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Ditaxiporinae Stach 1935
No full textSubfamily Ditaxiporinae Stach, 1935 Diagnosis. Colony erect, jointed, comprising sterile internodes that are mostly multizooidal and biserial, rarely unizooidal in part or all of the colony; fertile internodes always multizooidal and biserial. Frontal shield varied; multi-costate, or mostly with vestigial suboral costae so that there is an extensive gymnocyst, typically foraminate; or the gymnocyst reduced owing to expansion of pore-chambers frontally so that most of the shield is effectively cryptocystal. Ovicells likewise variable, ectooecium well-developed, smooth, with a suture or foramina, or ectooecium reduced, exposing extensive granular endooecium. Dorsal surface of zooids smooth or with pore-chambers variably developed. Avicularia typically lateral-oral and mostly with complete cross-bar. Remarks. Stach (1933, 1934, 1935) subdivided the Catenicellidae into several subfamilies, basing them primarily on the position of the ovicell within the segment. He named five subfamilies — Vittaticellinae Stach, 1933, Scuticellinae Stach, 1934, Catenicellinae Stach, 1935, Cornuticellinae Stach, 1935, and Ditaxiporinae Stach, 1935. Additionally, Cheetham (1963) introduced a new family-rank taxon, Ditaxiporinidae. Gordon & Braga (1994) commented on these arrangements, synonymising some. They regarded biseriality (i.e., multizooidal) as having more fundamental significance within the family than the position of the ovicell (or the morphology of the frontal shield), which they interpret as having differentiated within different lineages later. For these reasons they subsumed the Ditaxiporinidae into the Ditaxiporinae.Published as part of Vieira, Leandro M., Gordon, Dennis P. & Correia, Monica D., 2007, First record of a living ditaxiporine catenicellid in the Atlantic, with a description of Vasignyella ovicellata n. sp. (Bryozoa), pp. 49-58 in Zootaxa 1582 (1) on page 51, DOI: 10.11646/zootaxa.1582.1.5, http://zenodo.org/record/509762
Deuterosminthurus bicinctus sensu Stach 1956
No full textDeuterosminthurus bicinctus (Koch, 1840) sensu Stach 1956 * Sminthurus bicinctus Koch, 1840: 353. Bourletiella (Deuterosminthurus) flava Gisin, 1946a: 254. Deuterosminthurus bicinctus – Stach 1956: 171. DISTRIBUTION. — Essonne. Forêt de Sénart et parc du Laboratoire du MNHNà Brunoy (Vannier 1970, D. flavus; Ponge 1973, 1980, 1993; Betsch 1991). Gers. Vers Cravencères (Nayrolles 1996). Haute-Garonne. Toulouse (Cassagnau 1962a). Haute-Saône. Denis 1924d. Hautes-Pyrénées. Valléed’Aure (Cassagnau 1964a, c, D. flavus). Meurthe-et-Moselle. Picard 1962. Tarn, Aude. Montagne Noire (Cassagnau 1965). Tarn. Entre Lacabarède et Rouairoux, 440 m (Nayrolles 1996). TAXONOMIE. — Nayrolles 1996 (redescription). ÉCOLOGIE. — Picard 1962 (faune psammique); Cassagnau 1964c, 1965 (écologie, sol); Vannier 1970 (eau, sol); Ponge 1973, 1980, 1993; Betsch 1991 (privation delitière).Published as part of Thibaud, Jean-Marc, 2017, Catalogue des collemboles de France, pp. 297-436 in Zoosystema 39 (3) on page 400, DOI: 10.5252/z2017n3a1, http://zenodo.org/record/549627
Disparrhopalites Stach 1956
No full text<i>Disparrhopalites</i> Stach, 1956 <p> <i>Disparrhopalites</i> Stach, 1956:63.</p> <p> <i>Dietersminthurus</i> Palacios-Vargas, Cuéllar & Vázquez, 1998:13, <b>new synonym</b>.</p> <p> <b>Type species:</b> <i>Disparrhopalites patrizii</i> (Cassagnau & Delamare Deboutteville, 1953) <b>Other</b> <b>species:</b></p> <p> <i>D. enkerlinius</i> (Palacios-Vargas, Cuellar & Vazquez, 1998) <b>n. comb.</b> <i>D. naasaveqw</i> <b>n. sp.</b></p> <p> <i>D. tergestinus</i> Fanciulli, Colla & Dallai, 2005</p> <p> <i>Pararrhopalites</i> <i>patrizii</i> Cassagnau & Delamare Deboutteville, 1953 was transferred to the new genus <i>Disparrhopalites</i> by Stach (1956). This widespread European species (Dallai 1971, Fanciulli <i>et al.</i> 2005) has been collected and studied thoroughly since then (Christian 1998; Dallai 1970, 1971; Delamare Deboutteville & Bassot 1957; Fanciulli et al. 2005; Gama 1988, 2005; Marx & Weber 2013, 2015). However, <i>Disparrhopalites</i> was not compared to more recently described but clearly similar genera (<i>Dietersminthurus, Gisinurus, Songhaica, Soqotrasminthurus, Varelasminthurus</i>, see Tables 1 & 2 for authorities and dates), perhaps because the two known <i>Disparrhopalites</i> spp. lack a prominent cavity-like formation on the unguis. The type species, <i>D. patrizii</i>, possibly a weak troglophile, has a shallow linear cavity while the highly modified troglobiont <i>D. tergestinus</i> apparently lacks the cavity completely. <i>Disparrhopalites naasaveqw</i> <b>n. sp.</b>, a presumed obligate troglophile, is very similar to <i>D. patrizii</i> but has a distinct ungual cavity. The ungual cavity may be a synapomorphy for this small clade of species that secondarily disappears with increasing troglomorphy. This scenario could account for the reduction of the cavity in <i>D. patrizii</i> and its complete loss in <i>D. tergestinus</i>, as well as its retention in <i>D. enkerlinius</i> and <i>D. naasaveqw</i> <b>n. sp.</b></p> <p> The monospecific genus <i>Dietersminthurus</i> was differentiated from similar genera by the presence of 5+5 eyes and eight setae on the anterior face of the dens (Palacios-Vargas <i>et al.</i> 1998). The reduction in eye number is recognised in a few other sminthurid genera, and the number of dental setae can be species-specific in several genera, e.g., <i>Sminthurus s. str.</i> Latreille, 1802 (see Bretfeld 1999) and <i>Sminthurinus</i> Börner, 1901 (Christiansen & Bellinger 1998). Ungual structure in <i>D. enkerlinius</i> appears to be identical to that of <i>D. patrizii</i> except that the ungual cavity is well-developed in <i>D. enkerlinius</i> and weakly developed in <i>D. patrizii</i>, and <i>D. enkerlinius</i> possesses eight anterior dental setae while <i>D. patrizii</i> has nine. <i>Disparrhopalites naasaveqw</i> <b>n. sp.</b> bridges these species by having a <i>patrizii</i> -like unguis with a well-developed <i>enkerlinius</i> -like cavity and nine anterior dental setae as in other <i>Disparrhopalites</i> spp. Therefore, <i>Dietersminthurus</i> is considered a junior subjective synonym of <i>Disparrhopalites</i>, and its sole species, <i>Dietersminthurus enkerlinius</i>, becomes <i>Disparrhopalites enkerlinius</i> (Palacios-Vargas, Cuéllar & Vázquez, 1998) <b>n. comb.</b></p> <p> <i>Varelasminthurus</i> was separated from similar genera by the absence of the posterior pretarsal seta (Da Silva <i>et al.</i> 2015). The number of pretarsal setae of <i>Disparrhopalites naasaveqw</i> <b>n. sp.</b> varies (1 or 2) from leg to leg, and therefore this character may not have significant validity at the generic level in Songhaicinae. However, the tunica of the single species, <i>V. potiguarus</i> Da Silva, Palacios-Vargas & Bellini, 2015, is heavy and fused to an external crest-like, serrated pseudonychium (Da Silva <i>et al.</i> 2015). Therefore, despite its similarities with <i>Disparrhopalites</i> spp. it is maintained here as a valid genus.</p>Published as part of <i>Wynne, J. Judson, 2017, Disparrhopalites naasaveqw n. sp. from caves at Wupatki National Monument, Arizona, synonymy of Dietersminthurus Palacios-Vargas, Cuéllar & Vázquez, 1998 with Disparrhopalites Stach, 1956 and composition of Songhaicinae (Collembola: Sminthuridae), pp. 77-90 in Zootaxa 4319 (1)</i> on page 81, DOI: 10.11646/zootaxa.4319.1.4, <a href="http://zenodo.org/record/888621">http://zenodo.org/record/888621</a>
Endonura incolorata Stach 1951, comb. nov.
No full textEndonura incolorata Stach, 1951 comb. nov. Figs 50–59, Tab. 5 Biloba incolorata Stach, 1951: 31 Type material. Lectotype male on slide by present designation, Ukraine, East Carpathians, Chornohora Range, “Zaroślak”, 1935, leg. J. Stach, det. J. Stach,. Paralectotypes, female and 2 juveniles on slides, same data as lectotype. Stach described Biloba incolorata based on numerous specimens collected in the forest reserve “Zaroślak” in the Chornohora Mts., but he did not designate the holotype. In Stach’s collection, housed at the Institute of Systematics and Evolution Animals PAS (Cracow), 4 syntypes on slides were found. One of them, in a best condition, was designated here as the lectotype. Other material. Ukraine, East Carpathians, Chornohora Range, „Zaroślak”, 1400 m alt., Norway spruce forest (subalpine belt), decaying wood, under bark of rotting logs, 7.ix. 1999, 10.ix. 1999, 12.ix. 1999, leg. R. J. Pomorski, D. SkarŻyński, A. Smolis, numerous individuals on slides; East Carpathians, Chornohora Range, south slope of Howerla Mt., 1000 m alt., Carpathian beech forest (montane belt), under bark of logs and in rooting wood, 11.ix. 1999, leg. A. Smolis, 3 females and 3 juveniles on slides. Other material is deposited in the Department of Biodiversity and Evolutionary Taxonomy of Wrocław University, Poland. Diagnosis. Habitus typical of the genus Endonura. Dorsal tubercles present and well developed, except tubercles Di on th. I. 2 + 2 unpigmented eyes present. Buccal cone rather short. Labral chaetotaxy 4 / 2, 4. Mandible thin with 3 teeth. Head with 3 chaetae Oc, chaetae A, B, C and D. Chaeta O absent. Tubercles Dl and (L+So) on head wih 6 and 10 chaetae respectively. Tubercles De on thoracic terga II and III with 3 and 4 chaetae respectively. Tubercles L on abd. III and IV with 3 and 6 chaetae respectively. Abd. IV and V with 8 and 2 tubercles respectively. Claw without inner tooth. Tibiotarsi with chaetae B 4 and B 5 short. Redescription. Habitus typical of the genus. Body length (without antennae): females 1.0–1.7 mm, males 1.0–1.2 mm, I instars 0.5 mm. Colour of the body white. 2 + 2 small unpigmented eyes (Figs 50–51, 53). Types of dorsal ordinary chaetae. Macrochaetae Ml thin, relatively short, straight, narrowly sheathed, gradually tapered and apically pointed (Figs 50–51, 54, 57); macrochaetae Mc and Mcc thin, straight, apically pointed; mesochaetae and microchaetae short, thin and pointed. Macrochaetae very similar in I instars and in adults (Fig. 53). All macrochaetae feebly serrated. Same number and arrangement of chaetae in adults and I instars, except chaetotaxy of ant. IV (see: Tab. 5 c) and genital field (complete absence of chaetae in first instars). a) Cephalic chaetotaxy–dorsal side. b) Cephalic chaetotaxy—ventral side. c) Chaetotaxy of antennae. 5 8 bs, 5 miA 8 bs, 5 miA d) Postcephalic chaetotaxy. Terga Legs Di De Dl L Scx 2 Cx Tr Fe T th. I 1 2 1 - 0 3 6 13 19 th. II 3 2 +s 3 +s+ms 3 2 7 6 12 19 th. III 3 3 +s 3 + s 3 2 8 6 11 18 Sterna abd. I 2 3 + s 2 2 VT: 4 abd. II 2 3 + s 2 2 Ve: 5 Ve 1 - present abd. III 2 3 + s 2 3 Vel: 5 Fu: 4–5 me 4–6 mi abd. IV 2 2 + s 3 6 Vel: 4 Vec: 2 Vei: 2 Vl: 4 abd. V 8 +s Ag: 3 Vl: 1 L': 1 abd. VI 7 Ve: 12 – An: 1mi 13 Head. Buccal cone short (Fig. 59). Labrum rounded, with ventral sclerifications as in Fig. 56. Labrum chaetotaxy 4 / 2, 4. Chaetotaxy of labium as in Fig. 56. Maxilla styliform, mandible thin tridentate. Chaetotaxy of antennae in adults and I instars as in Tab. 5 c. Apical vesicle distinct, simple. Sensilla S on ant.IV subequal, short and rather thick. Chaetotaxy of head as in Tab. 5 a, b, and Figs 51, 56, 59. Tubercles Cl and Af separate (Fig. 51). Chaetae O and E absent. Chaetae D and L 4 free. Tubercle Dl with 6 chaetae, chaeta Dl 3 present (Fig. 59). Elementary tubercle BE absent. Chaeta A shorter than B. Thorax, abdomen, legs. Body sensilla fine and smooth, shorter than nearby macrochaetae (Figs 54, 57). Chaetotaxy of th. and abd. as in Tab. 5 d and in Figs 50, 52–55, 57– 58. Tubercles Di on th. I not differentiated (Figs 50, 53–54). Chaetae De 3 on abd. I– III shorter than De 2. Chaetae De 2 on th. II – III and De 3 on th. III connected with tubercle De. Chaetae De 3 on abd. I– III connected with tubercle De (Figs 54, 57). The line of chaetae De 1 –sensillum perpendicular to the dorsomedian line on abd. I–IV (Figs 54–55). Tubercle L on abd. III and IV with 3 and 6 chaetae respectively (Figs 57–58). One chaeta L on abd. IV free. Furca rudiment with 4–6 microchaetae.Tubercles Di on abd. V separate, fused with tubercles De, Dl and L (Figs 50, 53, 57). Chaeta L' on abd. V present. Cryptopygy absent. Chaetotaxy of legs as in Tab. 5 d. Tibiotarsi with short chaetae B 4 and B 5. Claw without inner tooth. Discussion. See: Discussion of E. tatricola. Variability. According to the original description (Stach 1951), chaeta Di 3 on abd. V was absent. Analysis of available material showed that the chaeta was absent only in approx. 50 % specimens and present in remaining ones. Distribution. E. incolorata was recorded from Ukraine (Eastern Carpathians, Stach 1951, Kaprus’ et al. 2006), Slovakia (Western Carpathians, Low Tatra Mts., Nosek 1964, 1969), Poland (Western Carpathians, Beskid Żywiecki Mts., Stach 1964) and recently from Rumania (Eastern Carpathians, Dányi et al. 2006). Ecological remarks. A mountain saproxylic species, occurs in beech, fir–spruce (montane belt) and spruce forests (subalpine belt), not reported above tree line. Found mostly in humid rotting wood and under bark of decaying logs, but also in litter and soil with fine woody debris. First instars were collected in September. Remarks. Stach (1951) described Biloba incolorata based on specimens from Czornohora Mountains (Ukrainian Carpathians). Later the species was considered as member of Neanura (e.g. Gisin 1960, Salmon 1964, Stach 1964, Massoud 1967), Adbiloba (Cassagnau 1979) or as “ Biloba ” incolorata (Babenko et al. 1988). A detailed analysis of types and a new material (collected in the type locality) made it possible to ascertain that B. incolorata undoubtedly belonged to the genus Endonura (see: Discussion of Endonura). Furthermore, this allowed me to redescribe this species using a set of modern, diagnostic characters.Published as part of Smolis, Adrian, 2008, Redescription of four Polish Endonura Cassagnau, 1979 (Collembola, Neanuridae, Neanurinae), with a nomenclature of the ventral chaetae of antennae, pp. 9-36 in Zootaxa 1858 on pages 31-34, DOI: 10.5281/zenodo.18360
Endonura taurica Stach 1951
No full textEndonura taurica (Stach, 1951) Figs 1–14, Tab. 1 Biloba taurica: Stach 1951: 34 Neanura taurica: Dunger 1974: 14 Material examined. 3 adult females and juvenile; originally labelled “Krym Rezerwat”, VI–X. 1929, leg. W. Bukowski, det. J.Stach (ISEA). Other material. Female on slide, leaf litter in beech forest, ca. 800 m a.s.l., valley of Avunda River, Partyzans’ke near Gurzuf, Crimea, Ukraine, 10.IX. 1997, leg. R. J. Pomorski, D. SkarŻyński, I. Kaprus’. 3 adult females and male on slides, leaf litter on mountain meadow, Nikits’kyi Pereval, ca. 1450 m a.s.l., neighborhood of Jalta, Crimea, Ukraine, 12.IX. 1997, leg. R. J. Pomorski, D. SkarŻyński, I. Kaprus’. Adult female and juvenile on slide, soil and litter in mixed forest, Jaltyns'kyi Reserve, neighborhood of Jalta, Crimea, Ukraine, 11.ix. 1997, leg. R. J. Pomorski, D. SkarŻyński, I. Kaprus’ (ZIWU). Diagnosis. Habitus typical of the genus Endonura. Dorsal tubercles present, well developed. 2 + 2 eyes present. Buccal cone not elongated, wide. Labral chaetotaxy 4 / 2, 4. Mandible strong with 4 teeth. Head with 3 chaetae Oc, chaetae A, B, C, D, E (chaeta O absent). Tubercle (L+So) on head with 9 chaetae. Tubercles De on thoracic terga II and III with 3 and 4 chaetae respectively. Tubercles L on abd. III and IV with 4 and 6–7 chaetae respectively. Claw with or without inner tooth. All tibiotarsi with long chaetae B 4 and B 5. Redescription. Habitus typical of the genus (Fig. 1). Body length (without antennae) 0.78–1.38 mm. Colour of the body greyish-blue. 2 + 2 medium, dark-pigmented eyes. Types of dorsal ordinary chaetae: macrochaetae Ml and Mc thickened, narrowly sheathed, feebly serrated and not pointed (Figs 1, 13–14); mesochaetae thin and pointed. Number of other cephalic chaetae: Vi, 6; Ve, 11; labrum, 4 / 2, 4; labium, 11, 0x; ant. I, 7; ant.II, 11; ant. III, 17 + 5 s; ant. IV, 8 S+i +or+ 12 mou. b) Postcephalic chaetotaxy. Sterna abd. I 2 3 + s 2 3 VT: 4 abd. II 2 3 + s 2 3 Ve: 4–6 Ve 1 present Head. Buccal cone short and wide (Fig. 2). Labrum chaetotaxy 4 / 2, 4 (Fig. 3), ventral sclerifications as in Fig. 2. Chaetotaxy of labium as in Fig. 2 and Tab. 1. Maxilla slender with two small apical teeth at apex (Fig. 5). Mandible strong, with 2 basal and 2 apical teeth, the subapical one very small (Fig. 4). Chaetotaxy of antennae as in Tab. 1. Apical vesicle distinct, trilobed. S-chaetae subequal, long and moderately thickened. Chaetotaxy of head as in Figs 1–2, 7– 8 and Tab. 1. Chaeta D free (Fig. 8) or connected with tubercle Cl (Figs 1, 7).Tubercle Af wide (Fig 1, 7–8), chaeta O absent. Tubercle (L+So) with 9 chaetae (chaeta So 2 absent). Thorax, abdomen, legs. Chaetotaxy of th. and abd. as in Figs 1, 6, 14 and Tab. 1. Tubercles Di on th.I present or absent (Fig. 1). Tubercles L on abd. III and IV with 4 and 6–7 chaetae respectively (Fig. 6). Cryptopygy absent. Chaetotaxy of legs as in Figs 9–12 and Tab. 1. Claw with or without small inner tooth (Figs 9– 12). All tibiotarsi with long chaetae B 5 and B 4. Discussion. (see discussion of E. gracilirostris sp. nov.) Remarks. E. taurica was briefly described from the Crimea (Alushta reserve forest) by Stach (1951) and redescribed by Dunger (1974). Since both mentioned descriptions provided poor information on chaetotaxy we decided to make a modern redescription of this species based on extensive material (including Stach’s specimens).Published as part of Smolis, Adrian, Skarżyński, Dariusz, Pomorski, Romuald J. & Kaprus', Ighor J., 2007, Redescription of Endonura taurica (Stach, 1951) and E. quadriseta Cassagnau & Péja, 1979, and description of two new species of the genus Endonura Cassagnau, 1979 (Collembola: Neanuridae: Neanurinae) from the Crimea (Ukraine), pp. 19-35 in Zootaxa 1442 on pages 20-24, DOI: 10.5281/zenodo.17603
Heteroisotoma Stach 1947
No full textGenus Heteroisotoma Stach, 1947 Type species: Isotoma andrei Mills, 1934 Diagnosis. White or with rare scattered pigmentation, with 0 + 0 to 4 + 4 ocelli. PAO boat-shaped: elongate, strongly protruding, with pointed ends, in extreme case subdivided into lobes. Maxillary outer lobe with trifurcate palp and 4 sublobal hairs. Labial palp with all papilla (A-E), 16 (more rarely 15) guards, and four processes: lateral one, as common for Collembola, medial at papilla A, and processes B' and D' on inner side of papillae B and D, respectively. The development of two last processes varies depending on species. Maxilla without expanded and elongated lamellae. Tibiotarsi without clavate tenent hairs, with 8–9 setae in distal whorl. Ventral tube with at least 30 setae. Tenaculum with more than 5 setae. Furca long, reaching ventral tube. Distal part of manubrium with 1 + 1 to 4 + 4 ventroapical spines or spiniform setae (excluding H. jacutica). Dens with or without spines along inner side. Body macrosetae long. Tergal sensilla usually well differentiated, thin, often elongated on the last abdominal segments. Abd. V with 4 sensilla in characteristic position as 2 as-sensilla in front and 2 medial accp-sensilla in posterior position. Setal glands, in understanding of Deharveng (1981), present under manubrial and dental spines. For synonyms of the genus see Potapov and Stebaeva (1992). Labial palp. The main part of labial palp, if not reduced, is strikingly similar in different families of Collembola (Fjellberg 1999) and consists of several setae, five papillae (A–E) each armed with terminal sensillum and several guard setae ("guards"), and a lateral process (lost in few families). In Isotomidae, the palp is the most complete and the least differentiated. In the family at whole, the labial palp varies depending on species and genus and appears to be of great taxonomical value (Fjellberg 1999). Unlike in other subfamilies, species of the subfamily Isotominae, however, show invariable labial palp having full set of labial papillae, 15–16 "guards", and a characteristic finger-like lateral process (with the genus Metisotoma Maynard as an exception). In the genus Heteroisotoma, however, the labial palp undergoes the multiplication of un-armed (i.e. those which lacking guards) processes. It bears totally 4 such processes (Figs 1–5, 33): one common finger-like lateral process, one medial process of the same shape, and two more broad ones B' and D'. Such a structure has not before been noted before in Isotomidae indicating the monophyletic origin of the genus Heteroisotoma. Length and width of the processes vary depending on species. Heteroisotoma sinorossica sp. nov. has smaller B' and D' processes, so does H. stebaevae with B' (but more material is wanted to confirm the intraspecifiс variability of the character). The un-armed additional process on labial palp is known only in Desoria trispinata (MacGillivray), which has socalled "a particular lobe" at the base of papilla D (Fig. 58 in Fjellberg 1999). The highly evolved labium of Archisotoma Linnaniemi (subfamily Anurophorinae s.l.) has numerous additional papillae but each is armed with associated guard (b 1–4, d 1–4, e 1–6). Dental spines. The generic position of the two new species described in the present paper is somewhat uncertain. Due to the presence of spines on dens H. sinorossica sp. nov. and H. heteronomica sp. nov. may formally belong to the genus Semicerura for which the modern generic diagnosis is unknown. Type species, Semicerura bishopi Maynard (USA), has strong papillate spines on dens and a full set of ocelli. In the most characters, two new species resemble members of the genus Heteroisotoma Stach according to Potapov and Stebaeva (1992). They share the shape of PAO, general pattern of sensillar chaetotaxy of last abdominal segments, spines on manubrium, tridentate mucro, and complete or partial loss of ocelli. Because S. bishopi was not redescribed using modern taxonomical characters we place two our species to the genus Heteroisotoma. The labial palp of S. bishopi is unknown but two species studied by us in Eastern Asia, S. goryshini and S. sp. cf. bishopi, have neither medial process nor lobes B' and D' (see Fig. 36 for S. goryshini). In addition, Semicerura has the two last abdominal segments fused (vs. separated in Heteroisotoma), a simple PAO (vs. boat-shaped), and roundish (vs. sharp) apical labral ridges. A tridentate mucro, reduced apical whorl of setae on tibiotarsus (8–10 in Semicerura), and spines on dens and manubrium, however, are shared characters of these genera. So far, Isotoma s.str, Pseudisotoma Handschin, Semicerura, Heteroisotoma, and possibly also Folsomotoma Bagnall form a natural group in Isotominae with a reduced whorl of distal setae on tibiotarsi and an independent tendency to form spines on different parts of furca. The taxonomical validity of Semicerura and Heteroisotoma needs further study, also their relation with "spined" southern taxa (Greenslade 1989) remains even less understood. Misturasotoma Bernard & Christiansen, 2010, a large-sized blind genus from North America, formally resembles Heteroisotoma. The former genus, however, has 11 apical setae on tibiotarsi, quadridentate mucro, dense cover of equally short sensilla on two last abdominal segments, and have no additional processes on labium, that indicate a similarity with Desoria Nicolet and its allies.Published as part of Jie, Ding, Potapov, Mikhail & Sokolova, Elena, 2011, Further study on the labial palp in the Isotomidae (Collembola) with reference to the genus Heteroisotoma Stach, pp. 49-61 in Zootaxa 2991 on page 50, DOI: 10.5281/zenodo.20394
Plusiocampa (Plusiocampa) spelaea Stach 1930
No full textPlusiocampa (Plusiocampa) spelaea Stach, 1930 Table 10 Plusiocampa spelaea Stach, 1930: 278. Diagnosis (Stach 1930; Condé 1954a; Paclt 1957) Non-troglomorphic species. Antennae with 20–22 antennomeres. Pronotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 3+3 la, 2+2 lp, 1+1 mp macrosetae; metanotum with 1+1 ma, 1+1 la, 2+2 lp, 1+1 mp macrosetae. Two ventral tibial macrosetae. Urotergites I–II with 1+1 post macrosetae; urotergite III with 2+2 post macrosetae; urotergites IV–VII with 1+1 la, 5+5 post macrosetae; urosternite I with 8+8?, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Remarks The study of five females and one juvenile collected in four caves (see Table S 2 in Supplementary Material) has allowed the completion of the former description, although new specimens from the type locality are needed for a proper redescription of this species. Nevertheless, a complementary and differential description has been made using the studied material. Non-troglomorphic species (Table 10). Antennae with 21 and 23 doliiform antennomeres, apical antennomere 1.3× as long as wide, with a small cupuliform organ occupying 1/7 of antennomere, with four simple olfactory chemoreceptors. Small ventral sensillum of third antennomere between d and e macrosetae. Non-protruding frontal process with non-tubercular setae. Labial palp suboval with a thick lateral anterior sensillum, two guard setae, up to five normal setae and 75 neuroglandular setae. Pronotum with 2+2 la to 3+3 la macrosetae; long, barbed notal macrosetae; notal clothing setae long, smooth or with one or two distal barbs. Calcars with two rows of long barbs. Smooth subapical tarsal setae. Slightly unequal claws (1.15–1.20) with large lateral crests and a small backward overhang. Urotergite IV with 4+4 post macrosetae (5+5 according to redescription in Paclt 1957); urotergites VI– VII with 2+2 la macrosetae (1+1 la according to Paclt 1957); urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 7+7, urosternites II–VII with 4+4, urosternite VIII with 2+2 macrosetae. Stylar setae with a few long, thin bars. Female with subcylindrical appendages bearing up to 9 glandular a 1 setae. Habitat and distribution Apparently, this species only lives in caves, but its non-troglomorphic shape reveals a more superficial habitat (MSS or soils). It has been found in six caves in the southern karst areas of the Western Carpathians, where it was described from the Domica-Baradla cave system in the Slovak and Aggtelek Karst, on the border between Slovakia and Hungary.Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 83-84, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/365982
Endonura transcaucasica Stach 1951
No full textEndonura transcaucasica (Stach, 1951) Figs 1–7, Tab. 1 Biloba transcaucasica: Stach 1951: 34. Type material. Lectotype by present designation: adult female on slide, Georgia, Caucasus, Tbilisi, 21.IV.1918, leg. Roszkowski, coll. J. Stach (ISEA). Diagnosis. Habitus typical of the genus Endonura. Dorsal tubercles present and well developed. 2+2 pigmented eyes. Buccal cone short, labrum nonogival. Head with chaetae A, B, D, E, F and G. Chaetae O and C absent. Tubercles Cl and Af separate. Chaetae D free. Tubercles Dl and (L+So) on head with 6 and 7 chaetae respectively. Tubercles Di and De on th. I present and separate. Tubercles De on th. II and III with 3 and 4 chaetae respectively. Tubercles L on abd. III and IV with 3 and 6 chaetae respectively. Abd. IV and V with 8 and 3 tubercles respectively. Claw without inner tooth. Tibiotarsi with chaetae B4 and B5 short. Redescription. Habitus typical of the genus. Body length of lectotype (without antennae): 1.65 mm. Colour of the body white. 2+2 large pigmented eyes (Fig. 1). Types of dorsal ordinary chaetae. Macrochaetae Ml relatively long, strongly thickened, almost cylindrical, arclike or straight, narrowly sheathed, densely serrated, apically rounded (Figs 1, 5–7); macrochaetae Mc and Mcc thickened, straight and rounded at apex or pointed; mesochaetae and microchaetae short, thin, feebly serrated and pointed. Head. Buccal cone short. Labrum nonogival, with ventral sclerifications as in Fig. 2. Labrum chaetotaxy 2/2, 4. Labium with four basal, three distal and four lateral chaetae, papillae x absent. Maxilla and mandible as in Figs 3, 4. Antennal segments I and II with 7 and 11 chaetae respectively. Chaetotaxy of antennal segments III and IV impossible to recognize. Apical vesicle poorly visible, trilobed. Chaetotaxy of head as in Tab. 1 a, b, and Fig. 1. Head with chaetae A, B, D, E, F and G. Chaetae O and C absent. Tubercles Cl and Af separate. Chaeta D free. Tubercle Dl with 6 chaetae, chaeta Dl3 present. Tubercle (L+So) with 7 chaetae, chaetae So2, So5 and L3 absent (Fig. 1). Elementary tubercles BE and CD present. Chaeta E connected with tubercle Af. Chaeta A shorter than B. Tubercle Oc with two chaetae, chaeta Oca absent (Fig. 1). Chaetae Ocp longer than A. Chaeta De2 as mesochaeta. Thorax, abdomen, legs. Body s–chaeta thin and smooth, distinctly shorter than nearby macrochaetae (Fig. 5). Chaetotaxy of th. and abd. as in Tab. 1 d and in Figs 1, 5–7. Tubercles Di on th.I well differentiated and not fused with De. Tubercles De on th. II and III with 3 and 4 chaetae respectively. Chaetae De2 on th. II–III and De3 on th. III connected with tubercle De. Chaetae De3 on abd. I–III connected with tubercle De (Fig. 5). Chaetae De3 on Th. III and abd. I-III slightly longer than De2. The line of chaetae De1–chaeta s perpendicular to the dorsomedian line on abd. I–III (Fig. 5). Furca rudiment without microchaetae. Abd. IV and V with 8 and 3 tubercles respectively, tubercles Di on abd. V fused (Fig. 6). Tubercle L on abd. IV with 6 chaetae (Fig. 7). Chaetae L' and Vl on abd. V present. No cryptopygy. Chaetotaxy of legs as in Tab. 1 d.. Claw without inner tooth. Tibiotarsi with chaetae B4 and B5 short. a) Cephalic chaetotaxy––dorsal side. b) Cephalic chaetotaxy––ventral side. c) Postcephalic chaetotaxy. Terga Legs Di De Dl L Scx2 Cx Tr Fe T th. I 1 2 1 - 0 3 6 1 2 1 9 th. II 3 2+s 3+s+ms 3 2 7 6 11 19 th. III 3 3+s 3+s 3 2 8 6 1 0 1 8 Sterna abd. I 2 3+s 2 3 VT: 4 abd. II 2 3+s 2 3 Ve: 5 Ve1 - present abd. III 2 3+ s 2 3 Vel:4 Fu: 5–6 me 0 mi abd. IV 2 2+ s 3 6 Vel: 3 Vec: 2 Vei: 2 Vl: 4 abd. V (3+3) 5+s Ag: 3 Vl: 1 L': 1 abd. VI 7 Ve:13-14 An: 2mi Discussion. Because of the absence of chaetae C and O on head, feature generally very rarely observed within Endonura, E. transcaucasica resembles E. baculifer Deharveng, 1979 described from Portugal. However, both species differ in a few characters, important from Neanurinae taxonomy point of view: presence/absence of chaetae E on head (in transcaucasica present, in baculifer absent) number of chaetae Dl on head (in transcaucasica 6, in baculifer 5) and number of chaetae (L+So) on head (in transcaucasica 8, in baculifer 7–9). In addition, the scientific name of Portugal species was derived from an unusual shape of its macrochaetae Di 1 of abdomen V which are extremely thickened and blunt at apex (Deharveng 1979: Figs 9 B, C).Published as part of Smolis, Adrian, 2016, On the identity of some poorly known species of the genus Endonura Cassagnau, 1979 (Collembola, Neanuridae, Neanurinae), pp. 237-250 in Zootaxa 4173 (3) on pages 238-239, DOI: 10.11646/zootaxa.4173.3.3, http://zenodo.org/record/15932
Checklist of Maltese Collembola with nomenclatural notes and new synonyms
No full textAbstract: Maltese Collembola records are reviewed and a checklist is provided. In a critical examination of previous publications considering Maltese Collembola, several nomenclatural issues were uncovered. Based on these, the following nomenclatural changes are proposed: Dicyrtoma dorsosignata Stach, 1924 stat. nov., with the following combinations as synonyms: D. fusca var. dorsosignata Stach, 1924 syn. nov.; D. fusca var. pallida Stach, 1924 syn. nov.; D. fusca f. strigata Stach, 1924 syn. nov.; D. melitensis Stach, 1957 syn. nov.; D. melitensis var. dorsosignata Stach, 1957 syn. nov.; D. melitensis var. pallida Stach, 1957 syn. nov. and D. melitensis Stach, 1967 syn. nov.. Entomobrya abrupta (Stach, 1924) stat. nov. with the synonym E. melitensis Stach, 1963 syn. nov. The following nomina nuda can be considered lapsus calami: Hypogastrura melitensis Stach, 1967 is a nomen nudum and refers to H. varians Stach, 1967. Triacanthella transilvatica Stach, 1967 is nomen nudum and refers to T. terrasilvatica Salmon, 1943. T. perfusa Stach, 1967 is nomen nudum and refers to T. purpurea Salmon, 194
Dicyrtomina minuta sensu Stach 1957
No full textDicyrtomina minuta (O. Fabricius, 1783) sensu Stach 1957 * Podura minuta O. Fabricius, 1783: 307. Dicyrtomina dorsimaculata Lubbock, 1873: 117. Dicyrtomina minuta – Willem 1900: 63. — Stach 1957: 43. DISTRIBUTION. — France. Bourlet 1841 (D. dorsimaculata). Ariège. GrottedeL’Herm (Cauchois & Théodoridés 1954). — Massif del’Arize, 400- 1400 m (Deharveng & Lek 1995). Auxois, Morvan, Côte-d’Or. Saint-Jean-de-Losne (Denis 1921, var. couloni). Bas-Rhin. Merkwiller-Pechelbronn (Gillet & Ponge 2005). Bouches-du-Rhône. Camargue (Poinsot 1973, 1979). Essonne. Parcdu Laboratoiredu MNHN à Brunoy etforêt de Sénart (Ponge 1973). — Forêtde Sénart (Ponge 1980, 1993; Salmon & Ponge 1999; Ponge etal. 2002). — Maresde Sénart (Pichard etal. 1989). — Parcdu Laboratoiredu MNHNà Brunoy (Vannier & Kilbertus 1981; Betsch 1991; Chauvat & Ponge 2002). Eure. Lyons-la-Forêt (Carl 1899). Finistère. Prat-ar-Coum prèsde Lannilis (Delamare Deboutteville 1950c). Haute-Garonne. Toulouse (Cassagnau & Delamare Deboutteville 1951; Cassagnau 1962a). — Ruisseaux de La Maure et de Ruau (Lek-Ang & Deharveng 2002). — Gar-Cagire près de Saint-Gaudens (Salmon etal. 2010). Hautes-Pyrénées. Valléesde Couplanet dela Gargante (Cassagnau 1964a). — Hautevalléedel’Aureetmassif du Néouvielle (Lauga- Reyrel & Lauga 1995). — Montagned’Arize (Deharveng 1996). Hauts-de-Seine. Maresde Clamart (Pichard etal. 1989). Landes. Roquefort (Jeannenot 1957). Loir-et-Cher. Forêt près de Salbris (Prat & Massoud 1980). Loiret. Forêtd’Orléans (Ponge etal. 1993). Nièvre. Parcrégionaldu Morvan (Ponge etal. 2003). Norddela France. Finot 1890 (Dicyrtoma dorsi-maculata). Pyrénées-Orientales. Forêtdela Massaneprèsde Banyuls-sur-Mer (Travé etal. 1954). Seine-et-Marne. Forêtde Foljuif (Massoud etal. 1984). Sud-Ouestdela France; Pyrénées. Cassagnau 1961. Tarn. Sidobre (Izarra 1969; Bonnet etal. 1972). Vendée. Île-d’Yeu (Denis 1922a). Vosges. Lorraine (Picard 1962). PHYSIOLOGIE. — Vannier & Kilbertus 1981 (migration substances organo-minérales). ÉCOLOGIE. — Picard 1962 (faune psammique); Bonnet et al. 1972 (écologie, muscicoles); Poinsot 1973, 1979; Ponge 1973, 1980, 1993; Prat & Massoud 1980 (sol forestier); Massoud etal. 1984 (piétinement); Pichard etal. 1989; Betsch 1991 (privation delitière); Ponge etal. 1993 (écologie expérimentale, litière), 2002 (toxicité, herbicide), 2003 (bioindicateur); Deharveng & Lek 1995; Deharveng 1996 (déboisement, reboisement); Salmon & Ponge 1999; Chauvat & Ponge 2002 (métaux lourds); Lek-Ang & Deharveng 2002 (zone humide); Gillet & Ponge 2005 (pollution); Salmon etal. 2010 (diversité, peuplement, endémisme).Published as part of Thibaud, Jean-Marc, 2017, Catalogue des collemboles de France, pp. 297-436 in Zoosystema 39 (3) on pages 402-403, DOI: 10.5252/z2017n3a1, http://zenodo.org/record/549627
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