Insecta: Hymenoptera: Formicidae: Myrmicinae), pp. 45-55 in Revue suisse de Zoologie 123 (1) on page 53, DOI: 10.5281/zenodo.4628
Pheidole acantha Eguchi 2001
Pheidole acantha Eguchi, 2001 Pheidole acantha Eguchi, 2001: 25-27. Type material examined: BMNH, MCZC, MSNG, NHMW, UMS; Malaysia, Sabah, Taman Kinabalu (KPHQ); K. Eguchi leg. [colony No.: Eg 97 -BOR- 404, type images: CASENT0901618 and CASENT0901619 of AntWeb (http://www.antweb.org/)]. Nontype material examined: ACEG; Malaysia, Sabah, Taman Kinabalu (KPHQ); K. Eguchi leg. [Eg 97 - BOR- 377 (Accession No. LC 020538: voucher specimen No. Ext 20131019 - 10), - 386], Mt. Kinabalu, ca. 1500 m alt., T. Kikuta leg. [7 IV 0714- 1 -B 1; 15 Q 12 S3, 15Q 22 B 5; 118 AC, 177 A, 179 A, 185 A], Mt. Kinabalu, ca. 1800 m alt., T. Kikuta leg. [593 A, 604 A, 607 A, 617 A, 626 A, 664 A, 668 A, 873 A]. Distinction from P. leloi: In the major, promesonotal dome with a conspicuous transverse ridge on its posterior slope; ventral face of petiole without a longitudinal carina. In the minor, dorsum of mesosoma in profile bearing less than 10 standing hairs.Published as part of Katsuyuki Eguchi, Bui Tuan Viet, Emiko Oguri & Seiki Yamane, 2016, The first discovery of the " Pheidole quadricuspis group " in the Indo-Chinese Peninsula (Insecta: Hymenoptera: Formicidae: Myrmicinae), pp. 45-55 in Revue suisse de Zoologie 123 (1) on pages 49-51, DOI: 10.5281/zenodo.4628
Pheidole foveolata Eguchi
Pheidole foveolata Eguchi Figs. 8a-g Pheidole foveolata Eguchi, 2006: 121-123. Holotype: major, "Y Linh Ho (a small fragment of forest), ca. 1100 m alt., Sa Pa, Lao Cai, Vietnam, Eg02-VN-210", IEBR, examined; paratypes: 19 majors, 21 minors & 1 dealate queen, same data as holotype, IEBR, MHNG, MCZC, BMNH, FSKU & ACEG, examined. Pheidole sp. eg-163. Bui & Eguchi 2003: 9 (checklist). Other material examined: Vietnam: Lao Cai: Y Linh Ho (a small fragment of forest), 1100 m alt., Sa Pa [Eg02-VN-220, -227]. Eguchi's informal species code " Pheidole sp. eg-163" has been applied to these specimens. Worker measurements & indices: Major (data from the original description). - HL 0.93-0.97 mm, HW 0.91-0.97 mm, CI 98-101, SL 0.46-0.49 mm, SI 48-54, FL 0.56-0.58 mm, FI 60-62. Minor (data from the original description). - HL 0.47-0.51 mm; HW 0.42-0.46 mm, CI 89-94, SL 0.42- 0.45 mm, SI 98-102, FL 0.42-0.47 mm, FI 100-102. Worker description Major. - Head in lateral view very weakly impressed on vertex; frons longitudinally rugose; vertex and dorsal and lateral faces of vertexal lobe weakly reticulate, with enclosures punctured; frontal carina absent, or present just as rugula(e); antennal scrobe absent; clypeus without a median longitudinal carina; median, submedian and lateral processes of hypostoma conspicuous; antenna with a 3-segmented club; maximal diameter of eye longer than antennal segment X. Promesonotal dome lacking a prominence/mound on its posterior slope; humerus weakly produced laterad; the dome at the humeri as broad as at the bottom, or a little broader at the bottom. Petiole much longer than postpetiole (excluding helcium); postpetiole not massive. First gastral tergite largely smooth and shining except a weakly punctured area around its articulation with postpetiole. Minor. - Head puncutured dorsally and laterally; preoccipital carina absent dorsally; median part of clypeus smooth and shining, usually with a weak or very weak median longitudinal carina; antenna with a3-segmented club; scape exceeding posterior margin of head by 0.5-1.5* length of antennal segment II; maximal diameter of eye a little longer than antennal segment X. Mesosoma punctured well dorsally and laterally; promesonotal dome in lateral view lacking a prominence/mound on its posterior slope; humerus in dorsaloblique view not or hardly produced; propodeal spine much reduced to a tiny dent (at most as long as maximal diameter of propodeal spiracle). Petiole much longer than postpetiole (excluding helcium); postpetiole not massive. Recognition: This species is characterized among Indo-Chinese species by the following characteristics: in the minor dorsal and lateral faces of head and mesosoma punctured; in the minor median part of clypeus smooth and shining; in the major hypostoma in the middle with a conspicuous median process and a pair of conspicuous submedian processes; in the major and minor promesonotal dome lacking a conspicuous prominence /mound on its posterior slope; in the minor propodeal spine much reduced to a small dent. It is similar to Pheidole mus Forel and P. sagei Forel (the type material of both species housed in MHNG was examined) and P parva Mayr (the type material housed in NHMW was examined) but distinguished from the latter three by the following characteristics: propodeal spine is rather developed in the minor of the latter three; the minor of P. mus having median portion of clypeus which is punctured weakly or dimly and not shining. Distribution & bionomics: Known from N. Vietnam. This species inhabits forest edges and nests in the soil (Eguchi 2005). Majors serve as repletes (e.g., the type series).Published as part of Eguchi, K., 2008, A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)., pp. 1-118 in Zootaxa 1902 on pages 34-3
Pheidole vulgaris Eguchi
Pheidole vulgaris Eguchi Figs. 29a-g Pheidole vulgaris Eguchi, 2006: 127-129. Holotype: major, My Yen Commune Forest, 21°35'N, 105°36'E, Na Hau Village, My Yen Commune, Thai Nguyen, Vietnam, Eg01-VN-155, IEBR, examined; paratypes: 19 majors & 20 minors, same data as holotype, IEBR, MHNG, MCZC, BMNH, FSKU & ACEG, examined. Pheidole sp. eg-111. Yamane et al. 2003: 58 (checklist), Bui & Eguchi 2003: 9 (checklist), Eguchi et al. 2004 (ecological study), Eguchi, Bui et al. 2005: 90 (checklist). Other material examined: S. China: Guangxi: Dayaoshan N.R., Jinxiu [J. Fellowes]; Guangdong: Dawuling N.R., Maoming [J. Fellowes]; Da Qiao Town, Ruyuan County[Latella]; Hong Kong: Taipo Kau N.P., New Territories [J. Fellowes]. N. Vietnam: Lao Cai: Y Linh Ho (a small fragment of forest), ca. 1100 m alt., Sa Pa [Eg02-VN-214, -230]; Cat Cat (a trail to Mt. Phansipan), ca. 1300-1400 m alt., Sa Pa [Eg02-VN- 265]; Thai Nguyen: My Yen Commune Forest, 21°35'N, 105°36'E), Na Hau Village [Eg01-VN-155]; Bac Giang: W. Yen Tu N.P. (= Tay Yen Tu N.P.), 21°10-11'N, 106°43-44'E, 170 m alt. [Eg03-VN-079], W. Yen Tu N.P., 210 m alt. [Eg03-VN-063], W. Yen Tu N.P., 370 m alt. [B&E03-52, -56, -57; Eg03-VN-147], W. Yen Tu N.P., 400 m alt. [Eg04-VN-106], W. Yen Tu N.P., 415 m alt. [B&E03-41], W. Yen Tu N.P., 435 m alt. [Eg04-VN-144]; Quang Ninh: Chua Yen Tu, 21°09'N, 106°43'E, 520-725 m alt. [Eg04-VN-035]; Ky Thuong N.R., 21°11'N, 107°07'E, ca. 105 m alt. [B&E03-73], Ky Thuong N.R., 160 m alt. [Eg03-VN-166, -173, - 179, -181], Ky Thuong N.R., 220 m alt. [Eg03-VN-182]; Vinh Phuc: Tam Dao N.P., 21°27'N, 105°38'E, 800- 900 m alt. [Eg99-VN-034], Tam Dao N.P., 900 m alt. [Eg99-VN-002; Eg01-VN-112], Tam Dao N.P., 900- 1100 m alt. [Eg99-VN-043]; Ha Tay (part of specimens mislabeled as "Ha Tai"): Ba Vi N.P., 21°03'N, 105°22'E, 400-600 m alt. [Eg99-VN-085, -089, -093; Eg01-VN-209], Ba Vi N.P., 400-800 m alt. [Eg99-VN- 120], Ba Vi N.P., 600 m alt. [Eg99-VN-103], Ba Vi N.P., 670 m alt. [Eg01-VN-224, -234], Ba Vi N.P., 700 m alt. [Eg02-VN-027], Ba Vi N.P., 800 m alt. [Eg02-VN-048]; Ninh Binh: Cuc Phuong N.P., 20°14'N, 105°36'E, 320 m alt. [Eg01-VN-193, -195]. Thailand: Chiang Mai: Doi Suthep-Pui N.P., 800-900 m alt. [Eg01-TH-079 (W. Jaitrong leg., 1997)]; Nakhonratchasima: Khao Yai N.P., 900-1000 m alt. [TH00-SKY- 34]. India: Utter Pradesh: Rajaji N.P. [A. Schulz & K. Vock, 1996]. Eguchi's informal species code " Pheidole sp. eg-111" has been applied to these specimens. Worker measurements & indices: Major (data from Eguchi 2006). - HL 1.18-1.38 mm; HW 1.06- 1.21 mm; CI 86-91; SL 0.60-0.68 mm; SI 53-59; FI 0.81-0.94 mm; FI 75-81. Minor (data from Eguchi 2006). - HL 0.54-0.61 mm; HW 0.46-0.53 mm; CI 85-91; SL 0.52-0.61 mm; SI 108-117; FL 0.57-0.67 mm; FI 116-127. Worker description Major. - Body covered with relatively long standing hairs. Head in lateral view not or very weakly impressed on vertex; dorsal surface of head variable in sculpture; frons obliquely rugose or largely smooth with sparse interrupted and irregular rugulae; vertex and vertexal lobe largely smooth, or weakly and discontinuously rugose/rugoso-reticulate obliquely toward posterolateral corner of the lobes, often with the interspaces punctured; frontal carina absent or inconspicuous (present just as rugulae); antennal scrobe absent; clypeus sometimes with an evanescent or weak median longitudinal carina; hypostoma with a low or inconspicous median process and a pair of conspicuous submedian processes in addition to a pair of conspicuous lateral processes; outer surface of mandible smooth and shining excluding its basal area, and covered relatively long decumbent hairs; antenna with a 3-segmented club; eye consisting of usually more than 20 ommatidia(but sometimes less than 20 ommatidia); maximal diameter of eye almost as long as, or a little shorter or a little longer than antennal segment X. Promesonotal dome at most with an inconspicuous mound on its posterior slope; humerus of the dome only very weakly produced; the dome usually a little narrower, but sometimes a little broader (as seen in the holotype), at the humeri than at the bottom. Petiole (a little) longer than postpetiole (excluding helcium); postpetiole not massive. First gastral tergite very weakly punctured at least around its articulation with postpetiole, and smooth or sometimes shagreened in the remainder. Minor. - Body yellowish-brown. Frons and vertex smooth and shining; preoccipital carina complete but weak dorsally; clypeus without a median longitudinal carina; antenna with a 3-segmented club; scape exceeding posterior margin of head by the length of second antennal segment or more; eye consisting of 12-18 ommatidia; maximal diameter of eye (a little) shorter than antennal segment X. Promesonotal dome largely smooth and shining, in lateral view without a mound on its posterior slope; humerus of the dome in dorso-oblique view not produced; mesopleuron punctured; metapleuron and lateral face of propodeum weakly punctured or almost smooth; propodeal spine small, elongate-triangular. Petiole (a little) longer than postpetiole (excluding helcium); postpetiole not massive but relatively large, in lateral view somewhat globular. Recognition: This species is characterized among Indo-Chinese species by the combination of the following features: in the major vertex and vertexal lobe largely smooth, or with weak and interrupted rugoso-reticulation directed towards the posterolateral corner of the lobes, often with interspaces punctured; in the minor head and promesonotal dome smooth and shining; in the major hypostoma with conspicuous submedian processes and one very low or vestigial median process; in the minor maximal diameter of eye (a little) shorter than antennal segment X; in the major and minor promesonotal dome lacking a conspicuous prominence/ mound on its posterior slope; in the major and minor petiole longer than postpetiole; in the major and minor postpetiole not massive. This species is very similar to Pheidole taipoana Wheeler and P. laevicolor Eguchi but well distinguished from the latter two by the following characteristics: the major of the latter two having dorsum of vertexal lobe reticulate; the minor of the latter two having maximal diameter of eye a little longer than the length of antennal segment X. This species is also similar to Pheidole vieti (see under P. vieti). Distribution & bionomics: Known from Vietnam, S. China, Thailand and Utter Pradish. This species occurs from forest edges to well-developed forests, and nests in the soil (Eguchi et al. 2004). Colonies are at least occasionally polygynous.Published as part of Eguchi, K., 2008, A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)., pp. 1-118 in Zootaxa 1902 on pages 103-10
Pheidole vulgaris Eguchi
Pheidole vulgaris Eguchi Figs. 29a-g Pheidole vulgaris Eguchi, 2006: 127-129. Holotype: major, My Yen Commune Forest, 21°35'N, 105°36'E, Na Hau Village, My Yen Commune, Thai Nguyen, Vietnam, Eg01-VN-155, IEBR, examined; paratypes: 19 majors & 20 minors, same data as holotype, IEBR, MHNG, MCZC, BMNH, FSKU & ACEG, examined. Pheidole sp. eg-111. Yamane et al. 2003: 58 (checklist), Bui & Eguchi 2003: 9 (checklist), Eguchi et al. 2004 (ecological study), Eguchi, Bui et al. 2005: 90 (checklist). Other material examined: S. China: Guangxi: Dayaoshan N.R., Jinxiu [J. Fellowes]; Guangdong: Dawuling N.R., Maoming [J. Fellowes]; Da Qiao Town, Ruyuan County[Latella]; Hong Kong: Taipo Kau N.P., New Territories [J. Fellowes]. N. Vietnam: Lao Cai: Y Linh Ho (a small fragment of forest), ca. 1100 m alt., Sa Pa [Eg02-VN-214, -230]; Cat Cat (a trail to Mt. Phansipan), ca. 1300-1400 m alt., Sa Pa [Eg02-VN- 265]; Thai Nguyen: My Yen Commune Forest, 21°35'N, 105°36'E), Na Hau Village [Eg01-VN-155]; Bac Giang: W. Yen Tu N.P. (= Tay Yen Tu N.P.), 21°10-11'N, 106°43-44'E, 170 m alt. [Eg03-VN-079], W. Yen Tu N.P., 210 m alt. [Eg03-VN-063], W. Yen Tu N.P., 370 m alt. [B&E03-52, -56, -57; Eg03-VN-147], W. Yen Tu N.P., 400 m alt. [Eg04-VN-106], W. Yen Tu N.P., 415 m alt. [B&E03-41], W. Yen Tu N.P., 435 m alt. [Eg04-VN-144]; Quang Ninh: Chua Yen Tu, 21°09'N, 106°43'E, 520-725 m alt. [Eg04-VN-035]; Ky Thuong N.R., 21°11'N, 107°07'E, ca. 105 m alt. [B&E03-73], Ky Thuong N.R., 160 m alt. [Eg03-VN-166, -173, - 179, -181], Ky Thuong N.R., 220 m alt. [Eg03-VN-182]; Vinh Phuc: Tam Dao N.P., 21°27'N, 105°38'E, 800- 900 m alt. [Eg99-VN-034], Tam Dao N.P., 900 m alt. [Eg99-VN-002; Eg01-VN-112], Tam Dao N.P., 900- 1100 m alt. [Eg99-VN-043]; Ha Tay (part of specimens mislabeled as "Ha Tai"): Ba Vi N.P., 21°03'N, 105°22'E, 400-600 m alt. [Eg99-VN-085, -089, -093; Eg01-VN-209], Ba Vi N.P., 400-800 m alt. [Eg99-VN- 120], Ba Vi N.P., 600 m alt. [Eg99-VN-103], Ba Vi N.P., 670 m alt. [Eg01-VN-224, -234], Ba Vi N.P., 700 m alt. [Eg02-VN-027], Ba Vi N.P., 800 m alt. [Eg02-VN-048]; Ninh Binh: Cuc Phuong N.P., 20°14'N, 105°36'E, 320 m alt. [Eg01-VN-193, -195]. Thailand: Chiang Mai: Doi Suthep-Pui N.P., 800-900 m alt. [Eg01-TH-079 (W. Jaitrong leg., 1997)]; Nakhonratchasima: Khao Yai N.P., 900-1000 m alt. [TH00-SKY- 34]. India: Utter Pradesh: Rajaji N.P. [A. Schulz & K. Vock, 1996]. Eguchi's informal species code " Pheidole sp. eg-111" has been applied to these specimens. Worker measurements & indices: Major (data from Eguchi 2006). - HL 1.18-1.38 mm; HW 1.06- 1.21 mm; CI 86-91; SL 0.60-0.68 mm; SI 53-59; FI 0.81-0.94 mm; FI 75-81. Minor (data from Eguchi 2006). - HL 0.54-0.61 mm; HW 0.46-0.53 mm; CI 85-91; SL 0.52-0.61 mm; SI 108-117; FL 0.57-0.67 mm; FI 116-127. Worker description Major. - Body covered with relatively long standing hairs. Head in lateral view not or very weakly impressed on vertex; dorsal surface of head variable in sculpture; frons obliquely rugose or largely smooth with sparse interrupted and irregular rugulae; vertex and vertexal lobe largely smooth, or weakly and discontinuously rugose/rugoso-reticulate obliquely toward posterolateral corner of the lobes, often with the interspaces punctured; frontal carina absent or inconspicuous (present just as rugulae); antennal scrobe absent; clypeus sometimes with an evanescent or weak median longitudinal carina; hypostoma with a low or inconspicous median process and a pair of conspicuous submedian processes in addition to a pair of conspicuous lateral processes; outer surface of mandible smooth and shining excluding its basal area, and covered relatively long decumbent hairs; antenna with a 3-segmented club; eye consisting of usually more than 20 ommatidia(but sometimes less than 20 ommatidia); maximal diameter of eye almost as long as, or a little shorter or a little longer than antennal segment X. Promesonotal dome at most with an inconspicuous mound on its posterior slope; humerus of the dome only very weakly produced; the dome usually a little narrower, but sometimes a little broader (as seen in the holotype), at the humeri than at the bottom. Petiole (a little) longer than postpetiole (excluding helcium); postpetiole not massive. First gastral tergite very weakly punctured at least around its articulation with postpetiole, and smooth or sometimes shagreened in the remainder. Minor. - Body yellowish-brown. Frons and vertex smooth and shining; preoccipital carina complete but weak dorsally; clypeus without a median longitudinal carina; antenna with a 3-segmented club; scape exceeding posterior margin of head by the length of second antennal segment or more; eye consisting of 12-18 ommatidia; maximal diameter of eye (a little) shorter than antennal segment X. Promesonotal dome largely smooth and shining, in lateral view without a mound on its posterior slope; humerus of the dome in dorsooblique view not produced; mesopleuron punctured; metapleuron and lateral face of propodeum weakly punctured or almost smooth; propodeal spine small, elongate-triangular. Petiole (a little) longer than postpetiole (excluding helcium); postpetiole not massive but relatively large, in lateral view somewhat globular. Recognition: This species is characterized among Indo-Chinese species by the combination of the following features: in the major vertex and vertexal lobe largely smooth, or with weak and interrupted rugoso-reticulation directed towards the posterolateral corner of the lobes, often with interspaces punctured; in the minor head and promesonotal dome smooth and shining; in the major hypostoma with conspicuous submedian processes and one very low or vestigial median process; in the minor maximal diameter of eye (a little) shorter than antennal segment X; in the major and minor promesonotal dome lacking a conspicuous prominence/ mound on its posterior slope; in the major and minor petiole longer than postpetiole; in the major and minor postpetiole not massive. This species is very similar to Pheidole taipoana Wheeler and P. laevicolor Eguchi but well distinguished from the latter two by the following characteristics: the major of the latter two having dorsum of vertexal lobe reticulate; the minor of the latter two having maximal diameter of eye a little longer than the length of antennal segment X. This species is also similar to Pheidole vieti (see under P. vieti). Distribution & bionomics: Known from Vietnam, S. China, Thailand and Utter Pradish. This species occurs from forest edges to well-developed forests, and nests in the soil (Eguchi et al. 2004). Colonies are at least occasionally polygynous.Published as part of Eguchi, K., 2008, A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)., pp. 1-118 in Zootaxa 1902 on pages 103-10
Pheidole elongicephala Eguchi, 2008, sp.n.
Pheidole elongicephala Eguchi sp.n. Figs. 4a-h Pheidole sp. B (aff. aglae Forel). Yamane et. al. 2003: 58 (checklist). Pheidole aglae Forel. Misidentification made by Eguchi et al. 2004 (ecological study). Type material examined: Holotype, major, Cuc Phuong N.P. ("Dong Nguoi Xua" Area), Ninh Binh, N. Vietnam [Eg09vi05-08 (K. Eguchi leg., 09/JUN/2005)] (IEBR); paratypes: 7 majors, 9 minors & 2 males, same data as holotype (IEBR, MHNG, MCZC, & ACKE). Other material examined: S. China: Hong Kong: Taipo Kau N.P., New Territory [Eg00-HK-028]. Vietnam: Thai Nguyen: My Yen Commune Forest, 21°35'N, 105°36'E, Na Hau Village [Eg01-VN-158]; Bac Giang: W. Yen Tu N.P. (=Tay Yen Tu N.P.), 21°10-11'N, 106°43-44'E, 190-370 m alt. [Eg03-VN-030, -040, -135, -153, -160; B&E03-12]; Ha Tay (mislabeled as Ha Tai): Ba Vi N.P., 21°03'N, 105°22'E, 400-700 m alt. [Eg02-VN-009, -022]; Ninh Binh: Cuc Phuong N.P. [Eg11iv05-14; Eg15vi05-07]. Part of specimens to which Eguchi's informal species code " Pheidole sp. eg-101" has been applied (Eguchi, Bui et al. 2005: 90) is P. elongicephala, and the remainder is P. ochracea. Worker measurements & indices: Holotype (major). - HL 1.76 mm; HW 1.56 mm; CI 89; SL 1.18 mm; SI 76; FL 1.56 mm; FI 100. Nontype major (n=4). - HL1.68-1.78 mm; HW 1.51-1.57 mm; CI 88-90; SL 1.13-1.16 mm; SI 74-75; FL 1.53-1.56 mm; FI 99-102. Minor (n=5, including one paratype minor). - HL 0.78-0.85 mm; HW 0.58-0.64 mm; CI 75-76; SL 1.06-1.19 mm; SI 176-187; FL 1.11-1.25 mm; FI 187-197. Worker description Major. - Body deep yellowish-brown, brown or deep reddish-brown, with paler appendages. Head in lateral view not or weakly impressed on vertex; vertexal lobes in full-face view relatively close to each other; frons and anterior part of vertex longitudinally rugose; posterior part of vertex and dorsum of vertexal lobe rugoso-reticulate or reticulate; clypeus without a conspicuous median longitudinal carina; frontal carina conspicuous; antennal scrobe very shallow; hypostoma with a very low or inconspicuous median process and a pair of low or inconspicuous submedian processes in addition to a pair of conspicuous lateral processes; antenna with a 3-segmented club; maximal diameter of eye shorter than antennal segment X. Promesonotal dome sparsely rugose transversely, with interspaces smooth and shining; a conspicuous prominence present on its posterior slope; humerus not or hardly produced laterad; the dome at the humeri much narrower than at the bottom; mesopleuron and metapleuron weakly punctured, overlain by rugoso-reticulation; propodeal spine narrowly based. Petiole longer than postpetiole (excluding helcium); postpetiole not massive. First gastral tergite smooth entirely, or shagreened only around its articulation with postpetiole. Minor. - Body yellowish-brown or brown, with paler appendages. Head in full-face view elongate-elliptical, tapered posterad behind eyes; frons and vertex smooth or shagreened; median portion of clypeus smooth; median longitudinal carina very weak or absent; occipital carina forming a well-developed collar; antenna with a 3-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye much shorter than antennal segment X; 8-9 ommatidia present on the long axis of eye; promesonotal dome smooth and shining, in lateral view with a low mound on its gentle posterior slope; the mound bearing usually 4 standing hairs; humerus in dorso-oblique view not raised/produced; mesopleuron, metapleuron and lateral face of propodeum punctured, often overlain by weak rugulae; propodeal spine elongate-triangular, directing upward; petiole (a little) longer than postpetiole (excluding helcium); postpetiole not massive. Recognition: Pheidole elongicephala,P. binghamii and P. ochracea sp.n. are morphologically very similar to each other (and probably very closely related to each other). Differences between P. elongicephala and P. binghamii are as follows: head behind eyes in full-face view more elongated and tapered posterad in the minor of the former than in that of the latter; mound on the posterior slope of promesonotal dome bearing usually4 standing hairs in the minor of the former, but usually only a pair of standing hairs in the minor of the latter. The body size of both the subcastes is almost constantly larger in P. elongicephala than in P. binghamii. The two species are allopatric or parapatric (range of P. binghamii: Myanmar, Thailand, S. Vietnam). Differences between P. elongicephala and P. ochracea sp.n. are as follows: head behind eyes in full-face view more elongated and tapered posterad in the minor of the former than in that of the latter; occipital carina of the minor much more developed in the former than in the latter; the long axis of eye having 8-9 ommatidia in the minor of the former, but 6-7 ommatidia in that of the latter. They are sympatric in S. China and N. Vietnam. Distribution & bionomics: Known from N. Vietnam and S. China. This species occurs in woody habitats, and nests in rotting wood material (logs, stubs) and in the soil.Published as part of Eguchi, K., 2008, A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)., pp. 1-118 in Zootaxa 1902 on pages 20-2
Pheidole elongicephala Eguchi, 2008, sp.n.
Pheidole elongicephala Eguchi sp.n. Figs. 4a-h Pheidole sp. B (aff. aglae Forel). Yamane et. al. 2003: 58 (checklist). Pheidole aglae Forel. Misidentification made by Eguchi et al. 2004 (ecological study). Type material examined: Holotype, major, Cuc Phuong N.P. ("Dong Nguoi Xua" Area), Ninh Binh, N. Vietnam [Eg09vi05-08 (K. Eguchi leg., 09/JUN/2005)] (IEBR); paratypes: 7 majors, 9 minors & 2 males, same data as holotype (IEBR, MHNG, MCZC, & ACKE). Other material examined: S. China: Hong Kong: Taipo Kau N.P., New Territory [Eg00-HK-028]. Vietnam: Thai Nguyen: My Yen Commune Forest, 21°35'N, 105°36'E, Na Hau Village [Eg01-VN-158]; Bac Giang: W. Yen Tu N.P. (=Tay Yen Tu N.P.), 21°10-11'N, 106°43-44'E, 190-370 m alt. [Eg03-VN-030, -040, -135, -153, -160; B&E03-12]; Ha Tay (mislabeled as Ha Tai): Ba Vi N.P., 21°03'N, 105°22'E, 400-700 m alt. [Eg02-VN-009, -022]; Ninh Binh: Cuc Phuong N.P. [Eg11iv05-14; Eg15vi05-07]. Part of specimens to which Eguchi's informal species code " Pheidole sp. eg-101" has been applied (Eguchi, Bui et al. 2005: 90) is P. elongicephala, and the remainder is P. ochracea. Worker measurements & indices: Holotype (major). - HL 1.76 mm; HW 1.56 mm; CI 89; SL 1.18 mm; SI 76; FL 1.56 mm; FI 100. Nontype major (n=4). - HL1.68-1.78 mm; HW 1.51-1.57 mm; CI 88-90; SL 1.13-1.16 mm; SI 74-75; FL 1.53-1.56 mm; FI 99-102. Minor (n=5, including one paratype minor). - HL 0.78-0.85 mm; HW 0.58-0.64 mm; CI 75-76; SL 1.06-1.19 mm; SI 176-187; FL 1.11-1.25 mm; FI 187-197. Worker description Major. - Body deep yellowish-brown, brown or deep reddish-brown, with paler appendages. Head in lateral view not or weakly impressed on vertex; vertexal lobes in full-face view relatively close to each other; frons and anterior part of vertex longitudinally rugose; posterior part of vertex and dorsum of vertexal lobe rugoso-reticulate or reticulate; clypeus without a conspicuous median longitudinal carina; frontal carina conspicuous; antennal scrobe very shallow; hypostoma with a very low or inconspicuous median process and a pair of low or inconspicuous submedian processes in addition to a pair of conspicuous lateral processes; antenna with a 3-segmented club; maximal diameter of eye shorter than antennal segment X. Promesonotal dome sparsely rugose transversely, with interspaces smooth and shining; a conspicuous prominence present on its posterior slope; humerus not or hardly produced laterad; the dome at the humeri much narrower than at the bottom; mesopleuron and metapleuron weakly punctured, overlain by rugoso-reticulation; propodeal spine narrowly based. Petiole longer than postpetiole (excluding helcium); postpetiole not massive. First gastral tergite smooth entirely, or shagreened only around its articulation with postpetiole. Minor. - Body yellowish-brown or brown, with paler appendages. Head in full-face view elongate-elliptical, tapered posterad behind eyes; frons and vertex smooth or shagreened; median portion of clypeus smooth; median longitudinal carina very weak or absent; occipital carina forming a well-developed collar; antenna with a 3-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye much shorter than antennal segment X; 8-9 ommatidia present on the long axis of eye; promesonotal dome smooth and shining, in lateral view with a low mound on its gentle posterior slope; the mound bearing usually 4 standing hairs; humerus in dorso-oblique view not raised/produced; mesopleuron, metapleuron and lateral face of propodeum punctured, often overlain by weak rugulae; propodeal spine elongate-triangular, directing upward; petiole (a little) longer than postpetiole (excluding helcium); postpetiole not massive. Recognition: Pheidole elongicephala, P. binghamii and P. ochracea sp.n. are morphologically very similar to each other (and probably very closely related to each other). Differences between P. elongicephala and P. binghamii are as follows: head behind eyes in full-face view more elongated and tapered posterad in the minor of the former than in that of the latter; mound on the posterior slope of premesonotal dome bearing usually4 standing hairs in the minor of the former, but usually only a pair of standing hairs in the minor of the latter. The body size of both the subcastes is almost constantly larger in P. elongicephala than in P. binghamii. The two species are allopatric or parapatric (range of P. binghamii: Myanmar, Thailand, S. Vietnam). Differences between P. elongicephala and P. ochracea sp.n. are as follows: head behind eyes in full-face view more elongated and tapered posterad in the minor of the former than in that of the latter; occipital carina of the minor much more developed in the former than in the latter; the long axis of eye having 8-9 ommatidia in the minor of the former, but 6-7 ommatidia in that of the latter. They are sympatric in S. China and N. Vietnam. Distribution & bionomics: Known from N. Vietnam and S. China. This species occurs in woody habitats, and nests in rotting wood material (logs, stubs) and in the soil.Published as part of Eguchi, K., 2008, A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)., pp. 1-118 in Zootaxa 1902 on pages 20-2
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