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    Cloeodes opacus Nieto & Richard

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    Cloeodes opacus Nieto & Richard Cloeodes opacus Nieto & Richard, 2008: 13 Diagnosis. Cloeodes opacus can be distinguished from the other species of the genus by the following combination of characters: 1) hind wing pads present; 2) segment III of labial palpi subquadrangular (Nieto & Richard, 2008: Fig. 46); 3) dorsal margin of femora with a row of blunt spines, apically with a subquadrangular projection and two blunt spines(Nieto & Richard, 2008: Fig. 48); 4) abdominal color pattern dark brown with segments I–IX with medial yellow markings as in Nieto & Richard, 2008: Fig. 43, segment X brown; 5) caudal filaments with flattened setae basally sclerotized, cerci with long spines toward external margin every two segments, terminal filament with long spines toward midline dorsally and ventrally every two segments. Material. One nymph: URUGUAY, Rivera, Mina de los Corrales, Arroyo Corrales, S 31 º 34´34 ´´, W 55 º 27´58 ´´, 135m, 19 / 2 / 2008, Emmerich & Molineri colls. Two nymphs: Treinta Tres, R 7, km 279, Arroyo La Yeguada, S 32 º 57´34 ´´, W 54 º 59´11 ´´, 232m, 21 / 2 / 2008, Emmerich & Molineri colls. One nymph: Rivera, Santa Ernestina, R. 29, cerca Mina Corrales, Arroyo sin nombre, S 31 º 32 ' 23.8 " W 55 º 33 ' 42 ", 146m, 19 / 2 / 2008, D. Emmerich & C. Molineri colls. Twenty-two nymphs: Rocha, Tahona, Finca Sr. Fernández, Cascada de los Helechos, S 34 º 20 ’ 49.5 ’’ W 54 º 25 ’ 40.8 ’’, 122m, 23 / 3 / 2010, D. Emmerich & C. Molineri colls. Seventeen nymphs: Treinta y Tres, Quebrada de los Cuervos, Arroyo Yerbal Chico, S 32 º 55 ’ 35.2 ’’, W 54 º 27 ’ 39.6 ’’, 78 m, 25 / 3 / 10, D. Emmerich & C. Molineri colls.Published as part of Nieto, Carolina & Emmerich, Daniel, 2011, Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay, pp. 57-65 in Zootaxa 2996 on page 63, DOI: 10.5281/zenodo.20543

    Econ 2016, 10o. Congreso Internacional de Economía y Gestión : expositor Daniel Ricardo Nieto Michel

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    Conferencia “Hacia la reconstrucción de las Estadísticas Públicas”, disertantes Jorge Todesca (Director INDEC); Daniel Ricardo Nieto Michel (Dirección Nacional de Estadísticas de Condiciones de Vida - INDEC

    Econ 2016, 10o. Congreso Internacional de Economía y Gestión : expositor Daniel Ricardo Nieto Michel

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    Conferencia “Hacia la reconstrucción de las Estadísticas Públicas”, disertantes Jorge Todesca (Director INDEC); Daniel Ricardo Nieto Michel (Dirección Nacional de Estadísticas de Condiciones de Vida - INDEC

    Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay

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    Nieto, Carolina, Emmerich, Daniel (2011): Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay. Zootaxa 2996: 57-65, DOI: 10.5281/zenodo.20543

    FIGURES 1–3. Nymphs, dorsal habitus. 1 in Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay

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    FIGURES 1–3. Nymphs, dorsal habitus. 1, Cloeodes dialutoi sp. nov. 2, Cloeodes guenoa sp. nov. 3, Cloeodes vaimaca sp. nov.Published as part of Nieto, Carolina & Emmerich, Daniel, 2011, Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay, pp. 57-65 in Zootaxa 2996 on page 59, DOI: 10.5281/zenodo.20543

    Cloeodes guenoa Nieto & Emmerich, 2011, sp. nov.

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    Cloeodes guenoa sp. nov. (Figs. 2, 15– 25) Nymph (Fig. 2). Length: body, 4.9–5.1 mm; cerci, 1.4–1.5 mm; terminal filament, 1.2–1.3 mm. Antennae broken. Head yellowish brown, longer than wide, compound eyes yellowish brown. Antennae pale yellow. Mouthparts (Figs. 15–20): labrum (Fig. 15 a) with one subapical seta centrally and one seta near lateral margin, anterior margin with bifid setae (Fig. 15 b). Left mandible (Fig. 16) without setae between prostheca and mola, thumb of molar area transverse to anterior margin. Right mandible (Fig. 17) without setae between prostheca and mola. Hypopharynx (Fig. 18): lingua with rounded projection and longer than superlinguae. Maxillae (Fig. 19), palpi as long as galealacinia, two segmented, segment II with a constriction. Labium (Fig. 20) with segment III of palpi rounded. Thorax pale yellow. Pleura yellowish brown, sterna pale yellow. Legs yellowish brown. Dorsal margin of femora with a row of pointed spines, apically with a subquadrangular projection and two blunt spines (Fig. 21). Tarsal claws 0.3 times the length of tarsi. Hind wing pads present. Abdomen yellowish brown (Fig. 2), segments III-VII with two whitish spots near midline, segments IV and VIII pale brown. Abdominal terga with covers, posterior margin of terga with spines as in Fig. 22. Paraprocts with 10–11 spines apically (Fig. 23). Gills translucent white, main and secondary trachea pigmented (Fig. 24), 2 times the length of each tergum. Caudal filament yellowish, with flattened setae basally sclerotized (similar to Fig. 14 b). Cerci with long spines toward external margin every two segments, terminal filament with long spines toward midline every two segments (Fig. 25). Adults. Unknown. Etymology. “ Guenoa ” is the name of the native people who inhabits the current districts of Treinta y Tres, Cerro Largo and Tacuarembó. Diagnosis. Cloeodes guenoa sp. nov. can be distinguished from the other species of the genus by the following combination of characters: 1) hind wing pads present; 2) lingua with rounded projection and longer than superlinguae (Fig. 18); 3) labium with segment III of palpi rounded (Fig. 20); 4) dorsal margin of femora with a row of pointed spines, apically with a subquadrangular projection and two blunt spines (Fig. 21); 5) abdominal color pattern as in Fig. 2, with segments III-VII with two whitish spots near midline, segments IV and VIII pale brown. Material. Holotype female mature nymph, URUGUAY, Tacuarembó, R 5, km 379, Arroyo Tranqueras, S 31 º 47´29 ´´, W 55 º 58´51 ´´, 124m, 22 / 2 / 2008, Emmerich & Molineri colls. Paratypes six nymphs (one dissected) same data as holotype. Three nymphs: Lavalleja, R 8, km 190, Arroyo Tapes Chico, S 33 º 59´18 ´´, W 54 º 46´54 ´´, 130m, 4 / 3 / 2008, Emmerich & Pérez colls. One nymph: Treinta y Tres, Quebrada de los Cuervos, Arroyo Los Helechos, S 32 º 55 ’ 27.3 ’’, W 54 º 27 ’ 34.4 ’’, 125 m. 8 / 11 / 2004, D. Emmerich & E. Morelli colls. Six nymphs: Maldonado, R 109, Cerro Cordillera, Arroyo sin nombre, S 34 º 20 ’ 29.4 ’’, W 54 º 37 ’ 40.5 ’’, 301 m, 23 / 3 / 10, D. Emmerich & C. Molineri colls.Published as part of Nieto, Carolina & Emmerich, Daniel, 2011, Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay, pp. 57-65 in Zootaxa 2996 on pages 61-63, DOI: 10.5281/zenodo.20543

    Paracloeodes morellii Emmerich & Nieto, 2009, sp. nov.

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    Paracloeodes morellii sp. nov. (Figs. 13–25) Nymph. Length: body: 2.8–3.3 mm; cerci 1.0– 1.3 mm. Antennae 0.7–0.8 mm, 1.5 times the head capsule. General color brownish orange. Head orange brown, with small brownish spots as in Figs. 13–14. Frontal keel present, ocelli brownish. Antennae: scape and pedicel pale yellow, flagellum pale. Mouthparts (Figs. 15–20): labrum (Figs. 15 a–b) dorsally with 2–3 subapical setae near midline, anterior margin with bifid setae. Mandibles (Figs. 16–17) without setae between prostheca and mola, incisors cleft in two sets, inner set at right angle with outer set. Left mandible (Fig. 16) with molars with constrictions, prostheca broad with 1–2 denticles. Right mandible (Fig. 17) with prostheca bifid. Hypopharynx (Fig. 18) with lingua subequal to superlinguae and rounded apically. Maxillae (Fig. 19) with palpi two segmented, 1.5 times the length of galealacinia. Labium (Fig. 20) with segment II of palpi with rounded projection, 2 times width of segment III, segment III conical. Thorax (Fig. 13) orange brown. Pronotum with a pair of brownish spots in the midline. Mesonotum shadowed with brown and with small brownish spots. Metanotum brownish. Pleura reddish brown, sterna whitish. Legs (Fig. 21) pale yellow. Tarsal claws 0.4 times the length of tarsi, with 2 rows of minute denticles (Fig. 22). Hind wing pads absent. Female (Fig. 14) paler than male and with a brown band in metanotum and anterior portion of fore wing pads. Abdomen orange brown. Color pattern: segment II–VII with medial elongate spot as in Fig. 13, segment IX brownish. Posterior margin of terga with spines as long as wide (Fig. 23). Sterna pale yellow, sternum IX brownish except at lateral sides. Gills whitish, translucent, oblong, 1.5 times the length of each tergum. Paraprocts (Fig. 24) with 8–10 spines apically. Terminal filaments pale yellow, with crown of small spines on each segment (Fig. 25). Adults. Unknown. Etymology. This species is dedicated to Dr. Enrique Morelli, entomologist of Universidad de la República (Uruguay) who continually stimulates the study of the aquatic insects of Uruguay. Discussion. This species can be distinguished from the other species of the genus by the following combination of characters: 1) frontal keel present; 2) antennae short, 1.5 times the head capsule; 3) segment II of labial palpi with rounded projection, 2 times width of segment III (Fig. 20); 4) tarsal claws with two rows of minute denticles (Fig. 22); 5) hind wing pads absent; 6) general color pattern with head and thorax with small spots, abdomen with segment II–VII with medial elongate spot as in Fig. 13; 7) posterior margin of terga with spines as long as wide (Fig. 23); 8) paraprocts with 8–10 spines apically (Fig. 24); 9) terminal filament with crown of small spines on each segment (Fig. 25). Material. Holotype, male nymph: URUGUAY, Deptos. Rivera-Tacuarembó, R 44, km 26.5, A° Carpintería, S 31°48´28´´, W 55°13´09´´, 121 m, 20/02/2008, Emmerich & Molineri col. Paratypes: 5 nymphs same data (mouthparts, legs and paraprocts of 1 nymph mounted on slide); 1 nymph (parts on slide Depto. Durazno, Sarandí del Yi, Río Yi, S 33°20´41´´, W 55°37´07´´, 132 m, 21/02/2008, same collectors; 2 nymphs (1 on slide): Depto. Cerro Largo, ruta 8, km 335, A° Otazo, S 32°50´00´´, W 54°14´55´´, 123 m, 20/02/2008, Same collectors. The holotype and 1 paratype female are deposited in the Museo de la Universidad de la República, Montevideo, Uruguay (MUR); remaining paratypes are housed in Instituto Miguel Lillo, Tucumán, Argentina (IML).Published as part of Emmerich, Daniel & Nieto, Carolina, 2009, First record of the genus Paracloeodes Day (Ephemeroptera: Baetidae) in Uruguay and description of two new species, pp. 57-62 in Zootaxa 2225 (1) on pages 60-62, DOI: 10.11646/zootaxa.2225.1.3, http://zenodo.org/record/532027

    Paracloeodes charrua Emmerich & Nieto, 2009, sp. nov.

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    Paracloeodes charrua sp. nov. (Figs. 1–12) Nymph. Length: body: 3.8–4.3 mm; cerci 1.8 mm. Antennae long, 2.1 mm, 3.5 times width of the head capsule. General color yellowish brown. Head yellowish brown with a characteristic pattern (Fig. 1). Frontal keel present, ocelli black. Antennae pale yellow. Mouthparts (Figs. 2–7): labrum (Figs. 2 a–b) dorsally with a pair of subapical setae near midline, anterior margin with bifid setae. Mandibles (Figs. 3–4) without setae between prostheca and mola, incisors cleft in two sets, inner set at right angle with outer set. Left mandible (Fig. 3) with molars with constrictions, prostheca with two denticles. Right mandible (Fig. 4) with prostheca slender and bifid. Hypopharynx with lingua shorter than superlinguae and rounded apically. Maxillae (Fig. 6) with palpi two-segmented, twice the length of galea-lacinia. Labium (Fig. 7) with segment II of palpi with rounded projection, 2 times width of segment III, segment III conical. Thorax yellowish brown, shadowed with brown (Fig. 1). Pleura brownish, sterna pale yellow. Legs (Fig. 8) pale yellow, femora with brownish transverse band apically (more conspicuous in female). Tarsal claws (Fig. 9) 0.5 times the length of tarsi, with 2 rows of small denticles basally and 4–5 denticles increasing in size apically. Hind wing pads present. Abdomen yellowish brown (Fig. 1). Color pattern: segment II–IX with 3 brownish spots forming a triangle, segment V and IX darker than others. Posterior margin of terga with spines 2 times longer than wide (Fig. 10). Sterna yellowish brown, sternum IX darker than others. Gills whitish, elongate, less than twice the length of each tergum, main branch of trachea pigmented. Paraprocts (Fig. 11) with 10–12 spines apically. Terminal filaments with crown of spines on each segment, cerci with long spines toward the external margin every four segments (Fig. 12). Adults. Unknown. Etymology. Charrua: ethnic group that inhabited Uruguay and was exterminated in the middle of the 19th Century. We dedicate this species to their memory. Discussion. This species can be distinguished from the other species of the genus by the following combination of characters: 1) frontal keel present; 2) antennae long, 3.5 times width of head capsule; 3) maxillae with palpi twice the length of galea-lacinia (Fig. 6); 4) segment II of labial palpi with rounded projection, 2 times width of segment III (Fig. 7); 5) tarsal claws 0.5 times the length of tarsi with two rows of small denticles basally and 4–5 denticles increasing in size apically (Figs. 8–9); 6) hind wing pads present; 7) abdomen with a characteristic color pattern: segment II–IX with 3 brownish spots forming a triangle (Fig. 1); 8) posterior margin of terga with spines twice longer than wide; 9) paraprocts with 10–12 spines apically; 10) cerci with long spines toward the external margin every four segments (Fig. 12). Material. Holotype, female nymph: URUGUAY, Depto. Cerro Largo, Arroyo Otazo, ruta 8, km 335, S 32°50´00´´, W 54°14´55´´, 123 m. 20/02/2008, Emmerich & Molineri cols. Paratypes: 4 nymphs same data (mouthparts, legs and paraprocts of one nymph mounted on slides); 4 nymphs: Depto. Rivera, Tranqueras, ruta 30, km 251, Río Tacuarembó, S 31°10´40´´, W 55°45´44´´, 147 m, 19/02/2008, same collectors; 5 nymphs: Depto. Tacuarembó, ruta 5, km 379, A° Tranqueras, S 31°47´29´´, W 55°58´51´´, 124 m, 22/02/ 2008, same collectors; 3 nymphs: Dpto Soriano, ruta 21, km 339, Aº Las Maulas, S 33º 23’ 32’’, W 58º 07’ 02’’, 16 m, 6/03/08, Emmerich & Pérez cols.; 11 nymphs: Dpto Paysandú, “Predio”, Aº Grande, S 32º 39’ 34.2’’, W 57º 14’ 56.7’’, 9/08/07, Morelli col. The holotype is deposited in the Museo de la Universidad de la República, Montevideo Uruguay (MUR), paratypes are deposited in Instituto Miguel Lillo, Tucumán, Argentina (IML).Published as part of Emmerich, Daniel & Nieto, Carolina, 2009, First record of the genus Paracloeodes Day (Ephemeroptera: Baetidae) in Uruguay and description of two new species, pp. 57-62 in Zootaxa 2225 (1) on pages 58-60, DOI: 10.11646/zootaxa.2225.1.3, http://zenodo.org/record/532027

    Cloeodes vaimaca Nieto & Emmerich, 2011, sp. nov.

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    <i>Cloeodes vaimaca</i> sp. nov. <p>(Figs. 3, 26–35)</p> <p>Nymph (Fig. 3). Length: body, 4.1–4.2 mm; cerci, 1.4–1.5 mm; terminal filament, 1.3–1.4 mm. Antennae partially broken. Head pale yellow, longer than wide, compound eyes pale yellow. Antennae pale yellow. Mouthparts (Figs. 26–31): labrum (Fig. 26 a) with one subapical seta centrally and two-three setae near lateral margin, anterior margin with bifid setae (Fig. 26 b). Left mandible (Fig. 27) without setae between prostheca and mola, thumb of molar area transverse to anterior margin. Right mandible (Fig. 28) without setae between prostheca and mola. Hypopharynx (Fig. 29): lingua with rounded projection and longer than superlinguae. Maxillae (Fig. 30), palpi scarcely shorter than galea-lacinia, two segmented, segment II with a constriction. Labium (Fig. 31) with segment III of palpi rounded.</p> <p>Thorax pale yellow. Pleura pale yellow, sterna whitish. Legs pale yellow. Dorsal edge of femora with a row of pointed spines, apically with a rounded projection and two blunt spines (Fig. 32). Tarsal claws 0.4 times the length of tarsi. Hind wing pads present.</p> <p>Abdomen pale yellow, segments I–V with a reddish spot in the midline. Abdominal terga with covers, posterior margin of terga with spines as in Fig. 33. Sterna pale yellow with trachea pigmented, sterna II-VI with a tuft of long fine setae laterally on each side. Paraprocts with spines apically (Fig. 34). Gills (Fig. 35) translucent, trachea pigmented, 2.5 times the length of each tergum. Caudal filaments pale yellow, with flattened setae basally sclerotized (similar to Fig. 14 b). Cerci with long spines toward external margin every two segments, terminal filament with long spines toward midline every two segments (similar to Fig. 35).</p> <p> <b>Etymology.</b> Vaimaca was one of the four charrúas taken to Paris in 1833 to be exhibited in the "Museum of the Man ". This fact crowned the humiliation and destruction of this people for the colonial Spanish model. We dedicate respectfully this species to his memory.</p> <p> <b>Diagnosis.</b> <i>Cloeodes vaimaca</i> <b>sp. nov.</b> can be distinguished from the other species of the genus by the following combination of characters: 1) hind wing pads present; 2) lingua with rounded projection and longer than superlinguae (Fig. 29); 3) labium (Fig. 31) with segment III of palpi rounded; 3) dorsal margin of femora with a row of pointed spines, apically with a rounded projection and two blunt spines (Fig. 32); 4) abdominal color pattern as in Fig. 3, with segments I-V with a reddish spot in the midline</p> <p> <b>Material.</b> Holotype male nymph: URUGUAY, Flores, R. 57, km 30, Arroyo Guardia Vieja, S 33º 42´12´´, W 57º 06´17´´, 120m, 6/ 3/ 2008, Emmerich & Pérez colls. Paratypes: three nymphs (one dissected) same data as holotype. One nymph: Paysandú, R. 3, km 366, Arroyo Sacra, S 32º 21´02´´, W 58º 03´04´´, 10m, 7/ 3/ 2008, Emmerich & Pérez colls. One nymph: Río Negro, R 3, Km 291, Camping. Arroyo San Esteban, S 32° 50’ 07’’, W 57º 30’ 59’’, 34 m, 2/ 3/ 0 8, D. Emmerich & S. Pérez colls.</p>Published as part of <i>Nieto, Carolina & Emmerich, Daniel, 2011, Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay, pp. 57-65 in Zootaxa 2996</i> on pages 63-65, DOI: <a href="http://zenodo.org/record/205436">10.5281/zenodo.205436</a&gt

    FIGURES 13–25 in First record of the genus Paracloeodes Day (Ephemeroptera: Baetidae) in Uruguay and description of two new species

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    FIGURES 13–25. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 15–20): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus.Published as part of Emmerich, Daniel & Nieto, Carolina, 2009, First record of the genus Paracloeodes Day (Ephemeroptera: Baetidae) in Uruguay and description of two new species, pp. 57-62 in Zootaxa 2225 (1) on page 61, DOI: 10.11646/zootaxa.2225.1.3, http://zenodo.org/record/532027
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