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Madecorphnus cuccodoroi Frolov 2010
No full textMadecorphnus cuccodoroi Frolov, 2010 (Figs. 24 A–F) Madecorphnus cuccodoroi Frolov, 2010a: 64. Type material examined. Holotype (Figs. 24 A, C–E), male, “ MADAGASCAR, Mt Ambre, 3 km NE N.P. camp site, 12°33'09''S, 49°09'40''E, 1300 m. 28.II.2003 #3a G. Cuccodoro / Holotypus Madec. cuccodoroi Frolov 2010 ” (MHNG). Paratypes: one male, “ Madagascar, Mt Ambre, 2,5 km SE N.P. camp site, 12°31'12''S, 49°11'42''N, 1050 m. 11.III.2003, G. Cuccodoro ” (MHNG); one male, “ Madagascar, Mt Ambre, 1 km NW N.P. camp site, 12°31'55''S, 49°10'32''N, 1000 m. 2.III.2003, G. Cuccodoro ” (MHNG); two males, “ Madagascar, Mt Ambre, 3 km NE N.P. camp site, 12°32'30''S, 49°10'02''N, 1250 m. 15.III.2003, G. Cuccodoro ” (MHNG, ZIN); one female, “ Madagascar, Mt Ambre, 2,5 km SE N.P. camp site, 12°31'15''S, 49°11'40''N, 950 m. 03.III.2003, G. Cuccodoro ” (MHNG); two females, “ Madagascar, Mt Ambre, 0,5 km NE N.P. camp site, 12°31'50''S, 49°10'25''N, 1000 m. 0 1.III.2003, G. Cuccodoro” (MHNG, ZIN). Additional material examined. MADAGASCAR: Antsiranana: six males, “ MADAGASCAR: Province d'Antsiranana, Parc National Montagne d'Ambre 3.6 km 235° SW Joffreville Elev 925 m 20–26 Jan 2001 / 12°32′4″S, 49°10′46″E coll. Fisher, Griswold et al. California Acad. of Sciences, sifted litter, montane rainforest, code: BLF2564” (CASC). Diagnosis. This species is externally similar to M. peyrierasi Frolov, 2010 and M. perinetensis Frolov, 2010 and can reliably be separated from them only by the shape of the parameres and, especially, the internal sac armature. In M. peyrierasi and M. perinetensis, the internal sac armature consists of 2 larger sclerites basally; 2 smaller, comma-shaped sclerites apically; and an area of numerous minute spinules situated between the sclerites (Figs. 26 D–E, 27 D–E). In M. cuccodoroi, the 2 larger sclerites are probably homologous to those in the previous species, but they differ slightly in shape. The area composed of numerous minute spinules is larger in M. cuccodoroi, the spinules are somewhat coarser, and a part of the apical spinules is more robust and they are almost tooth shaped (Fig. 24 C). Madecorphnus cuccodoroi also differs in the shape of the parameres, which are somewhat wider and more curved at apices (Figs. 24 D, 26B, 27B). Description. Male. Body length 5.0– 5.5 mm. Color uniformly dark brown, almost black, legs somewhat paler (Fig. 24 A). Right mandible slightly longer than left, without tooth behind apex. Labrum trapezoidal, with slightly rounded sides, length about 1/6 width (in dorsal view). Frontoclypeus slightly asymmetrical, apically obtuse, with 2 long and 2 shorter setae on the apical margin. Frontoclypeus slightly depressed apicomedially. Head finely punctate with minute punctures separated by greater than 4 times their diameter. Pronotum approximately 1.5 times wider than long, widest medially. Disc of pronotum convex, without any depressions, tubercles, or ridges. Punctation on pronotum finer than on head, almost indistinct. Margins with relatively wide border, lateral margins with 4 long setae: 1 seta on basal angle, 1 seta approximately in the middle of lateral margin, and 2 setae on the apical angle. Scutellum triangular, angulate apically, about 1/12 length of elytra. Elytra convex, with distinct humeral and apical, widest at basal third. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra sparsely punctate with relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytron with border from scutellum to humeral callus. Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex of protibia with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape to each other. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and as long as 2 tarsomeres in metathoracic legs. Parameres with small teeth laterally, narrowly rounded apically in dorsal view and curved downwards in lateral view (Fig. 24 D). Internal sac with 2 long sclerites and a large number of small spinules; some spinules more robust and form 2 indistinctly separated clusters in apical part (Fig. 24 C). Female. Females (Fig. 24 B) differ from males in having the apical spur of the protibiae that is slightly longer and more robust than apical seta in males. Variation. Body length of the examined specimens varies from 5.0– 5.5 mm. Left and right mandibles of some males are almost equal in length and similar to those in females. Distribution and habitat. All specimens were collected in the Amber Mountain National Park in Antsiranana Province (northern Madagascar, Fig. 24 F). The park occupies the Madagascan northernmost remnant of indigenous humid forest covering about 200 km 2. All specimens were collected by sifting forest litter. The number of the collected specimens suggests that the population density of M. cuccodoroi is low at least during the season when the collecting was done.Published as part of Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, pp. 1-93 in Zootaxa 4207 (1) on pages 48-50, DOI: 10.11646/zootaxa.4207.1.1, http://zenodo.org/record/20044
Madecorphnus brunneus Frolov 2010
No full textMadecorphnus brunneus Frolov, 2010 (Figs. 25 A–F) Madecorphnus brunneus Frolov, 2010c: 1108. Type material examined. Holotype (Figs. 25 A–E), male, “ANDRINGITRA Est Anjavidilava 1850–1950 m 18- XII / 15-I-1971 / Holotypus Madecorphnus brunneus Frolov 2009 ” (MNHN). Diagnosis. Madecorphnus brunneus is similar to M. perinetensis and M. peyrierasi but differs from them in having the parameres less acute in lateral view and in the different internal sac armature. Description. Male. Body length 6.2 mm. Color brown, pronotum and head dark brown (Fig. 25 A). Right mandible about 2 times longer than left, without tooth basad of apex. Labrum trapezoidal, its length about 1/5 width (in dorsal view). Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long setae. Frontoclypeus slightly depressed in the middle anteriorly, finely punctate. Pronotum 1.9 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, minutely punctate. Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra with sparse punctation. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral callus. Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and as long as 2 tarsomeres in metathoracic legs. Parameres acutely rounded in lateral view (Fig. 25 B). Internal sac of the aedeagus with 1 longer sclerite and 1 smaller, somewhat bifurcated sclerite (Figs. 25 D–E). Female. Unknown. Distribution and habitat. Madecorphnus brunneus is known from one locality in the Adringitra Massif, southeastern Madagascar (Fig. 25 F).Published as part of Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, pp. 1-93 in Zootaxa 4207 (1) on page 50, DOI: 10.11646/zootaxa.4207.1.1, http://zenodo.org/record/20044
Madecorphnus peyrierasi Frolov 2010
No full textMadecorphnus peyrierasi Frolov, 2010 (Figs. 27 A–F) Madecorphnus peyrierasi Frolov, 2010c: 1109. Type material examined. Holotype (Figs. 27 A–E), male, “ MADAGASCAR EST PERINET XI.1972 A.PEYRIERAS / Holotypus Madec. peyrierasi Frolov 2009 ” (MNHN). Paratypes: one female with the same data as the holotype (MNHN); one female, “ Madagascar East Perinet A.Peyrieras X-1972 ” (MNHN); one male, “La Mandraka A. Peyrieras 25-III-1975 ” (MNHN); one male, “ Madagascar Est route d'Anosibe A. Peyrieras 6.IV.1975 ” (MNHN). Diagnosis. This species is similar to M. perinetensis externally and in the shape of the internal sac armature, but can be separated from it by the different shape of the parameres and somewhat smaller eyes (Figs. 27 A–B). Description. Male. Body length 4.6–6.2 mm. Color uniformly brown, head slightly darker (Fig. 27 A). Right mandible about 1.7–1.2 times longer than left, without tooth behind apex. Labrum trapezoidal, length about 1/6 width (in dorsal view). Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long and 2 shorter setae on the anterior margin. Frontoclypeus slightly depressed in the middle anteriorly, finely punctate. Pronotum 1.8 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, minutely punctate. Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra with sparsely punctate with relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral callus. Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and as long as 2 tarsomeres in metathoracic legs. Parameres tapering apically, acute in lateral view (Fig. 27 B). Internal sac with 2 longer and 2 smaller sclerites (Figs. 27 D, E). Female. Females differ from males in having short subsymmetrical mandibles, long protibial spur, and wider elytra. Distribution and habitat. Madecorphnus peyrierasi is known from one locality in Adringitra Massif, southeastern Madagascar (Fig. 27 F).Published as part of Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, pp. 1-93 in Zootaxa 4207 (1) on page 54, DOI: 10.11646/zootaxa.4207.1.1, http://zenodo.org/record/20044
Madecorphnus hanskii Frolov 2012
No full textMadecorphnus hanskii Frolov, 2012 (Figs. 23 A–F) Madecorphnus hanskii Frolov, 2012b: 160. Type material examined. Holotype (Figs. 23 A–E), male, “ Madagascar Ambohitantely [18°10' S, 47°17' E] 24.3.2005, wet forest, I. Hanski-group leg. / Holotypus Madec. hanskii Frolov 2009 ” (ZIN). Diagnosis. Madecorphnus hanskii is similar to M. falciger and M. punctatus in having internal sac of the aedeagus with 3 sclerites: 1 relatively long, spur-like, and 2 comma-shaped or somewhat bifurcate (Figs. 16 E–F, 22D–E, 23D–E). It can be separated from these species by the shape and relative size of these sclerites and, from the latter, by the shape of parameres and sparse punctation of head and pronotum. Description. Male. Body length 5.3 mm. Color uniformly brown (Fig. 23 A). Right mandible as long as left, without tooth behind apex. Labrum trapezoidal, with broadly rounded sides, length about 1/6 width (in dorsal view). Frontoclypeus slightly asymmetrical, apically obtuse, with 2 long and 6 shorter setae on the apical margin. Canthus and frontoclypeus almost flat, minutely depressed apicomedially. Head without traces of frontoclypeal suture, finely punctate with minute punctures separated by greater than 5 times their diameter. Pronotum approximately 1.6 times wider than long, widest medially. Disc of pronotum convex, without any depressions, tubercles, or ridges. Punctation on pronotum similar to that on head. Margins with relatively wide border, lateral margins with 4 long setae: 1 seta on basal angle, 1 seta approximately in the middle of lateral margin, and 2 setae on the apical angle. Scutellum triangular, right angled apically, about 1/13 length of elytra. Elytra convex, with distinct humeral humps, widest at basal third, glabrous. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra with a few relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytron with border. Wings fully developed. Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Protibial apex with robust, spurlike seta and 2 smaller setae basally. Mesothoracic and metathoracic legs similar in shape to each other, metathoracic legs somewhat longer. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and longer than 2 basal tarsomeres in metathoracic legs. Parameres with teeth laterally, narrowly rounded apically in dorsal view and curved downwards in lateral view (Fig. 23 B). Internal sac with a long, spur-like sclerite and 2 smaller, somewhat comma-shaped sclerites (Figs. 23 D– E). Female. Unknown. Distribution and habitat. The type specimen of M. hanskii was collected in wet forest in Ambohitantely special reserve (Fig. 23 F). The reserve is situated at altitudes of about 1500 m and houses the last remnants of indigenous forest on the central plateau of Madagascar. Madecorphnus hanskii is apparently a forest litter dweller and, due to severe deforestation of the central plateau, its current range may be limited to this small forest remnant.Published as part of Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, pp. 1-93 in Zootaxa 4207 (1) on pages 46-47, DOI: 10.11646/zootaxa.4207.1.1, http://zenodo.org/record/20044
Madecorphnus perinetensis Frolov 2010
No full textMadecorphnus perinetensis Frolov, 2010 (Figs. 26 A–F) Madecorphnus perinetensis Frolov, 2010c: 1108. Type material examined. Holotype (Figs. 26 A–E), male, “ Madagascar East Perinet A.Peyrieras XI-1972 / Holotypus Madec. perinetensis Frolov 2009 ” (MNHN). Paratypes: one male and two females with the same data as the holotype (MNHN); one male “ Madagascar East Perinet A.Peyrieras X-1972 ” (MNHN). Additional material examined. MADAGASCAR: Toamasina: male, “ Madagascar Est: Analamazoatra 6- 11.I.2011 Ballerio leg.” (ABCB). Diagnosis. This species is similar to M. peyrierasi externally and in the shape of the internal sac armature, but can be separated from it by the different shape of the parameres and somewhat larger eyes (Figs. 26 A–B). Description. Male. Body length 5.5–5.8 mm. Color uniformly brown (Fig. 26 A). Right mandible about 2 times longer than left, without tooth behind apex. Labrum trapezoidal, length about 1/ 6 width (in dorsal view). Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long and 2 shorter setae on the anterior margin. Frontoclypeus slightly depressed in the middle anteriorly, finely punctate. Pronotum 1.8 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, minutely punctate. Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra sparsely punctate with relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral callus. Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and as long as 2 basal tarsomeres in metathoracic legs. Parameres less tapering apically than in the previous species, wider in dorsal view, and acute in lateral view. Internal sac with 2 longer and 2 smaller sclerites (Figs. 26 D–E). Female. Females differ from males in having short subsymmetrical mandibles, long protibial spur, and wider elytra. Distribution and habitat. Madecorphnus perinetensis is known from a small area near Andasibe, eastern Madagascar, in primary rain forest habitat (Fig. 26 F).Published as part of Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, pp. 1-93 in Zootaxa 4207 (1) on pages 50-53, DOI: 10.11646/zootaxa.4207.1.1, http://zenodo.org/record/20044
Madecorphnus simplex Frolov 2010
No full text<i>Madecorphnus simplex</i> Frolov, 2010 <p>(Figs. 19 A–F)</p> <p> <i>Madecorphnus simplex</i> Frolov, 2010c: 1106, Frolov 2013a: 5.</p> <p> <b>Type material examined.</b> Holotype (Figs. 19 D, E), male, “ MADAGASCAR CENTRE AMBATOFITORAHANA 1800 M F D H M / 31.XII.1972 / A. PEYRIERAS / Holotypus <i>Madec.</i> <i>simplex</i> Frolov 2009 ” (MNHN).</p> <p> <b>Additional material examined.</b> Male, “ Madagascar, Fianarantsoa Prov., Foret d'Atsirakambiaty, 7.6 km 285° WNW Itremo, elev 1550 m., 22–26 January 2003 / 20°35′36″S, 046°33′48″E colls. Fisher Griswold <i>et al.</i> Calif. Acad. of Sci. pitfall trap—montane rainforest collection code BLF7150” (CASC); male, “ MADAGASCAR: Province Fianarantsoa, Parc National Ranomafana, at broken bridge, elev 1110 m 19–26 March 2002 / 21°13′57″S, 47°22′19″E collector R. Harin'Hala California Acad. of Sciences, malaise trap in high altitude rainforest, MA-02- 09A-21” (CASC).</p> <p> <b>Diagnosis.</b> This species can be separated from other <i>Madecorphnus</i> species by the distinct shape of its parameres and the internal sac armature.</p> <p> <b>Description. Male.</b> Body length 5.5 mm. Color reddish brown (Fig. 19 A).</p> <p>Right mandible 1/5 longer than left, without tooth behind apex. Labrum trapezoidal, length about 1/4–1/5 width (in dorsal view).</p> <p>Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long and a number of smaller setae. Frontoclypeus slightly depressed in the middle anteriorly, sparsely punctate (punctures separated by about 5 puncture diameters).</p> <p>Pronotum 1.8 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, sparsely punctate.</p> <p>Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral humps. Elytra sparsely punctate except for a few relatively large punctures.</p> <p>Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally.</p> <p>Mesothoracic and metathoracic legs similar in shape. Longer tibial spur as long as 2 basal tarsomeres in mesothoracic legs and shorter than 3 tarsomeres in metathoracic legs. Metafemora with a few pectinate setae on the inner side apically.</p> <p>Parameres rounded apically (in lateral view), with small teeth laterally (Fig. 19 B). Internal sac with 1 sclerite composed of 2 long slender spinules connected basally (Figs. 19 D–E).</p> <p> <b>Female.</b> Unknown.</p> <p> <b>Distribution and habitat.</b> <i>Madecorphnus simplex</i> is known from 3 localities in central and eastern Madagascar (Fig. 19 F).</p>Published as part of <i>Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, pp. 1-93 in Zootaxa 4207 (1)</i> on page 40, DOI: 10.11646/zootaxa.4207.1.1, <a href="http://zenodo.org/record/200444">http://zenodo.org/record/200444</a>
Eduard Frolov, the Outstanding St. Petersburg Ancient Historian
Get PDFThe article reveals the most important results of the scientific work of the outstanding researcher
of Ancient history and culture, Doctor of historical sciences, Professor Eduard Davidovich
Frolov (1933–2018) as one of the most prominent representatives of the St. Petersburg
school of Classics. The scientific work of Ed. D. Frolov is shown as a very important
stage in the development of the St. Petersburg school of Classics. The scholarly writings by
Ed. D. Frolov, dealing with the political history of Ancient Greece, social thought, Ancient
culture, Russian historiography of Antiquity, are fully and thoroughly described in this article.
The scientific works of Ed. D. Frolov covers the number of the important problems, such as the
political history of Ancient Greece, social thought, Ancient culture, Russian historiography of
Antiquity. Ed. D. Frolov has investigated the formation of the polis in Archaic Greece in the
8th — 6th centuries BC, the problem of the crisis of a political organization in the 4th century
BC, the place and role of tyranny in Ancient Greek history and others. All his scholarly works
have always been carried out with exhaustive completeness, on the basis of brilliant knowledge
of the entire mass of sources. The scientific activities of Ed. D. Frolov have resulted in
more than 350 publications. Among them are 15 monographs, many chapters in collective
monographs, translations from Ancient Greek and German, many articles and reviews. Finally,
Ed. D. Frolov has created his own scientific school, having brought up a large number of
pupils — Candidates and Doctors of science
Madecorphnus dentatus Frolov 2010
No full textMadecorphnus dentatus Frolov, 2010 (Figs. 17 A–F) Madecorphnus dentatus Frolov, 2010c: 1105. Type material examined. Holotype (Figs. 17 A–E), male, “ Chaînes Anosyennes, S.O. du Trafonaomby plateau Andohahelo 1770–1950 m, V-1972 / MUSEUM PARIS Madagascar Est mission C.N.R. S. R. C.P. n°225 / Holotypus Madec. dentatus Frolov 2009 ” (MNHN). Paratypes: two specimens, female with the same data as the holotype and male, “ Madagascar Est: N.- O. de Fort-Dauphin, massif de l'Andohahelo, forêt d'Andranomangara, R.N.I. n 11, 1750 m, 20/ 25-I-1974 A. Peyrieras ” (MNHN). Diagnosis. This species can be separated from other Madecorphnus species by the distinct shape of the parameres and internal sac armature. Description. Male. Body length 6.0 mm. Color reddish brown, head, legs, and apices of elytra somewhat darker. Right mandible slightly longer than left, without tooth behind of apex. Labrum trapezoidal, length about 1/4–1/ 5 width (in dorsal view). Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long and a number of smaller setae. Canthus and frontal suture indistinct. Frontoclypeus somewhat depressed in the middle anteriorly, sparsely punctate. Pronotum 1.8 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, punctate (punctures separated by 2–3 puncture diameters). Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra with sparse punctures. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral callus. Elytra minutely punctate except for a few relatively large punctures in the middle of each elytron near stria 1. Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape. Longer tibial spur as long as 2 basal tarsomeres in mesothoracic legs and as long as 3 basal tarsomeres in metathoracic legs. Aedeagus. Parameres symmetrical, tapering apically in lateral view, with large teeth laterally (Fig. 17 B). Internal sac of aedeagus with 1 sclerite composed of 2 long slender spinules connected basally and with 3 small spinules (Fig. 17 D–E). Female. Except for shorter mandibles, females differ from males in having 2 setae on hind angles of pronotum and in robust protibial spur. Variation. Male paratype has shorter mandibles similar to those in female. Distribution and habitat. Madecorphnus dentatus is known from two localities in southeastern Madagascar (Fig. 17 F).Published as part of Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, pp. 1-93 in Zootaxa 4207 (1) on pages 35-38, DOI: 10.11646/zootaxa.4207.1.1, http://zenodo.org/record/20044
Metacatharsius janssensi Frolov 2014, comb. nov.
No full text<i>Metacatharsius janssensi</i> (Frey), comb. nov. <p>(Figure 9E)</p> <i>Delopleurus janssensi</i> Frey, 1963 <p> <i>Delopleurus janssensi</i> was described from two specimens from Warder, Somali region of Ethiopia. The holotype, female, has all the diagnostic characters of the genus <i>Metacatharsius</i> Paulian (large dorsal eye parts can be seen on the habitual photo, Figure 9E). The paratype is similar to and apparently conspecific with the holotype. In the original description, the author compared this species with <i>D. pullus</i> and <i>D. gilleti</i> but did not provide characters specific for the genus <i>Delopleurus.</i> However, the original description and locality data agree well with the type specimens. There is no doubt that <i>Delopleurus janssensi</i> was described from misidentified <i>Metacatharsius</i> specimens and the new generic combination is therefore established here.</p> <p> It is possible that <i>M. janssensi</i> is conspecific with another described species of the genus. There are up to 18 species of this genus known from adjacent Somalia (Ferreira 1972) and the genus is currently a subject of taxonomic revision (François Génier, pers. comm.). It is out of the scope of the present work to specify the taxonomic status of <i>M. janssensi</i> more exactly.</p> <i>Type material</i> <p>Holotype, female with the labels: ‘ Wardere Somalia’, ‘Holotype’, ‘TYPE’, ‘Brit. Mus 1963-277’, ‘ Delopleurus janssensi det. G. Frey 1963, n. sp. Typus’ (BMNH) and paratype with the labels: ‘Wardere Somalia Oct. 1943, T.H.E.Jackson’, ‘ Delopleurus janssensi det. G. Frey 1963, p. Typus’, ‘Cotype’ (NHMB).</p>Published as part of <i>Frolov, Andrey V., 2014, Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa, pp. 129-154 in Journal of Natural History 49 (3)</i> on pages 152-153, DOI: 10.1080/00222933.2014.909072, <a href="http://zenodo.org/record/4002597">http://zenodo.org/record/4002597</a>
Triodontus inexpectatus Frolov, Montreuil & Akhmetova, new species
No full textTriodontus inexpectatus Frolov, Montreuil & Akhmetova, new species (Figs. 46 A–F) Type material. Holotype (46 A–E), male, “ MADAGASCAR E. Masoala, 50–470 m R. Antsamanarana 3–7/xii/ 1993, Flight Intercept Trap BMNH(E) 1994-138 / Holotypus Triodontus inexpectatus Frolov et al. 2010 ” (BMNH). Diagnosis. Triodontus inexpectatus is most similar to T. hanskii but can be separated from it in having inner lobes of the parameres relatively larger (in lateral view) and outer lobes as long as the inner lobes. Description. Male. Body length 12.5 mm, width 6.5 mm. Body strongly shiny, uniformly colored (Figs. 46 A– B). Color of head, pronotum, and elytra dark brown; legs, antennae, and underside of body brown. Frontoclypeus convex anteriorly, rounded laterally, anterior margin crenulate and setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small, dorsal and large, ventral parts. Frontoclypeus with tubercles mediad of each eye and with short obtuse horn in center of frontoclypeus. Pronotum convex, without tubercles or ridges, with shallow concavity in the middle. Lateral margins with wide border appearing somewhat crenulate in dorsal view. Anterior margin with wide, smooth border. Posterior margin with fine border; not crenulate; punctate with narrow, longitudinal punctures laterally. Surface of pronotum mainly smooth, anterior angles with a few coarse punctures. Scutellum rounded apically, visible part is about 1/15 length of elytra. Elytra convex, with distinct humeral humps. Maximum width approximately at middle. Elytra with 10 feebly visible striae and with oblique line from base of 6th stria to approximately middle of 8th stria. Stria marked with small, indistinct punctures. Intervals with minute punctures, almost smooth. Base of elytra with an irregular row of coarse, semicircular punctures each bearing small, yellow seta. Sternite 8 medially about 2 times as short as other sternites combined, slightly sinuate apically, without tubercles or concavities in the middle (Fig. 46 E). Parameres with apices of upper lobes being obtusely rounded, strongly sclerotized; upper lobes deeply sinuate laterally, the sinuation forms a tooth in dorsal view (Fig. 46 D). Internal sac of aedeagus with a somewhat pentagonal main sclerite. Female. Unknown. Distribution. The species in known from a single specimen collected in the Masoala Peninsula (Fig. 46 F). Etymology. From Latin, “ inexpectatus ” for unexpected. This name is an adjective in the nominative singular.Published as part of Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, pp. 1-93 in Zootaxa 4207 (1) on pages 88-91, DOI: 10.11646/zootaxa.4207.1.1, http://zenodo.org/record/20044
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