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    Platytropesa Macquart 1851

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    Genus Platytropesa Macquart The genus was known from 3 very similar species variously distributed in the Moluccas, New Guinea, the Bismarck Archipelago and North Queensland. One more of these is here recorded from Australia.Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 98, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species

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    Figure 5. Distribution of Amenia imperialis and close relatives (overlapping records omitted). Filled circles = A. imperialis; open squares = "arid form"; filled triangles = Amenia albosquamata.Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 90, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Paramenia angustifrons Crosskey 1965

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    Paramenia angustifrons Crosskey I have seen two of the the three specimens from Highvale (14 miles NW of Brisbane), Queensland, that Crosskey (1965) thought might represent a new species, largely on the basis of head shape. Using my own system of measurement (which I believe to be more accurate) I find little difference between these specimens and typical Paramenia angustifrons, with which they cluster closely in a PCA ordination (not shown). The range of the species is shown in Fig. 27. It seems restricted to the tropics and subtropics.Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 113, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Amenia longicornis

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    Amenia longicornis (Malloch) As showninPart 1, the species comprises two fairly distinct geographic forms: an eastern one, found around the head of Spencer Gulf and north through the Flinders Ranges to the MacDonnell ranges of the Northern Territory, and a western one in places west of about 135° longitude. Although qualifying as geographic subspecies, I see no point in naming them formally. Two specimens from the Flinders Ranges were reared from snails of the species Sinumelon hawkerana Solem (Camaenidae) (near Jonburgh, SouthAustralia, M. Newton, 27 May 1992)Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 111, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Paraplatytropesa rieki

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    Paraplatytropesa rieki (Paramonov) Crosskey did not mention the profuse setulae of the propleuron (not present in some other species). Also, the abdominal sternites have profuse long hairs, not just "a few". The male genitalia are shown in Fig. 12; note the veryshortsurstyles, whichbarelyprojectbeyondthelateral lobes ofT9. I have two additional specimens, also from central Queensland (Bundaberg, March 1973, and Bin Bin Range, west of Biggenden, December 1974, both colI. H. Frauca). Distribution (Fig. 26, p. 108). Queensland: Palm Island, Springsure, Collinsville, Bundaberg, Biggenden.Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 100, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Amenia albomaculata Colless 1998, rev.stat.

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    Amenia albomaculata (Macquart) rev.stat. Amenia leonina albomaculata, Crosskey, 1965: 115. There is no difficulty in identifying the species, using the combination of characters given in the key. The submedian bristles on abdominal T3 alone serve for all specimens of A. albomaculata seen to date. Of the remaining male attributes, the semi-erect setulae on T3 (Fig. 25) are also reliable, as are the others if the "New England " form (see below) is excluded. Also of some use is the not infrequent reduction or absence of the presutural silvery patch (on the suture, mesad of the presutural bristle), which seems usually present in leonina. I have three enigmatic males from the vicinity of Brisbane that resemble the "New England " form (see below) in having the parafrontal setae as in leonina, but the postorbits are intermediate between that form and the typical one. Their status remains unclear.Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 111, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Amenia Robineau-Desvoidy 1830

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    Genus Amenia Robineau-Desvoidy Formosiomima Enderlein, 1936: 444. Type species: Formosiomima imitatrix Enderlein, 1936: 444, (= Amenia nigromaculata Malloch, 1929), by original designation. Despite the considerable differences in ornamentation between Malloch's nigromaculata and other species of Amenia, the structural differences are not great. I am therefore reluctant to follow Crosskey in accepting the generic status proposed by Enderlein (see below) and therefore place his Formosiomima as a junior synonym. Despite much further collecting, Amenia has been found (as reported by Crosskey, 1965) only in Australia, especially Queensland and New South Wales; it is present in the Torres Strait islands, but absent from Tasmania. It is sharply separated from other genera by the characters given in the key. See also the note on setulae of the suprasquamal ridge under Paramenia (below). In descriptions of species, I have confined myself to salient, diagnostic features, rather than repeat the detail provided by Crosskey (1965).Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 105, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Amenia chrysame

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    Amenia chrysame (Walker) Like the several preceding species, A. chrysame has a wide distribution in eastern Australia (Fig. 10). To test whether it, too, might display significant geographic variation, a preliminary analysis was made of 6 measurements and 4 bristle characters on a small set of specimens. This suggested that frons-width (Frw, as defined above) does, indeed, vary geographically. The ratio FrwlPdc was therefore measured on 100 specimens, selected to cover the species' range. The ratio proved to have a highly significant regression on latitude (p<O. OI). However, a scatter-plot (Fig. 11) shows that the regression reflects, not a regular c1ine, but an abrupt shift in average frons width at about 26 degrees south. Separate analyses of specimens from north and south of the 26th. parallel show no significant regression on latitude. However, the mean frons ratios were 0.27 and 0.20 respectively-a difference that, although small, is statistically highly significant (p<O.OI for t-test). On present evidence, I would interpret this as simple geographic variation; but the reason for its occurrence is not at all obvious. Perhaps it reflects some ancient physiographic discontinuity that has long since disappeared-indeed, perhaps the same one that gave rise to A. leonina and A. albomaculata. The matter is further discussed in Part 3. Figure 11 shows several other interesting features: (a) a deficiency of specimens with relatively narrow frons in places between 28 and 31 degrees south; (b) an apparent surplus of such specimens in places between 26 and 28 degrees south-i.e., in the general vicinity of Brisbane; (c) a deficiency of specimens with relatively wide frons in places between 31 and 35 degrees south. If these represent more than fortuitous statistical fluctuations, I am at a loss to explain them. (midline marked in degrees). PART 2. FORMAL TAXONOMY In this section I formalise the taxonomic decisions foreshadowed in the section on morphometrics and also describe several new species that have come to hand, together with new records and occasional notes on other species. "Specimens examined" are listed only for new species and diagnoses are left to be inferred from the keys. Original descriptions and synonymies are not cited, since they were more than adequately handled by Crosskey (1965, 1969, 1972) and have been recently collated by Kurahashi (1989).Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on pages 93-95, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Paraplatytropesa grandicornis Colless 1998, n.sp.

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    Paraplatytropesa grandicornis n.sp. Holotype: male; 14°49'S 125°50'E, Mining Camp, Mitchell Plateau, WesternAustralia, 9-19 May 1983, LD. Naumann & J. Cardale, ex ethanol. ANIC No. 6152. Other specimens examined. WESTERN AUSTRALIA: 1 0, 2¥ ¥, as for holotype; 14°35'S 125°45'E, Lone Dingo, Mitchell Plateau, Western Australia, 9-19 May 1983, LD. Naumann & J. Cardale, 10. Generally resembling P. rieki, with significant attributes as follows: Male. Head (Figs 20, 21). Carina short, narrow, not confluent with epistome, not protruding past antennal segment 3 in lateral view, dark brown with light silvery pollinosity. Facial ridge with setulae only close to vibrissa. Frons relatively broad as compared with head width, but rather short (Table 3), with 2 strong proclinate frontoorbitals and 1 reclinate. Antennal segment 3 short, only as long as that of female. Vibrissae set a little above level of epistome margin, conspicuously long and strong, incurved, decussate. Genae, parafacials, parafrontals, and postorbits with silver pollen; gena conspicuously short, truncate (Table 3). Thorax. Propleuron with several setulae. Scutellar setulae as for P. tropicalis. Lower calypter with exposed portion dark. Lappet hairs of posterior spiracle 1-2 in number, inconspicuous. Legs. Fore tibia without p seta. Mid tibia with 2-3 strong a setae. Postero-dorsal strip of hind coxa setulose. Wing. Setulae on basal node of R 3+4 extending half or more the distance to r-m (in other species, largely confined to the swollen base of the vein). Abdomen. T 1+2 dark purplish; T 3 metallic blue with median dark vitta and apical dark band, and conspicuous, pale silvery lateral patches (in posterior view) extending almost to midline; T 4 similarto T 3, but pale patches less conspicuous or absent; T 5 metallic purplish. S3 with only 4-6 long hairs, S4 without any. Genitalia practically identical with those of P. tropicalis. Female. Generally similar to male. Distribution (Fig. 26, p. 108). Known only from the Kimberleys, Western Australia.Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 103, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288

    Amenia dubitalis Malloch 1927, rev.stat.

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    Amenia dubitalis Malloch rev.stat. Amenia dubitalis Malloch, 1927: 343. Amenia imperialis dubitalis Crosskey, 1965: 111. Differs from A. imperialis in the attributes given in the key. The narrow frons of the male is particularly striking and in both sexes the difference in facial pollinosity, while seemingly rather trivial, is conspicuous under the stated conditions of lighting. For both sexes the M-bend index is 0.7-1.2; the antennal segment ratio is 2.2-3.1 for males and 2.4-2.6 for females. In the one male dissected the surstylus had a subbasal group of 3-4 bristles on the anterior surface, as opposed to a single bristle for other members of the group. I have one new rearing record: a male (24 km SE of Bonshaw, NSW, 31 March 1990, PH. Colman; AM) from Strangesta sp. (Rhytididae). Distribution. Eastern Australia, from about Ingham in the north to southeastern Victoria; restricted almost entirely to the coast and adjacent ranges (Fig. 3) A close examination of records shows a marked deficiency from the more inland localities of southeastern Queensland (Fig. 4), as compared with A. imperialis (Fig. 6). The area is well-collected and the difference is clearly significant, presumably reflecting some difference in host snails. Notes. The difference between females from the border area between Queensland and New South Wales and further south, which almost invariably have submedian marginal bristles on abdominal T3, and those from further north, which almost invariably lack them, could well be used to establish two separate subspecies. However, I see no point in taking this step. The change is quite abrupt, at about the latitude of Brisbane: between there and the 25th. parallel, six out of 13 specimens had such bristles, and furthernorth, only one out of 21 (and that very small). Per contra, only two out of 47 specimens from more southern localities (Sydney and Narrabri) lacked such bristles, two had only a single bristle, and two had three bristles.Published as part of Colless, D. H., 1998, Morphometrics in the genus Amenia and revisionary notes on the Australian Ameniinae (Diptera: Calliphoridae), with the description of eight new species, pp. 85-123 in Records of the Australian Museum 50 (1) on page 110, DOI: 10.3853/j.0067-1975.50.1998.1275, http://zenodo.org/record/465288
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