3,884 research outputs found
Agra delgadoi Erwin, new species
No full textAgra delgadoi Erwin, new species (Figs. 26 a, 26 b, 26 c, 26 d) Holotype. Male, Panamá, Panamá Province, Cerro Campana, 800m, 08º 40 'N 79 º 56 'W, July, (H. Stockwell)(USNM: ADP 59923). Diagnosis. This is the only species of Agra in Costa Rica with metallic purple nearly truncate elytra, and completely pale antennae. Description. Color and luster: Rufopiceous. Elytra metallic purple. Venter and appendages paler; antennomeres 411 testaceous. Surface shiny. Form: Head (Fig. 26 a) of female behind eyes short, hind angles rounded. Pronotum and prosternum finely punctate, prothorax widest at middle. Elytron with apex (Fig. 26 b) slightly oblique, lateral tooth small, sutural apex rounded, not projected acutely, apical margin very slightly emarginate. Aedeagus (Fig. 26 d) with ostium short, apex rounded, shorter than wide; parameres unisetose. Size: small, 8.5 to 10.0mm in length, 2.30 to 2.66mm in width. Other specimens examined. Three female paratypes. Costa Rica: Tortuguero and Cordillera Volcánica Central Conservation Areas. Costa Rica: Limón, Tortuguero National Park, Cerro Tortuguero, 0m 120m, LN 285000,588000, April (R. Delgado)(INBio: CRI 000242885); Cartago, Turrialba, CATIE, 600m, 09º 53 'N 83 º 38 'W, May (H. & A. Howden)(USNM: ADP 66237). Panamá, Panamá Province, Cerro Campana, 800m, 08º 40 'N 79 º 56 'W, July, (H. Stockwell)(USNM: ADP 68252). Specific epithet. The specific epithet, delgadoi, is the Latinized genitive form of the family name of the collector of one of the secondary type specimens, parataxonomist, Roberto Delgado of Cartago, Costa Rica.Published as part of Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119 on pages 58-59, DOI: 10.5281/zenodo.16280
Agra giesberti Erwin, new species
No full textAgra giesberti Erwin, new species (Figs. 3 a, 3 b, 3 c, 3 d, 3 e, 3 f) Holotype. Female. COSTA RICA: Cartago, 15km NE Turrialba, 9 º 53 'N 83 ° 38 'W, April (E. Giesbert) (CAS: ADP 62298). Diagnosis. This is the only species of the buqueti speciesgroup known from Costa Rica. Its alternating interneurs with large and small fovea provide an easy recognition attribute, as does the elongate, multisetiferous and anteriorly tubiform prothorax (Fig. 3 e). Description. Color and luster: Piceous. Head rufinistic, appendages rufotestaceous, and femora infuscated in some adults. Surface very shiny. Head (Fig. 3 a) behind eyes moderately short, markedly tapered in male, more rounded in female. Prothorax (Fig. 3 e) at anterior third somewhat depressed. Elytron (Fig. 3 b) with apex shallowly obliquely arcuate, sutural angle produced. Sternum VI (Fig. 3 d) shallowly vnotched in both sexes. Aedeagus (Fig. 3 f) markedly narrowed apically, ostium of medium length, apex subscimitariform. Stylomere 2 (Fig. 3 c). Size: large, 17.8 to 21.5mm in length, 4.0 to 5.8mm in width. Other specimens examined. Costa Rica: 2 paratypes, 1 male, 1 female, from the following Conservation Areas: Guanacaste, Tortuguero. Costa Rica, Cartago, 1 f, Turrialba, 15.0 km NE, 83 ° 38 'W, 9 ° 53 'N, April, (E. Giesbert)(CAS: ADP 62298); 1 f, Limón, Cariari, nr, Hacienda La Suerte, May, (F.T. Hovore)(FTHC: ADP 70512). I have seen additional specimens (paratypes) from Guatemala and Panamá. Specific epithet. The specific epithet, giesberti, is the Latinized genitive form of the family name of the late Edward Giesbert, amateur Coleopterist extraordinaire, who took time from his beloved cerambycids to collect an Agra or two. Notes. This species ranges from sea level (Panamá) to 2000m (Guatemala).Published as part of Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119 on pages 19-20, DOI: 10.5281/zenodo.16280
Agra fugax Erwin, new species
No full textAgra fugax Erwin, new species (Figs. 8 a, 8 b, 8 c, 8 d) Holotype. Male. COSTA RICA: Puntarenas, Carara Biological Reserve, Estacíon Quebrada Bonita, 50m, 09° 46 ' 35 ”N 84 ° 36 ' 11 ”W, June (R. Zuñiga)(INBio: CRI 000 349227). Diagnosis. Femur concolorous; elytron with sutural and lateral apices modestly dentate, margin between them slightly arcuate. Head markedly convex and nearly quadrate basally. Elytral interneurs each a series of large shiny (brassy) foveae. Description. Color and luster: Piceous with testaceous appendages, femur concolorous; elytral interneurs each a series of large shiny (brassy) foveae. Form: Head (Fig. 8 a) behind eyes markedly convex and nearly quadrate posteriorly. Elytron (Fig. 8 b) with apex moderately oblique, lateral and sutural dentiform projections moderately large, acute; interneurs each a series of large shiny (brassy) foveae. Abdominal sternum VI (Fig. 8 c) shallowly Unotched in female. Aedeagus (Fig. 8 d) with ostium moderately short, apex simple, cordiform. Size: medium, 13.6 to 13.8 mm in length, 3.2 to 3.8 mm in width. Other specimens examined. Costa Rica: 1 paratype, female, from the Cordillera Volcánica Central Conservation Area: 1 f, Cartago, Turrialba, 83 ° 38 'W 9 ° 53 'N, (WAR: ADP 4214). Specific epithet. The specific epithet, fugax, meaning swift, is a manuscript name from Max Liebke's collection in WAR that I am retaining, however, I have chosen a different Holotype specimen than he labeled with his manuscript name, a male that is in better condition.Published as part of Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119 on pages 28-29, DOI: 10.5281/zenodo.16280
Agra pitilla Erwin, new species
No full textAgra pitilla Erwin, new species (Figs. 24 a, 24 b, 24 c, 24 d) Holotype. Male. COSTA RICA: Guanacaste, P.N. Guanacaste, Estacíon Pitilla, 9.0 km S Santa Cecilia, 700m, 10 ° 59 ' 26 ”N, 85 ° 25 ' 40 ”W, January, (GNP Biodiversity Survey)(INBio: CRI 001024216). Diagnosis. Black fore body with rufinistic elytral apex, the apical margin infuscated and markedly lobed at middle. Hind trochanter sparsely setiferous. Description. Color and luster: Black. Apical 2 / 3 of elytra rufinistic, margin of elytral apex, abdominal sternum VI, and tergum VI infuscated; tarsi and abdomen rufotestaceous. Form: Labrum flat. Head (Fig. 24 a) markedly short behind eyes, quadrate in both sexes. Elytra with markedly and evenly punctulate interneurs, apex (Fig. 24 b) slightly lobed between lateral and sutural angles. Metasternum of male densely pubescent; abdominal sterna each with long scattered setae. Apex of sternum VI (Fig. 24 c) in male and female acutely angulate apicolaterally, caudal margin emarginate, slightly unotched in male, vnotched in female. Male middle tibia with small apicomedial brush. Aedeagus (Fig. 24 d) ostium short, slightly left pleuropic; apex of phallus a simple diamondshaped lobe; left paramere unisetose, right paramere glabrous. Size: medium, 13.0 to 13.5 mm in length, 3.1 o 3.7 mm in width. Other specimens examined. Costa Rica: 7 paratypes, 5 males, 2 females, from the Guanacaste Conservation Area. Costa Rica, Alajuela, 1 f, Dos Rios, 2.0 km W, N slope of Volcán de Rincon, 550m, 10 ° 50 ' N, 085° 22 ' W, May (J.T. Doyen & P.A. Opler)(UCB: ADP 80686); Guanacaste, 1 f, Santa Cecilia, 9.0 km S, 700m, 10 ° 59 ' 26 N, 85 ° 25 ' 40 ' W, LN 330200,380200, June (C. Moraga)(INBio: CRI 002047455), 6 males, P.N. Guanacaste, Estacíon Pitilla, April (P. Rios)(INBio: CRI 001794228), March (P. Rios, C. Moraga & R. Blanco)(INBio: CRI 000 211428), December (C. Moraga)(INBio: CRI 000 637527), May (P. Rios & M. Zumbado) (CRI 000 269888), LN 330200, 380200 # 3262, October (P. Rios)(INBio: CRI 001 997986), January (GNP Biodiversity Survey)(INBio: CRI 00 1024216). Specific epithet. The specific epithet, pitilla, is a noun in apposition based on the name of the INBio Field Station at which the Holotype was collected. Notes. All specimens are from the Guanacaste Conservation Area, 7 of them from Estacíon Pitilla.Published as part of Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119 on pages 53-54, DOI: 10.5281/zenodo.16280
Agra santarosa Erwin, new species
No full textAgra santarosa Erwin, new species (Figs. 16 a, 16 b, 16 c, 16 d) Holotype. Male, COSTA RICA: Guanacaste, Santa Rosa National Park, 280m, 10 º 50 'N 85 º 37 'W, May [1980], (D.H. Janzen & W. Hallwachs) (USNM: ADP 55010). Diagnosis. Black. Femora rufous, tibiae rufopiceous, knees and tarsi piceous. Elytral interneurs with regularly spaced and slightly cribriform punctulae; apex with more or less similarsized lateral and sutural dentiform projections. Eyes very large, hemispherical, head short behind eyes. Description. Color and luster: Black. Antennomeres 411 bicolored, red and black, tibiae rufous. surface Shiny. Form: Labrum entire. Head (Fig. 16 a) behind eye moderately short, tapered in both sexes, more so in male. Prothorax "inflated" at basal third. Prothorax subtubular, short, robust, sutures effaced, dorsal surface flattened and densely punctate. Elytral apex (Fig. 16 b) with more or less similar acute dentiform projections; interneurs with regularly spaced and slightly cribriform punctulae. Sternum VI (Fig. 16 c) deeply vnotched in male, shallowly vnotched in female. Aedeagus (Fig. 16 d) with ostium elongate, apex a double scimitar, parameres glabrous. Size: moderately large, 17.5 to 20.0mm in length, 4.86 to 5.16mm in width. Other specimens examined. Costa Rica: 2 paratypes, both females, from the following Guanacaste Conservation Area. Costa Rica, Guanacaste, 1 f, June, (D.H. Janzen & W. Hallwachs)(INBIO: ADP 55016), 1 f, Estacíon Maritza, Volcán de Orosi, East slope, 600m, LN 326900,373000, June, (GNP Biodiversity Survey)(INBIO: CRI 000 134050). Specific epithet. The specific epithet, santarosa, is a noun in apposition based on the type locality, Santa Rosa National Park. Notes. An illustration of famulagroup stylomere 2 is on the Agra fortuna web page.Published as part of Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119 on pages 40-41, DOI: 10.5281/zenodo.16280
Agra liv Erwin, new species
No full textAgra liv Erwin, new species (Figs. 15 a, 15 b, 15 c, 15 d) Holotype. Female, COSTA RICA: Puntarenas, Manual Antonio National Park, Quepos, 80m, 09° 23 'N 84 °09'W, January, (Bogenberger)(ZSBS: ADP 6893). Diagnosis. This is the only species of the famula speciesgroup in Costa Rica that is entirely black with pale tibiae and tarsi and with normal elytral intervals. Description. Color and luster: Black. Tips of mandibles, sides of antennomeres 511, and in some specimens sides of abdominal sterna rufopiceous. Surface shiny. Form: Labrum entire. Head (Fig. 15 a) behind eyes elongate, rounded in both sexes. Prothorax subtubular, short, robust, sutures effaced, widest at middle, dorsal surface moderately convex and sparsely punctate, punctures markedly small. Elytron (Fig. 15 b) with apex with more or less similar acute dentiform projections, interneurs with small regularly spaced and slightly cribriform punctulae, adjacent intervals not catenate. Sternum VI (Fig. 15 c) moderately deeply vnotched in male, shallowly vnotched in female. Aedeagus (Fig. 15 d) ostium elongate, apex a double scimitar, parameres glabrous. Size: medium, 14.0 to 15.5mm in length, 3.4 to 4.6mm in width. Other specimens examined. Holotype above, and a male and female from Panamá are labeled as Paratypes. Specific epithet. The specific epithet, liv, is a noun in apposition based on the first name of the actress Liv Tyler, starlet of the movie, Armageddon. The existence of this species of elegant beetle is dependent upon the rainforest not undergoing an Armageddon which is too much to hope for. Notes. The single Costa Rican record above is from the Pacifico Central Conservation Area. Famulagroup stylomere 2 can be found at the Agra fortuna web page.Published as part of Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119 on pages 39-40, DOI: 10.5281/zenodo.16280
Agra zuniga Erwin, new species
No full textAgra zuniga Erwin, new species (Figs. 31 a, 31 b, 31 c, 31 d, 31 e) Holotype. Female. COSTA RICA: Puntarenas, Manual Antonio National Park, Quepos, 80m, 09° 23 'N 84 °09'W, February (R. Zuñiga)(INBio: CRI 000 625744). Diagnosis. A. zuniga adults do not have a raised median keel on the anterior portion of the pronotal disc (Fig. 31 c) as in A. pia, although the area appears “polished.” Description. Color and luster: Head and pronotum black, elytra metallic purple, legs and venter very dark rufous, infuscated; surface shiny. Form: Head (Fig. 31 a) behind eye moderately elongate, quadrate, slightly rounded in female. Pronotum (Fig. 33 c) without a raised median keel on the anterior portion. Elytron (Fig. 31 b) with apex slightly oblique, projected acutely at both sutural and lateral corners, dentiform projections small, margin nearly straight medially. Male venter with extensive accessory patches of setae on metasternum, posterior coxae, trochanter, and abdominal sterna IIVI. Sternum VI (Fig. 31 d) deeply Unotched in male, shallowly Vnotched in female. Aedeagus (Fig. 31 e) with apical orifice moderately short, left pleuropic; apex a transverse ellipse. Size: small, 12.0 to 12.1mm in length, 3.2mm in width. Other specimens examined. Costa Rica: 1 paratype, male, also from the Pacifico Central Conservation Area. Costa Rica, Puntarenas, 1m, Res. Biol. Carara, Estacíon Quebrada Bonita, 50m, LN 194500,469850, May, (R. Zuñiga)(INBIO: CRI 000 195164). Specific epithet. The specific epithet, zuniga, is used as a noun in apposition honoring the collector of the Holotype specimen, parataxonomist, Rodolfo Zuñiga of Orotina, Alajuela, Costa Rica. Notes. Both known specimens are damaged. I selected the female as Holotype because it is a more complete specimen.Published as part of Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119 on pages 65-66, DOI: 10.5281/zenodo.16280
Agra ubicki Erwin, new species
No full text<i>Agra ubicki</i> Erwin, new species <p>(Figs. 18a, 18 b, 18c, 18d)</p> <p> <b>Holotype.</b> Male, <b>COSTA RICA:</b> Puntarenas, 3 km NE Golfito, 8º 39'N 83º10'W, 100m, May, (D. Ubick)(CAS: ADP4553).</p> <p> <b>Diagnosis</b>. Color piceous. Head markedly impressed on the occiput, almost foveate. Prothorax without the numerous long setae found in adults of <i>A. hilaris</i> Liebke. Elytral intervals not catenate.</p> <p> <b>Description</b>. Color and luster: Color piceous. Surface shiny. Form: Head (Fig. 18a) behind eyes moderately long, markedly rounded, occiput markedly impressed on the occiput, almost foveate. Prothorax widest at basal third, pronotum somewhat flattened with moderately dense coarse punctures. Elytron with apex (Fig. 18 b) almost truncate, lateral tooth small, sutural angle rounded, not projected acutely. Sternum VI (Fig. 18 c) moderately emarginate and v­notched in both sexes. Aedeagus (Fig. 18 d) with short ostium, apex of phallus a very broad triangular lobe, parameres glabrous. Size: small, 8.0 to 9.5mm in length, 2.48 to 3.24mm in width.</p> <p> <b>Other specimens examined.</b> Costa Rica: 4 paratypes, 1 female, 3 males, from the Osa Conservation Area. Costa Rica, Puntarenas, 1m, P.N. Corcovado, Madrigal, 3.0 km W, 83°36'W 8°29'N, May, (D. Ubick)(CAS: ADP4554), 1m, Peninsula de Osa, 83°36'W, 8°29'N, August, (D.R. Whitehead)(USNM: ADP59571), 1m, Golfito, 3.0 km NE, 100m, 83°10'W, 8°39'N, May, (D. Ubick)(CAS: ADP4545), shaking foliage, tropical wet forest, 1f, May, (D. Ubick)(CAS: ADP4552).</p> <p> <b>Specific epithet.</b> The specific epithet, <i>ubicki,</i> is the Latinized genitive form of the family name of the collector of the Holotype, Darryl Ubick of the Department of Entomology, California Academy of Sciences.</p> <p> <b>Notes.</b> Adults of this species group are commonly found in suspended dry leaves in the understory of Neotropical forests.</p> <p> The <b>palmata</b> species­group</p> <p> Diagnostic combination of the <i>palmata</i> species­group includes the following: Labrum flat; mentum with lateral lobes acute, not spiniform. Head elongate or not, broader across middle than occiput, eyes large; head posterior to eyes normally elongate, not ballooned (lateral aspect), eyes less than 1.5 times their own diameter from cervical constriction. Tibia moderately or slightly compressed, laterally barely expanded, apex never prolonged; all tibiae latero­apically oblique. Elytral apex bidentate, dentiform projections small.</p> <p> The <i>palmata</i> species­group was revised by Erwin (1984). At that time, it contained 39 species whose composite range extends from southern México to Paraguay. Since then, 13 new species have been discovered from throughout the overall range, plus another one from Costa Rica, described below.</p> <p> Four other species of the <i>palmata</i> species­group are known from Costa Rica, <i>A. atriperna</i> Erwin, <i>A</i>. <i>costaricensis</i> Liebke, <i>A. flavipes</i> Straneo, and <i>A. kayae</i> Erwin.</p>Published as part of <i>Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119</i> on pages 43-44, DOI: <a href="http://zenodo.org/record/162803">10.5281/zenodo.162803</a>
Agra dable Erwin, new species
No full textAgra dable Erwin, new species (Figs. 14 a, 14 b, 14 c, 14 d) Holotype. Male, COSTA RICA: Heredia, Estacíon Magsasay, Parque Nacional Braulio Carrillo, 200m, 10 ° 24 '04”N 84 °02' 57 ”W, March, (M. Zumbado)(INBio: CRI 000 351513). Diagnosis. Labrum entire. All black. Elytron with regular interneurs, 2, 4, and 6 with series of moderately impressed setigerous punctures, adjacent intervals catenate; interneurs with regularly spaced and moderately cribriform punctulae; apex with more or less similarsized lateral and sutural acute dentiform projections. Description. Color and luster: Black. Tips of mandibles, sides of antennomeres 411, and in some specimens, sides of abdominal sterna rufopiceous. Surface very shiny. Form: Labrum entire. Head (Fig. 14 a) behind eye elongate, taperrounded in both sexes. Prothorax subtubular, short, robust, sutures effaced, dorsal surface convex and densely punctate and transversely rugose, punctures very small. Elytron with apex (Fig. 14 b) moderately oblique, with more or less similar lateral and sutural acute dentiform projections, margin between dentiform projections moderately lobed; interneurs with regularly spaced and moderately cribriform punctulae and 2, 4, and 6 with series of moderately impressed setigerous punctures. Metasternum with scattered setae; abdominal sterna IIIVI of male with patches of discal setae. Sternum VI (Fig. 14 c) moderately vnotched in both sexes. Aedeagus (Fig. 14 d) with ostium elongate, apex a double scimitar, the distal with rounded sides, parameres glabrous. Size: medium, 14.5 to 18.0mm in length, 4.28 to 4.70mm in width. Other specimens examined. Costa Rica: 5 paratypes, 3 males, 2 females from the following Conservation Areas: Cordillera Volcánica Central, Guanacaste, Tortuguero. Costa Rica, Guanacaste, 1m, P.N. Guanacaste, Estacíon Pitilla, Santa Cecilia, 9.0 km S, 700m, 85 ° 25 ' 40 'W 10 ° 59 ' 26 "N, LN 330200,380200, June, (C. Moraga)(INBIO: CRI 000 819152), 1 f, May, (GNP Biodiversity Survey)(INBIO: CRI 001024226), 1m, P.N. Rincón de la Vieja, Estacíon Las Pailas, 800m, July, (K.E. Taylor)(INBIO: CRI 001133380), 1 f, October, (M. A. Zumbado)(INBIO: CRI 000515130), 1m, Limón, P.N. Tortuguero, Estacíon Cuatro Esquinas, 0m, 83 ° 32 'W 10 ° 34 'N, LN 280000,590500, June, (E. Quesada)(INBIO: CRI 000363904). Specific epithet. The specific epithet, dable, is part of the Spanish word, agradable, meaning “pleasing.” Notes. Adults of this species are remarkably similar to those of Agra solisi (below) except for leg color and obliqueness of the elytral apex. An illustration of famulagroup stylomere 2 is on the Agra fortuna web page.Published as part of Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), pp. 1-68 in Zootaxa 119 on pages 37-38, DOI: 10.5281/zenodo.16280
Figure 5 from: Erwin TL, Henry SC (2017) Hyboptera Chaudoir 1872 of the Cryptobatida group of subtribe Agrina: A taxonomic revision with notes on their ways of life (Insecta: Coleoptera, Carabidae, Lebiini). ZooKeys 714: 61-127. https://doi.org/10.3897/zookeys.714.15113
No full textFigure 5 -
Digital Photo-illustration, male aedeagus in repose, dorsal, ventral, left lateral aspects. A Hyboptera angulicollis Chaudoir, ADP007330 B Hyboptera biolat Erwin & Henry, sp. n., ADP007443 C Hyboptera vestiverdis Henry & Erwin, sp. n., ADP014856 D Hyboptera tiputini Erwin & Henry, sp. n., ADP135845 E Hyboptera viridivittis Chaudoir, ADP007601. Legend, lp, left paramere; rp, right paramere; pb, phallobase; bo, phallobase orifice; ps, phalloshaft; a, phalloapex. Scale line = 0.25 mm
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