147 research outputs found

    Neoarctidae de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia 1992

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    1.5 NEOARCTIDAE de Zio Grimaldi, D’Addabbo Gallo & Morone De Lucia, 1992 (emended by Bello & Grimaldi de Zio 1998). Considered as subfamily Neoarctinae assigned to the family Stygarctidae by de Zio Grimaldi et al. (1992) was elevated to family level by Bello & Grimaldi de Zio (1998). Arthrotardigrada with a complete set of cephalic cirri and three pairs of clavae. Body covered with five thin plates: A cephalic plate, a scapular plate (segmental plate I) which bears two long cirrophores with the primary clavae and cirri A, two other plates (segmental plates II and III), and the caudal plate (segmental plate IV). Median cirrus, internal and external cirri, with the secondary and tertiary clavae, located on the cephalic plate. Stylet supports absent. Legs non digitate, lacking true telescopy. Three claws on all legs attached to the tarsus with a dorsal basal membrane. Claws of different length: two longer, on the middle of the tarsus, and a short lateral one. A very short and thin dorsal bristle32 on the two medial claws. Sense organs on every leg. Cirrus E with a coarse accordion-shaped articulation. 1.5.1 Neoarctus de Zio Grimaldi, D’Addabbo Gallo & Morone De Lucia, 1992 (Figs 9 D, 10G) Generic diagnosis as that of the family. Type species: Neoarctus primigenius de Zio Grimaldi, D’Addabbo Gallo & Morone De Lucia, 1992 Selected literature for further information: de Zio Grimaldi et al. (1992).Published as part of Fontoura, Paulo, Bartels, Paul J., Jørgensen, Aslak, Kristensen, Reinhardt Møbjerg & Hansen, Jesper Guldberg, 2017, A dichotomous key to the genera of the Marine Heterotardigrades (Tardigrada), pp. 1-45 in Zootaxa 4294 (1) on page 31, DOI: 10.11646/zootaxa.4294.1.1, http://zenodo.org/record/83154

    Neostygarctus acanthophorus Grimaldi

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    154. Neostygarctus acanthophorus Grimaldi de Zio, D'Addabbo Gallo & Morone De Lucia, 1982 a Neostygarctus acanthophorus n. gen., n. sp. (Grimaldi de Zio et al. 1982 a) Neostygarctus acanthophorus Grimaldi de Zio et al. 1982 (Grimaldi de Zio et al. 1983 a) Neostygarctus acanthophorus, Grimaldi de Zio et alii 1982 (Grimaldi de Zio et al. 1983 b) Neostygarctus acanthophorus Grimaldi de Zio et al., 1982 (D’Addabbo Gallo et al. 1987, Grimaldi de Zio & D’Addabbo Gallo 1987, Grimaldi de Zio & D’Addabbo Gallo 2001, Gallo et al. 2007) Neostygarctus acanthophorus (Accogli et al. 2009) Terra typica: Ionian Sea (Italy, Europe) Adriatic Sea: • 40 ° 42 ′ 49 ′′N, 17 ° 47 ′ 58 ′′E; 20 m bsl: [FAO 37] Italy, Brindisi Province, Torre Guateco [Riserva Naturale Torre Guateco], sand of Posidonia seagrass meadow. Grimaldi de Zio et al. (1990) • 40 ° 12 ′ 18 ′′N, 18 ° 27 ′ 28 ′′E; 20 m bsl: [FAO 37] Italy, Lecce Province, near Otranto, sand of Posidonia seagrass meadow. Grimaldi de Zio et al. (1990) Indian Ocean: • 0 4 ° 15 ′N, 72 ° 49 ′E; 37 m bsl: [FAO 51] Republic of the Maldives, Ari Nord Atoll, coarse sand. Gallo et al. (2007) • 0 3 ° 57 ′N, 73 ° 25 ′E; 5–41 m bsl: [FAO 51] Republic of the Maldives, Male Sud Atoll, coarse sand. Gallo et al. (2007) Ionian Sea: • 40 ° 29 ′00′′N, 17 °03′ 25 ′′E; 35 m bsl: [FAO 37] Italy, Tarento Province, Armeleia Shoal, coarse coralligenous detritus. Grimaldi de Zio et al. (1990) • 40 ° 29 ′00′′N, 17 °03′ 25 ′′E; 15 m bsl: [FAO 37] Type locality: Italy, Taranto Province, coast of Taranto, offshore of Chiatona, Secca dell’Armeleia, subtidal, organogenic, Amphioxus debris. Grimaldi de Zio et al. (1982 a, 1983 a,b) • 40 °09′N, 17 ° 58 ′E; 0-20 m bsl: [FAO 37] Italy, Lecce Province, Nard, Corvine cave, coarse sand. Accogli et al. (2009) • 40 °08′ 44 ′′N, 17 ° 58 ′05′′E; 12 m bsl: [FAO 37] Italy, Lecce Province, Nard, Grotta Cattedrale, subtidal, detritus. Grimaldi de Zio et al. (1982 a, 1983 b) • 39 ° 52 ′00′′N, 16 ° 42 ′ 30 ′′E; 20–200 m bsl: [FAO 37] Italy, Cosenza Province, High Ionian Sea offshore, Calabrian Coast, Amendolara Shoal, coralligenous sand, coralligenous sand mixed with debris, coarse medium or fine coralligenous sand (few samples). D’Addabbo Gallo et al. (1987), Grimaldi de Zio & D’Addabbo Gallo (1987), Grimaldi de Zio et al. (2003) Tyrrhenian Sea: • 40 ° 20 ′ 30 ′′N, 09° 40 ′ 15 ′′E; 21–40 m bsl: [FAO 37] Italy, Sardinia Island, Nuoro Province, Orosei Gulf, sand in Posidonia seagrass meadow and coralligenous detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 40 °09′00′′N, 09° 39 ′ 19 ′′E; 20 m bsl: [FAO 37] Italy, Sardinia Island, Nuoro Province, Orosei Gulf, coarse detritus. Grimaldi de Zio et al. (1990) • 38 ° 36 ′N, 14 ° 55 ′E; 40 m bsl: [FAO 37] Italy, Aeolian Islands, Secca del Capo, coralligenous bank, subtidal zone, coarse coralligenous detritus. Grimaldi de Zio & D’Addabbo Gallo (2001) Record numbers (Sea/Ocean classification): Adriatic Sea: 2, Indian Ocean: 2, Ionian Sea: 5, Tyrrhenian Sea: 3; total: 12. Record numbers (FAO classification): FAO 37: 10, FAO 51: 2; total: 12. Remarks: This is a subtidal species (most often from 15 to 40 m bsl) known mainly from the Mediterranean basin, but also reported from tropical localities in the Indian Ocean. It has been collected from different substrates such as coralligenous detritus, Amphioxus debris, and sand in Posidonia seagrass meadows.Published as part of Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena & Nelson, Diane R., 2015, The Zoogeography of Marine Tardigrada, pp. 1-189 in Zootaxa 4037 (1) on pages 133-134, DOI: 10.11646/zootaxa.4037.1.1, http://zenodo.org/record/23351

    Parastygarctus renaudae Grimaldi

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    159. Parastygarctus renaudae Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia & Daddabbo, 1987 Parastygarctus renaudae n. sp. (Grimaldi de Zio et al. 1987) Parastygarctus renaudae Grimaldi de Zio et al., 1987 (Gallo et al. 2007) Terra typica: Indian Ocean (Republic of Mauritius, Africa)Published as part of Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena & Nelson, Diane R., 2015, The Zoogeography of Marine Tardigrada, pp. 1-189 in Zootaxa 4037 (1) on page 136, DOI: 10.11646/zootaxa.4037.1.1, http://zenodo.org/record/23351

    Halechiniscus macrocephalus Grimaldi

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    78. Halechiniscus macrocephalus Grimaldi de Zio, D’Addabbo Gallo & Morone De Lucia, 1988 Halechiniscus macrocephalus n. sp. (Grimaldi de Zio et al. 1988) Halechiniscus macrocephalus Grimaldi de Zio et al., 1988 (D’Addabbo Gallo et al. 2001, Grimaldi de Zio & D’Addabbo Gallo 2001, Accogli et al. 2011) Terra typica: Ionian Sea (Italy, Europe) Adriatic Sea: • 42 °06 ′ N, 15 ° 29 ′E; 9 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Viole, ca. 10 m from the cave’s entrance, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) Ionian Sea: • 39 ° 52 ′ 27 ′′N, 16 ° 43 ′ 25 ′′E; 80–100 m bsl: [FAO 37] Type Locality: Italy, Cosenza Province, High Ionian Sea, Amendolara Shoal, coralligenous detritus with mud. Grimaldi de Zio et al. (1988) • 39 ° 51.26 ′N, 18 ° 23.55 ′E; 20 m bsl: [FAO 37] Italy, Lecce Province, Gagliano, coarse sand. Accogli et al. (2011) • 39 ° 51 ′ 20 ′′N, 18 ° 23 ′ 59 ′′E; 0 m bsl: [FAO 37] Italy, Lecce Province, Gagliano del Capo. Grimaldi de Zio et al. (2003) Tyrrhenian Sea: • 40 ° 20 ′ 30 ′′N, 09° 40 ′ 15 ′′E; 0–60 m bsl: [FAO 37] Italy, Sardinia Island, Nuoro Province, Orosei Gulf, sand and coralligenous detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 38 ° 36 ′N, 14 ° 55 ′E; 40 m bsl: [FAO 37] Italy, Aeolian Islands, Secca del Capo, coralligenous bank, subtidal zone, coarse coralligenous detritus. Grimaldi de Zio & D’Addabbo Gallo (2001) Record numbers (Sea/Ocean classification): Adriatic Sea: 1, Ionian Sea: 3, Tyrrhenian Sea: 2; total: 6. Record numbers (FAO classification): FAO 37: 6; total: 6. Remarks: This is a rare Mediterranean species reported mainly from the subtidal zone (including a marine cave) down to ca. 100 m bsl, from sand and coralligenous and organogenic detritus.Published as part of Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena & Nelson, Diane R., 2015, The Zoogeography of Marine Tardigrada, pp. 1-189 in Zootaxa 4037 (1) on pages 77-78, DOI: 10.11646/zootaxa.4037.1.1, http://zenodo.org/record/23351

    Batillipes adriaticus Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo & Grimaldi 1979

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    Batillipes adriaticus Grimaldi de Zio, Morone De Lucia, D’Addabbo Gallo & Grimaldi, 1979 Material examined PORTUGAL: 1 ♀ collected at Portinho da Arrábida Beach, mounted in PVA (slide C. IX- 71), and 14 specimens (9 ♀♀, 1 ♂, 3 juveniles and 1 four-toed larva, all mounted in glycerol), collected at Vasco da Gama Beach (slides C. IX- 75, C. IX- 80 – C. IX- 91). Remarks The specimens clearly correspond to the original description of Grimaldi de Zio et al. (1979). The main characteristic of this species is its dorsal cuticle sculpture constituted by large pillars, appearing as large tubercles (diameter about 2–3 µm); ventrally the cuticle exhibits the fine punctuation as do the majority of Batillipes species. Cephalic appendages have a lance-like tip and papillary secondary clavae are present. Between legs III and IV, an acute sculptured conical body projection is present. Lateral body projections between the first three pairs of legs are not referred to by Grimaldi de Zio et al. (1979). However, in some Portuguese specimens, small blunt ventro-lateral body projections are visible. The caudal appendage is a long spine inserted on a cylindrical base. Middle toes (3 and 4) on feet of legs IV are of similar length. Sensory organs are present on all legs, considerably long on legs IV. The specific characters of the species: dorsal sculpture, conical lateral body projection between legs III–IV and spine-like caudal appendage are already present in the four-toed larva. Distribution Up to now, this species had only been reported from the Mediterranean Basin (Adriatic Sea, Ionian Sea and Tyrrhenian Sea, see Kaczmarek et al. 2015). This is its first record for the Atlantic Ocean. Associated species Batillipes phreaticus and H. greveni.Published as part of Santos, Erika, Rubal, Marcos, Veiga, Puri, da Rocha, Clélia M. C. & Fontoura, Paulo, 2018, Batillipes (Tardigrada, Arthrotardigrada) from the Portuguese coast with the description of two new species and a new dichotomous key for all species, pp. 1-32 in European Journal of Taxonomy 425 on pages 4-5, DOI: 10.5852/ejt.2018.425, http://zenodo.org/record/380613

    Halechiniscus chafarinensis Grimaldi

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    74. Halechiniscus chafarinensis Grimaldi de Zio & Villora-Moreno, 1995 Halechiniscus chafarinensis n. sp. (Grimaldi de Zio & Villora-Moreno 1995) Halechiniscus chafarinensis Grimaldi de Zio & Villora Moreno, 1995 (D’Addabbo Gallo et al. 2001, Grimaldi de Zio & D’Addabbo Gallo 2001) Halechiniscus chafarinensis (Sandulli et al. 1999) Halechiniscus chafarinensis de Zio Grimaldi & Villora Moreno, 1995 (Accogli et al. 2011) Terra typica: Alboran Sea (Morocco, Africa) Adriatic Sea: • 42 °06′N, 15 ° 28 ′E [42 °08′N, 15 ° 30 ′E]; 5–20 m bsl: [FAO 37] Italy, Foggia Province, between Tremiti Islands, Posidonia seagrass meadow, medium and fine sand. Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °09′00′′N, 15 ° 30 ′00′′E [42 °08′N, 15 ° 31 ′E]; 5–38 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, Caprara [Capraia] Island, Cala dei Turchi, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °07′N, 15 ° 31 ′E [42 °08′N, 15 ° 31 ′E]; 5–20 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Nicola, Punta del Cimitero, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °06′N, 15 ° 29 ′E [42 °07′N, 15 ° 29 ′E]; 5–20 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Sale Cave, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °08′N, 15 ° 29 ′E [42 °07′N, 15 ° 30 ′E]; 5–20 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, I Pagliai. coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °07′N, 15 ° 30 ′E; 5–20 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Cala Spino, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °06 ′ N, 15 ° 28 ′E; 3 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Murene, cave’s entrance, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °06 ′ N, 15 ° 28 ′E; 3 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Murene, the middle of the cave, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °06 ′ N, 15 ° 28 ′E; 2 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Murene, the end of the cave, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °06′N, 15 ° 28 ′E; 2–4 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Punta Zio Cesare, cave Grotta delle Murene, 0–10 cm depth, 80 % coarse organogenic sand, 3 % silt and clay, (mean diameter = 0.125 mm). Sandulli et al. (1999), Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °06 ′ N, 15 ° 29 ′E; 9 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Viole, ca. 10 m from the cave’s entrance, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °06 ′ N, 15 ° 29 ′E; 5 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Viole, internal opening, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °06′N, 15 ° 29 ′E; 5–20 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Punta del Diavolo, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 40 ° 34.38 ′N, 18 ° 0 9.49 ′E ; 10 m bsl: [FAO 37] Italy, Brindisi Province, Mattarelle [Torre Mattarelle], ENEL power station, coarse sand. Accogli et al. (2011) • 40 ° 34 ′ 34 ′′N, 18 °02′ 90 ′′E [40 ° 34 ′N, 18 °03′E]; 0 m bsl: [FAO 37] Italy, Brindisi Province, Mattarelle [Torre Mattarelle]. Grimaldi de Zio et al. (2003) Alboran Sea: • 35 ° 11 ′N, 02° 26 ′W; 4.5 m bsl: Type Locality: [FAO 37] Morocco, Chafarinas Islands, Isable II Island [Isla Isabel II], shallow sublittoral pool between Posidonia seagrass meadows, basaltic coarse sand. Grimaldi de Zio & Villora-Moreno (1995) Ionian Sea: • 40 ° 12.94 ′N, 17 ° 5.07 ′E [40 ° 13 ′N, 17 ° 55 ′E]; 20 m bsl: [FAO 37] Italy, Lecce Province, San Isidoro, medium sand. Accogli et al. (2011) • 39 ° 51.26 ′N, 18 ° 23.55 ′E; 20 m bsl: [FAO 37] Italy, Lecce Province, Gagliano, coarse sand. Accogli et al. (2011) • 39 ° 51.26 ′N, 18 ° 23.55 ′E; 50 m bsl: [FAO 37] Italy, Lecce Province, Gagliano, fine sand. Accogli et al. (2011) • 39 ° 51 ′ 20 ′′N, 18 ° 23 ′ 59 ′′E; 0 m bsl: [FAO 37] Italy, Lecce Province, Gagliano del Capo. Grimaldi de Zio et al. (2003) • 39 ° 50.14 ′N, 18 °06.45′E ; 20 m bsl: [FAO 37] Italy, Lecce Province, Torre Mozza (industry for fish processing), coarse sand. Accogli et al. (2011) Mediterranean Sea: • 35 ° 54 ′N, 14 ° 32 ′E; 41 m bsl: [FAO 37] Republic of Malta, SE coast of Malta, Sicily Channel, 1.75 km SE of the entrance of Grand Harbour and 1.4 km NW of the coastal settlement Xghajra, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001) Record numbers (Sea/Ocean classification): Adriatic Sea: 15, Alboran Sea: 1, Ionian Sea: 5, Mediterranean Sea: 1; total: 22. Record numbers (FAO classification): FAO 37: 22; total: 22. Remarks: This is a Mediterranean species reported mostly from the subtidal zone (to ca. 40 m bsl). Although H. chafarinensis was described from the Alboran Sea (coast of Chafarinas Islands), it was subsequently reported mainly from the Adriatic Sea in the subtidal zone and marine caves around the Tremiti Islands. It has been collected from a range of sand and detritus.Published as part of Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena & Nelson, Diane R., 2015, The Zoogeography of Marine Tardigrada, pp. 1-189 in Zootaxa 4037 (1) on pages 70-72, DOI: 10.11646/zootaxa.4037.1.1, http://zenodo.org/record/23351

    Archechiniscus minutus Grimaldi

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    3. Archechiniscus minutus Grimaldi de Zio & D'Addabbo Gallo, 1987 Archechiniscus minutus n. sp. (Grimaldi de Zio & D’Addabbo Gallo 1987) A. minutus Grimaldi de Zio & D'Addabbo Gallo, 1987 (D’Addabbo Gallo et al. 1987) Terra typica: Ionian Sea (Italy, Europe) Adriatic Sea: • 40 ° 43 ′N, 17 ° 48 ′ 30 ′′E; 20–34 m bsl: [FAO 37] Italy, Brindisi Province, Torre Guateco [Riserva Naturale Torre Guateco], coarse coralligenous debris. Grimaldi de Zio & D’Addabbo Gallo (1987) • 40 °08′N, 18 ° 32 ′E; 9–15 m bsl: [FAO 37] Italy, Lecce Province, near Otranto, coarse coralligenous debris. Grimaldi de Zio & D’Addabbo Gallo (1987) • 40 °08′N, 18 ° 32 ′E; 4–10 m bsl: [FAO 37] Italy, Lecce Province, near Otranto, eel-grass meadow, sand. Grimaldi de Zio & D’Addabbo Gallo (1987) Indian Ocean: • 03° 57 ′N, 73 ° 25 ′E; 5–41 m bsl: [FAO 51] Republic of the Maldives, Male Sud Atoll, coarse sand. Gallo et al. (2007) Ionian Sea: • 39 ° 51 ′N, 18 °07′E; 9–15 m bsl: [FAO 37] Italy, Lecce Province, Ugento [near Fontanelle and Torre Mozza], eel-grass meadow, sand. Grimaldi de Zio & D’Addabbo Gallo (1987) • 39 ° 52 ′00′′N, 16 ° 42 ′ 30 ′′E; 20–35 m bsl: [FAO 37] Type Locality: Italy, Cosenza Province, High Ionian Sea offshore, Calabrian Coast, Amendolara Shoal, coralligenous, coralligenous mixed with debris, coarse medium or fine coralligenous. D’Addabbo Gallo et al. (1987), Grimaldi de Zio & D’Addabbo Gallo (1987), Grimaldi de Zio et al. (2003) Tyrrhenian Sea: • 40 ° 20 ′ 30 ′′N, 09° 40 ′ 15 ′′E; 0–80 <m bsl: [FAO 37] Italy, Sardinia Island, Nuoro Province, Orosei Gulf, pure sand, coastal organogenic debris, coralligenous debris mud or Posidonia seagrass meadows. D’Addabbo Gallo et al. (1989), Grimaldi de Zio & D’Addabbo Gallo (2001) • 38 ° 36 ′N, 14 ° 55 ′E; 40 m bsl: [FAO 37] Italy, Aeolian Islands, Secca del Capo, coralligenous bank, subtidal zone, coarse coralligenous detritus. Grimaldi de Zio & D’Addabbo Gallo (2001) Record numbers (Sea/Ocean classification): Adriatic Sea: 3, Indian Ocean: 1, Ionian Sea: 2, Tyrrhenian Sea: 2; total: 8. Record numbers (FAO classification): FAO 37: 7, FAO 51: 1; total: 8. Remarks: This species is known mainly from the Mediterranean basin with a single record in the Indian Ocean (Maldives). It is an intertidal or subtidal species collected from different types of sand from 0 to 80 m bsl (most often ca. 20 m bsl).Published as part of Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena & Nelson, Diane R., 2015, The Zoogeography of Marine Tardigrada, pp. 1-189 in Zootaxa 4037 (1) on page 9, DOI: 10.11646/zootaxa.4037.1.1, http://zenodo.org/record/23351

    Styraconyx sardiniae D'Addabbo Gallo, Morone De Lucia & Grimaldi

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    116. Styraconyx sardiniae D’Addabbo Gallo, Morone De Lucia & Grimaldi de Zio, 1989 Styraconyx sardiniae new species (D’Addabbo Gallo et al. 1989) Styraconyx sardiniae D’Addabbo Gallo et al., 1989 (D’Addabbo Gallo et al. 1999 a, 2001, Grimaldi de Zio & D’Addabbo Gallo 2001, D’Addabbo Gallo et al. 2007, Gallo et al. 2007) Styraconyx sardiniae (Sandulli et al. 1999) Styraconyx sardiniae D’Addabbo Gallo et al., 1988 (Accogli et al. 2011) Terra typica: Tyrrhenian Sea (Italy, Europe) Adriatic Sea: • 42 °06′N, 15 ° 28 ′E [42 °08′N, 15 ° 30 ′E]; 11–20 m bsl: [FAO 37] Italy, Foggia Province, between Tremiti Islands, Posidonia seagrass meadow, medium and fine sand. Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °09′00′′N, 15 ° 30 ′00′′E [42 °08′N, 15 ° 31 ′E]; 5–56 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, Caprara [Capraia] Island, Cala dei Turchi, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °08′00′′N, 15 ° 31 ′00′′E; 5–56 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, Caprara [Capraia] Island, Cala Sorrentino, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °07′ 18 ′′N, 15 ° 29 ′ 49 ′′E; 0 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Cala delle Arene. Grimaldi de Zio et al. (2003) • 42 °08′N, 15 ° 29 ′E [42 °07′N, 15 ° 30 ′E]; 5–20 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, I Pagliai, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °07′N, 15 ° 30 ′E; 5–20 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Cala Spino, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °07′N, 15 ° 31 ′E; 5–20 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Nicola, Ferraio Cave, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °06 ′ N, 15 ° 28 ′E; 3 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Murene, the middle of the cave, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °06′N, 15 ° 28 ′E; 2–4 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Punta Zio Cesare, cave Grotta delle Murene, 0–10 cm depth, 80 % coarse organogenic sand, 3 % silt and clay, (mean diameter = 0.125 mm). Sandulli et al. (1999), Grimaldi de Zio & D’Addabbo Gallo (2001) • 42 °06 ′ N, 15 ° 29 ′E; 10 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Viole, external cave’s entrance, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °06 ′ N, 15 ° 29 ′E; 7 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, southern area of San Domino Island, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Viole, ca. 20 m from the cave’s entrance, coarse organogenic detritus. D’Addabbo Gallo et al. (2001), Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 42 °06′N, 15 ° 29 ′E; 5–10 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Punta Zio Cesare, cave Grotta delle Viole, 0–10 cm depth, 80 % coarse organogenic sand, 3 % silt and clay, (mean diameter = 0.125 mm). Sandulli et al. (1999), Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 40 ° 31.59 ′N, 18 °05. 51 ′E; 20 m bsl: [FAO 37] Italy, Brindisi Province, Lendinuso, coarse sand in Posidonia meadows. Accogli et al. (2011) Balearic Sea: • 40 ° 20 ′N, 08° 22 ′E; 20 m bsl: [FAO 37] Italy, Sardinia, Sassari or Oristano Province, Gulf of Alghero, medium sand. D’Addabbo Gallo et al. (2007) Indian Ocean: • 04° 19 ′S, 55 ° 52 ′E; 0 m bsl: [FAO 51] Republic of Seychelles, Coco [Cocos] Island, coarse sand. Gallo et al. (2007) Ionian Sea: • 39 ° 55 ′ 55 ′′N, 18 °02′ 71 ′′E [39 ° 54 ′N, 18 °06′E]; 0 m bsl: [FAO 37] Italy, Lecce Province, San Giovanni [Torre San Giovanni]. Grimaldi de Zio et al. (2003) • 39 ° 50.46 ′N, 18 °07.38′E ; 20 m bsl: [FAO 37] Italy, Lecce Province, Torre Mozza (industry for fish processing), sand, coarse organogenous debris. Accogli et al. (2011) Ligurian Sea: • 43 ° 32 ′ N, 10 ° 12 ′E; 7 m bsl: [FAO 37] Italy, Livorno Province, 300 m NW of Meloria Shoals Lighthouse, sediment samples. D’Addabbo Gallo et al. (1999 a) Mediterranean Sea: • 35 ° 54 ′N, 14 ° 32 ′E; 41 m bsl: [FAO 37] Republic of Malta, SE coast of Malta, Sicily Channel, 1.75 km SE of the entrance of Grand Harbour and 1.4 km NW of the coastal settlement Xghajra, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001) Tyrrhenian Sea: • 40 ° 20 ′ 30 ′′N, 09° 40 ′ 15 ′′E; 0–80 m bsl: [FAO 37] Type Locality: Italy, Sardinia Island, Nuoro Province, Orosei Gulf, organogenic debris and Posidonia seagrass meadow. D’Addabbo Gallo et al. (1989), Grimaldi de Zio & D’Addabbo Gallo (2001) • 40 ° 13 ′N, 09° 38 ′E; 30 m bsl: [FAO 37] Italy, Sardinia, Nuoro Province, Gulf of Orosei, coarse sand. D’Addabbo Gallo et al. (2007) • 38 ° 36 ′N, 14 ° 55 ′E; 40 and 46 m bsl: [FAO 37] Italy, Aeolian Islands, Secca del Capo, coralligenous bank, subtidal zone, coarse coralligenous detritus and medium carbonate sand. Grimaldi de Zio & D’Addabbo Gallo (2001) • 38 ° 34 ′N, 14 ° 52 ′E [38 ° 35 ′N, 14 ° 52 ′E]; 1–5 and 40 m bsl: [FAO 37] Italy, Aeolian Islands, Salina Island, Punta di Magne, subtidal zone, siliceous sand. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 38 ° 26 ′N, 14 ° 55 ′E [38 ° 26 ′N, 14 ° 57 ′E]; 1.5–5: [FAO 37] Italy, Aeolian Islands, Lipari Island, Pietra Lunga South, subtidal zone, siliceous sand. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) Record numbers (Sea/Ocean classification): Adriatic Sea: 13, Balearic Sea: 1, Indian Ocean: 1, Ionian Sea: 2, Ligurian Sea: 1, Mediterranean Sea: 1, Tyrrhenian Sea: 5; total: 24. Record numbers (FAO classification): FAO 37: 23, FAO 51: 1; total: 24. Remarks: This is a Mediterranean species from the sargassi group reported almost exclusively from the Mediterranean basin, except for one relatively recent report from the Indian Ocean. It has been found in the intertidal and subtidal zones, down to 80 m bsl, in different types of sand and sediments (also present in marine caves).Published as part of Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena & Nelson, Diane R., 2015, The Zoogeography of Marine Tardigrada, pp. 1-189 in Zootaxa 4037 (1) on pages 108-110, DOI: 10.11646/zootaxa.4037.1.1, http://zenodo.org/record/23351

    Actinarctus physophorus Grimaldi

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    128. Actinarctus physophorus Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia, Vaccarella & Grimaldi, 1982 a sensu lato Actinarctus physophorus sp. n. (Grimaldi de Zio et al. 1982 a) A. physophorus, Grimaldi de Zio et alii 1982 (Grimaldi de Zio et al. 1983 b) Actinarctus physophorus Grimaldi de Zio et al., 1982 (D’Addabbo Gallo et al. 1987, Grimaldi de Zio & D’Addabbo Gallo 1987, Villora-Moreno & Grimaldi de Zio 1996, D’Addabbo Gallo et al. 2007) Actinarctus cf. physophorus Grimaldi de Zio, 1982 (Kristensen & Neuhaus 1999) Actinarctus cf. physophorus Grimaldi de Zio, 1984 (Hansen et al. 2001) Terra typica: Tyrrhenian Sea (Italy, Europe) Atlantic Ocean: • 61 ° 17 ′ 42 ′′N, 08° 32 ′ 18 ′′W; 104–260 m bsl: [FAO 27] Faroe Islands, Faroe Bank, shell gravel, coarse shell gravel, fine carbonate sand, carbonate sand, or fine shell sand. Hansen et al. (2001) • 61 °05.61′N, 08° 35.57 ′W / 61 °06′N, 08° 36 ′W; 103.3 m bsl: [FAO 27] Faroe Islands, Faroe Bank, BIOFAR station 787. Kristensen & Neuhaus (1999) Alboran Sea: • 35 ° 11 ′N, 02° 26 ′W; 0 m bsl: [FAO 37] Morocco, Chafarinas Islands, Congreso Island [Isla del Congres], sublittoral mixed environment, Amphioxus gravel (2 sites). Villora-Moreno & Grimaldi de Zio (1996) • Balearic Sea: • 40 ° 20 ′N, 08° 22 ′E; 20 m bsl: [FAO 37] Italy, Sardinia, Sassari or Oristano Province, Gulf of Alghero, medium sand. D’Addabbo Gallo et al. (2007) Ionian Sea: • 39 ° 52 ′00′′N, 16 ° 42 ′ 30 ′′E; 20–200 m bsl: [FAO 37] Italy, Cosenza Province, High Ionian Sea offshore, Calabrian Coast, Amendolara Shoal, coralligenous sand, coralligenous sand mixed with debris, coarse medium or fine coralligenous sand (few samples). D’Addabbo Gallo et al. (1987), Grimaldi de Zio & D’Addabbo Gallo (1987), Grimaldi de Zio et al. (2003) Tyrrhenian Sea: • 40 ° 35 ′ 10 ′′N, 14 ° 19 ′ 38 ′′E; 12 m bsl: [FAO 37] Type Locality: Italy, Province of Naples, Sorrento Peninsula, Massa Lubrense, Cala di Mitigliano, cave (Grotta di Mitigliano), subtidal, organic detritus and 33 % gravel. Grimaldi de Zio et al. (1982 a, 1983 b) Record numbers (Sea/Ocean classification): Atlantic Ocean: 2, Alboran Sea: 1, Balearic Sea: 1, Ionian Sea: 1, Tyrrhenian Sea: 1; total: 6. Record numbers (FAO classification): FAO 27: 2, FAO 37: 4; total: 6. Remarks: This intertidal-subtidal species is known mainly from the Mediterranean region although it has also been reported from the deep subtidal zone in the Atlantic Ocean. However, the deep water specimens of both Kristensen & Neuhaus (1999) and Hansen et al. (2001) were reported as A. cf. physophorus, with the suggestion that these specimens from the Faroe Bank (Atlantic Ocean) could belong to a new species. This is an interstitial species found mainly in sand or gravel.Published as part of Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena & Nelson, Diane R., 2015, The Zoogeography of Marine Tardigrada, pp. 1-189 in Zootaxa 4037 (1) on pages 120-121, DOI: 10.11646/zootaxa.4037.1.1, http://zenodo.org/record/23351
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