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Lycoriella acutostylia Mohrig & Menzel 1990
<i>Lycoriella acutostylia</i> Mohrig & Menzel, 1990 <p>Figs 4A, 13B, 14A</p> <p> <b>Literature</b>. <i>Lycoriella</i> (<i>Lycoriella</i>) <i>acutostylia</i> Mohrig & Menzel — Menzel <i>et al.</i> (1990): 340, fig. 7a–c; Menzel & Mohrig (2000): 385, 611. <i>Lycoriella acutostylia</i> Mohrig & Menzel —Menzel <i>et al.</i> (2000): 77; Mukkala <i>et al.</i> (2005): 7, 16, 32; Heller <i>et al.</i> (2009): 41; Vilkamaa & Menzel (2019): 51.</p> <p> <b>Material studied</b>. CZECH REPUBLIC, Bílina, Vétrak, 50°33’29”N, 13°45’57”E, dump restoration SW, 200 m, 8–10. VI.1997, M. Barták, 1 male (in SDEI); same data but 9–23.IV.1997, 2 males (in SDEI). FINLAND, Lkor (Lapponia kemensis orientalis), Pelkosenniemi, Kemihaara, Sadinvaara (Grid 7445310:3537269), Malaise trap, 31. VII –29.IX.2015, J. Salmela, 3 males (in LMM; specimen IDs JS-COI-2016-0011, JS-COI-2016-0012 and JS-COI-2016-0242); Ks (Regio kuusamoensis), Kuusamo, Matinjärvi (Grid 7367909:3615372), Malaise trap, 1. VII – 3.VIII.2005, J. Salmela, 1 male (in MZH, http://id.luomus.fi/GE.1824); Ab (Regio Aboensis), Karkkila, Iilammi, rich fen, Malaise trap, 6. VI.2004, J. Salmela, 1 male (in LMM).</p> <p> <b>BIN</b>. BOLD:ACV2466.</p>Published as part of <i>Menzel, Frank & Vilkamaa, Pekka, 2021, New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region, pp. 501-530 in Zootaxa 5072 (6)</i> on page 524, DOI: 10.11646/zootaxa.5072.6.1, <a href="http://zenodo.org/record/5751541">http://zenodo.org/record/5751541</a>
Trichocoelina brevicubitalis Vilkamaa & Menzel 2019, comb. n.
Trichocoelina brevicubitalis (Lengersdorf, 1926) comb. n. Literature. Sciara brevicubitalis Lengersdorf—Lengersdorf (1926): 6; Gerbachevskaja-Pavluchenko (1986): 61. Lycoria (Neosciara) brevicubitalis (Lengersdorf) — Lengersdorf (1928 –30): 61. Neosciara brevicubitalis (Lengersdorf) —? Lackschewitz (1934): 155; Soot-Ryen (1942): 77. Lycoriella (Hemineurina) brevicubitalis (Lengersdorf) — Frey (1942): 36; Menzel & Mohrig (2000): 408. Redescription. Female. Head. Dark brown. Eye bridge united, 2–3 facets wide. Maxillary palpus long, with 3 segments, all segments narrow, 1 st segment with 4 sharp setae, with a dorsal unbordered patch of sensilla. Scapus and pedicellus dark bown and roundish; antennal flagellomers missing in the specimen studied. Thorax. Postpronotum non-setose. Scutellum with 8 long and strong setae. Legs. Strong, with long tibial spurs; tibial spurs of mid and hind tibiae unequally long [one spur is about 1/4 shorter than the other]; tarsal claws without teeth. Wing. Large, anal lobe strong, hind veins distinct, stCuA short, 1/3 of bM; bM = r-m, bM non-setose, r-m with 4–5 setae distally, c/w 0.50. Body length 2,7 mm, wing length 2,5 mm. BIN. Unknown. Discussion. The species was described from females from Northern Norway by Lengersdorf (1926): for the lectotype designation and literature, see Menzel & Mohrig (2000). After a study of the lectotype, which so far is the only specimen of the species (in ZFMK) available for study, Menzel & Mohrig (2000) stated that the scutellum has many long and strong setae, the 1 st palpal segment has a unbordered patch of sensilla, the fore tibia has spine-like setae, and that R 1 /R complex is long (R 1 ends near the level of the base of the M-fork). We have studied the lectotype again and have found characters that might be helpful in identification of the male of this species and for supporting its placement in the genus Trichocoelina. Of the characters given above, only the somewhat unequally long spurs of the mid and hind tibiae as well as the high number of long scutellars are unusual for Trichocoelina.Published as part of Vilkamaa, Pekka & Menzel, Frank, 2019, Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species, pp. 1-67 in Zootaxa 4665 (1) on page 19, DOI: 10.11646/zootaxa.4665.1.1, http://zenodo.org/record/400055
Trichocoelina janetscheki Vilkamaa & Menzel 2019, comb. n.
Trichocoelina janetscheki (Lengersdorf, 1953) comb. n. Figs 10 A, 10 B, 17 D Literature. Neosciara janetscheki Lengersdorf—Lengersdorf (1953): 167, fig. 1; Janetschek (1956): 471. Lycoriella (Hemineurina) janetscheki (Lengersdorf) — Tuomikoski (1959a): 35; Gerbachevskaja-Pavluchenko (1986): 31; Franz (1989): 14; Menzel & Mohrig (2000): 410; Wirta et al. (2016): appendix, unpaginated p. 21 (table S1) and unpaginated p. 39 (cladogram). Material studied. CANADA, Nunavut, Sverdrup Islands, Elles Ringnes Island, Isachsen, 70.79°N, 103.55°W, lemming burrow, 17.VI.1960, J.R. Vockeroth, 4 males (3 in CNC, 1 in MZH); same locality but moss near very small stream, 14.VII.1960, J.F. McAlpine, 2 males (in CNC); Nunavut, Ellesmere Island, Fosheim Peninsula, Hot Weather Creek, 79°58’N, 84°28’W, 2.VII.1990, F. Brodo, 1 male (in MZH); NE Greenland, Mestersvig, 72.24°N, 23,92°W, C. Vibe (in ZMUC); NE GREENLAND, Zackenberg, 74°28’N, 20°34’W (UTM8265500:0513778), 44 m, 2–11.VII.2011, T. Roslin & G. Várkonyi, 1 male (in MZH); same locality but UTM 8265758:0513786, 37 m, Malaise trap, 11–20.VII.2011, T. Roslin & G. Várkonyi, 2 males (in MZH, SDEI); same locality but 74.50°N, 21.00°W, 44 m, 7.VII.2011, T. Roslin & G. Várkonyi, 1 male (BOLD Sample ID GRPV16, in DAUH); same locality but 74.4667°N, 20.5667°W, 48 m, 24.VII.2011, T. Roslin & G. Várkonyi, 3 males (BOLD Sample IDs ZA2012- 50010, ZA2012-50011 and ZA2012-50012, in DAUH); USA, Colorado, Mt. Evans, 14 000 ft, 25.VII.1961, W.R.M. Mason, 1 male (in USNM). Redescription. Male. Head. Face brown, antenna dark brown maxillary palpus pale yellowish. Eye bridge 2–3 facets wide. Face with 15–21 setae. Clypeus with 1–4 setae. Maxillary palpus with 3 segments, 1 st segment as long as or longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 3–9 setae, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.95–2.75x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Dark brown, setae pale. Anterior pronotum with 5–11 setae. Proepisternum with 7–24 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.2–3.1 mm. Width/length 0.35–0.40. Anal lobe weak. Veins distinct. c/w 0.50–0.60. R 1 /R 0.50–0.90. stM shorter than fork of M. r-m longer than bM, bM non-setose, r-m non-setose or with 1–2 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large patch in shallow depression. Fore tibial spur as long as the tibial width. Abdomen. Pale brown, setae pale, short and fine. Hypopygium (Fig. 10 B). Brown, as abdomen. Intergonocoxal area long, with two short setose lobes. Gonocoxa broad, as long as gonostylus, medial margin basally smoothly curved, with short and fine setosity. Gonostylus (Fig. 10 A) voluminous, apically truncate, impressed; with short setosity, a short apical tooth, and numerous megasetae in the medial impression; megasetae slightly curved or straight, short and slender; with 1 well-differentiated whiplash seta basally at ventromedial margin. Tegmen (Fig. 17 D) longer than broad, laterally straight, apically with a narrow hyalinous acuminate process, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. BOLD:ACK5495. Discussion. In Fig. 10 A the gonostylus is flattened and distorted to show the arrangement of the megasetae. In its extremely voluminous gonostylus, Trichocoelina janetscheki (Lengersdorf, 1953) resembles most T. incrassata sp. n., for distinguishing characters, see under the latter.Published as part of Vilkamaa, Pekka & Menzel, Frank, 2019, Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species, pp. 1-67 in Zootaxa 4665 (1) on page 33, DOI: 10.11646/zootaxa.4665.1.1, http://zenodo.org/record/400055
Lycoriella eurystylata Menzel & Vilkamaa 2021, sp. n.
Lycoriella eurystylata sp. n. Figs 4B, 6C, 13A Material studied. Holotype male. CANADA, Quebec, Old Chelsey, 3.VIII.1958, J. R. Vockeroth (in CNC). Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 3 facets wide. Face with 14 setae. Clypeus with 4 setae. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 5 setae, with small dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.2x as long as wide, neck slightly shorter than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 5 setae. Proepisternum with 8 setae. Scutellum with 2 long and some short and fine setae. Wing. Fumose. Length 1.7 mm. Width/length 0.45. Anal lobe small. Veins distinct, except for stM. c/w 0.75. R1/R not detectable in the specimen studied. stM shorter than fork of M. bM and r-m subequal in length, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Brown; setae pale. Fore tibial organ forming large patch of many setae in demarcated depression. Fore tibial spur slightly shorter than apical width of tibia. Abdomen. Pale brown; setae pale and moderately long. Hypopygium (Fig. 4B). Brown, like abdomen. Intergonocoxal area rather long, with short setose lobe. Gonocoxa longer than gonostylus, slightly roundish laterally, with rather dense and fine setosity. Gonostylus (Fig. 6C) widest subbasally, narrowed towards apex, strongly impressed medially, densely setose apically, with strong and curved apical tooth; with 5 medial megasetae in apical half, megasetae slender, long and straight, oblique, the apicalmost (with sockets) shorter than the others, as long as apical tooth; with well-differentiated long whip-lash seta in basal third of gonostylus. Tegmen (Fig. 13A) much wider than long, smoothly curved and membraneous apically, straight laterally, sclerotized basolaterally, with small area bearing a few aedeagal teeth. Aedeagal apodeme moderately long. BIN. Unknown. Discussion. Lycoriella eurystylata sp. n. resembles L. taimyrensis sp. n. and L. tundrae sp. n. in having the tegmen smoothly curved apicolaterally, whereas L. acutostylia Mohrig & Menzel, 1990 [described in Menzel et al. (1990)], L. jakovlevi sp. n. and L. kinbasketi sp. n. have the tegmen distinctly angled apicolaterally (Fig. 13). L. eurystylata differs from L. taimyrensis and L. tundrae in having the tegmen very short, strongly sclerotized basolaterally, semicircular apically and without tooth-like structures, whereas the other two have their tegmen only slightly sclerotized laterally, less curved apically and with tooth-like structures. Furthermore, L. eurystylata has five, L. tundrae four and L. taimyrensis eight to nine gonostylar megasetae and the fore tibial organ of L. eurystylata is more distinctly demarcated than that of L. taimyrensis and L. tundrae. L. eurystylata is similar to L. acutostylia, L. jakovlevi and L. kinbasketi in having rather long antennal flagellomeres (4 th flagellomere 2.1–2.3 times as long as wide), whereas L. taimyrensis and L. tundrae have shorter flagellomeres (4 th flagellomere 1.4–1.6 times as long as wide). To distinguish L. taimyrensis and L. tundrae, see under the former, and to distinguish L. acutostylia, L. jakovlevi and L. kinbasketi, see under L. jakovlevi sp. n. Etymology. The name is derived from the Greek words eurys, broad, and stylos, style, referring to the thick gonostylus of the male hypopygium.Published as part of Menzel, Frank & Vilkamaa, Pekka, 2021, New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region, pp. 501-530 in Zootaxa 5072 (6) on pages 507-509, DOI: 10.11646/zootaxa.5072.6.1, http://zenodo.org/record/575154
Trichocoelina vitticollis Vilkamaa & Menzel 2019, comb. n.
Trichocoelina vitticollis (Holmgren, 1883) comb. n. Synonyms: = glacialis (Lundbeck, 1898) [as Sciara; preocc., not Sciara glacialis Rübsaamen, 1898]; = permutata (Lundbeck, 1900) [as Sciara; new name for Sciara glacialis Lundbeck, 1898]. Literature. Sciara glacialis Lundbeck—Lundbeck (1898): 254, pl. 6, fig. 13. Sciara humicola Lundbeck—Lundbeck (1898): 252, pl. 6, fig. 11 [misidentification]. Sciara permutata Lundbeck—Lundbeck (1900): 313. Bradysia permutata (Lundbeck) — McAlpine (1964): 128. Bradysia (Hemineurina) permutata (Lundbeck) — Frey (1948): 66, 84; pl. 18, fig. 106. Lycoriella (Hemineurina) permutata (Lundbeck) — Tuomikoski (1959a): 36; Tuomikoski, (1960): 75, 76; Stone & Laffoon (1965): 232; Tuomikoski (1967): 48; Gerbachevskaja-Pavluchenko (1986): 31 [not sensu Mohrig et al. (1983a) 15; misidentification, = T. subpermutata (Mohrig & Mamaev, 1990)]. Sciara vitticollis Holmgren—Holmgren (1883): 182; Jacobson (1898): 190; Gerbachevskaja-Pavluchenko (1986): 71. Lycoria (Neosciara) vitticollis (Holmgren) — Lengersdorf 1928 –30: 59. Lycoriella (Hemineurina) vitticollis (Holmgren) — Menzel & Mohrig (2000): 411, figs 380, 381; Coulson & Refseth (2004): 103; Coulson (2008): 162; Coulson (2013): 154; Mohrig et al. (2013): 271; Vilkamaa (2015): 551; Wirta et al. (2016): appendix, unpaginated p. 21 (table S1) and unpaginated p. 39 (cladogram). New records. CANADA, Northwest Territories, Mackenzie Delta, Reindeer Depot, 29.VI.1948, J.R. Vockeroth, 1 male (in MZH); NE GREENLAND, Zackenberg, 74°28’N, 20°34’W, 1.VIII.1991, J. Böcher, 1 male (in ZMUC); NW GREENLAND, Thule, 8.VII.1940, C. Wibe, 1 male (in ZMUC); SE GREENLAND, Skoldängen, 14–27.VII.1992, 2 males (1 in MZH, 1 in ZMUC); SW GREENLAND, Nugssuak, 18.VII.1949, C. Wibe, 1 male (in ZMUC); Saputit, 6.VII.1949, C. Wibe, 1 male (in MZH); W GREENLAND, Arnangarnup kua, 11.VII.1984, J. Böcher, 1 male (im ZMUC); Skjoklungen, ‘ Bygder’, 19–27.VII.1992, S. Andersen, 1 male (in ZMUC); Söndre Stormfjord, 21.VI.1952, C. Wibe, 1 male (in MZH); same locality but 67°02’N, 50°40’W, 2.VIII.1992, J. Böcher, 1 male (in MZH); NORWAY, ‘ Svalbard, Bjorndalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Bjørndalen W of Adventfjorden], 78.2320°N, 15.3270°E, 13.VII.2012, G.E.E. Søli, 1 male (NHMO 261403 and BOLD Sample ID SV985, in NHMO); ‘ Svalbard, Bolterdalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Bolterdalen on the southern side of Adventdalen], 78.1640°N, 15.9900°E, 100 m, 10.VII.2012, G.E.E. Søli, 3 males (NHMO 260984 and BOLD Sample ID SV1061, NHMO 260985 and BOLD Sample ID SV1062, 2 in NHMO; BOLD Sample ID SV1064, in SDEI); ‘Svalbard, Colesbukta’ [= Svalbard, Spitsbergen, Nordenskiöld Land, bay Colesbukta on the southern side of Isfjorden] 78.1120°N, 15.0290°E, 11.VII.2012, T. Ekrem, E. Stur & G.E.E. Søli, 1 male (NHMO 261328 and BOLD Sample ID SV910, in NHMO); ‘ Svalbard, Hanaskogdalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Hanaskogdalen on the eastern side of Adventfjorden], 78.2830°N, 15.6050°E, 25 m, 12.VII.2012, T. Ekrem, E. Stur & G.E.E. Søli, 4 males (NHMO 261385 and BOLD Sample ID SV967, NHMO 261390 and BOLD Sample ID SV972, 2 in NHMO; BOLD Sample ID SV945, 1 in NTNU-VM; BOLD Sample ID SV974, 1 in SDEI); ‘Svalbard, Krossfjorden, 14. juli bukta’ [= Svalbard, Spitsbergen, bay Fjortende Julibukta (northern side) on the eastern side of Krossfjorden], 79.1284°N, 11.8582°E, 3 m, 18.VII.2013, T. Ekrem, K. Harsaker & G.E.E. Søli, 1 male (NHMO 261046 and BOLD Sample ID SV1137, in NHMO); ‘ Svalbard, Lakselva’ [= Svalbard, Bjørnøya, river Lakselva], 74.4994°N, 18.9776°E, 15 m, 28.VII.2009, T. Ekrem, 1 male (NTNU-VM 50612 and BOLD Sample ID BJ262, in NTNU-VM); same locality but 3.VIII.2009, T. Ekrem, 2 males (NTNU-VM 50606 and BOLD Sample ID BJ256, NTNU-VM 50608 and BOLD Sample ID BJ258, in NTNU-VM); ‘ Svalbard, Lillehoeoekfjorden, Nilspynten’ [Sval- bard, Spitsbergen, Albert I Land, Lillehøkfjorden, E part of Mitrahalvøya, Nilspynten], 79.2660°N, 11.5690°E, 10 m, 18.VII.2013, G.E.E. Søli, 6 males (NHMO 261244 and BOLD Sample ID SV825, NHMO 261245 and BOLD Sample ID SV826, NHMO 261253 and BOLD Sample ID SV834, NHMO 261254 and BOLD Sample ID SV835, 4 in NHMO; BOLD Sample ID SV828, in SDEI; BOLD Sample ID SV 831 in MZH); ‘ Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2170°N, 15.6180°E, 16.VII.2012, G.E.E. Søli, T. Ekrem & E. Stur, 2 males (NHMO 261508 and BOLD Sample ID SV1263, NHMO 261175 and BOLD Sample ID SV1266, in NHMO); ‘Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2088°N, 15.5889°E, 14.VII.2012, G.E.E. Søli, 2 males (NHMO 261508 and BOLD Sample ID SV-NHMO91, in NHMO; BOLD Sample ID SV-NHMO94, in SDEI); ‘ Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2130°N, 15.6040°E, 14.VII.2012, E. Stur, 2 males (NHMO 261191 and BOLD Sample ID SV1282, in NHMO; BOLD Sample ID SV1283, in MZH); Svalbard, NW part of Spitsbergen, southern coast of Kongsfjord, W of Ny Ålesund, yellow pan trap, 2–15.VII.1974, Stephan, 2 males (in NHMO, SDEI); SWEDEN, Lapland, Abisko, Naturvetenskapliga Station, meadow at the station, sweep-net, 28.VI.1988, M. von Tschirnhaus, 4 males (2 in NHMO, 2 in SDEI). BIN. BOLD:ABA5288. Discussion. The species was described from one female from Novaya Zemlya by Holmgren (1883) and later described again based on some males and females from Greenland as Sciara glacialis (Lundbeck 1898) and from Spitsbergen as Sciara permutata (Lundbeck 1900). The species was redescribed with a list of literature, synonyms and combinations by Menzel & Mohrig (2000). Trichocoelina vitticollis (Holmgren) can be distinguished from all other species of the genus by its very large, setose intergonocoxal lobe of hypopygium and in having the gonostylus with a group of slender, oblique medial megasetae. See also under Trichocoelina tecta sp. n.Published as part of Vilkamaa, Pekka & Menzel, Frank, 2019, Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species, pp. 1-67 in Zootaxa 4665 (1) on pages 47-50, DOI: 10.11646/zootaxa.4665.1.1, http://zenodo.org/record/400055
Trichocoelina vitticollis Vilkamaa & Menzel 2019, comb. n.
Trichocoelina vitticollis (Holmgren, 1883) comb. n. Synonyms: = glacialis (Lundbeck, 1898) [as Sciara; preocc., not Sciara glacialis Rübsaamen, 1898]; = permutata (Lundbeck, 1900) [as Sciara; new name for Sciara glacialis Lundbeck, 1898]. Literature. Sciara glacialis Lundbeck—Lundbeck (1898): 254, pl. 6, fig. 13. Sciara humicola Lundbeck—Lundbeck (1898): 252, pl. 6, fig. 11 [misidentification]. Sciara permutata Lundbeck—Lundbeck (1900): 313. Bradysia permutata (Lundbeck) — McAlpine (1964): 128. Bradysia (Hemineurina) permutata (Lundbeck) — Frey (1948): 66, 84; pl. 18, fig. 106. Lycoriella (Hemineurina) permutata (Lundbeck) — Tuomikoski (1959a): 36; Tuomikoski, (1960): 75, 76; Stone & Laffoon (1965): 232; Tuomikoski (1967): 48; Gerbachevskaja-Pavluchenko (1986): 31 [not sensu Mohrig et al. (1983a) 15; misidentification, = T. subpermutata (Mohrig & Mamaev, 1990)]. Sciara vitticollis Holmgren—Holmgren (1883): 182; Jacobson (1898): 190; Gerbachevskaja-Pavluchenko (1986): 71. Lycoria (Neosciara) vitticollis (Holmgren) — Lengersdorf 1928 –30: 59. Lycoriella (Hemineurina) vitticollis (Holmgren) — Menzel & Mohrig (2000): 411, figs 380, 381; Coulson & Refseth (2004): 103; Coulson (2008): 162; Coulson (2013): 154; Mohrig et al. (2013): 271; Vilkamaa (2015): 551; Wirta et al. (2016): appendix, unpaginated p. 21 (table S1) and unpaginated p. 39 (cladogram). New records. CANADA, Northwest Territories, Mackenzie Delta, Reindeer Depot, 29.VI.1948, J.R. Vockeroth, 1 male (in MZH); NE GREENLAND, Zackenberg, 74°28’N, 20°34’W, 1.VIII.1991, J. Böcher, 1 male (in ZMUC); NW GREENLAND, Thule, 8.VII.1940, C. Wibe, 1 male (in ZMUC); SE GREENLAND, Skoldängen, 14–27.VII.1992, 2 males (1 in MZH, 1 in ZMUC); SW GREENLAND, Nugssuak, 18.VII.1949, C. Wibe, 1 male (in ZMUC); Saputit, 6.VII.1949, C. Wibe, 1 male (in MZH); W GREENLAND, Arnangarnup kua, 11.VII.1984, J. Böcher, 1 male (im ZMUC); Skjoklungen, ‘ Bygder’, 19–27.VII.1992, S. Andersen, 1 male (in ZMUC); Söndre Stormfjord, 21.VI.1952, C. Wibe, 1 male (in MZH); same locality but 67°02’N, 50°40’W, 2.VIII.1992, J. Böcher, 1 male (in MZH); NORWAY, ‘ Svalbard, Bjorndalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Bjørndalen W of Adventfjorden], 78.2320°N, 15.3270°E, 13.VII.2012, G.E.E. Søli, 1 male (NHMO 261403 and BOLD Sample ID SV985, in NHMO); ‘ Svalbard, Bolterdalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Bolterdalen on the southern side of Adventdalen], 78.1640°N, 15.9900°E, 100 m, 10.VII.2012, G.E.E. Søli, 3 males (NHMO 260984 and BOLD Sample ID SV1061, NHMO 260985 and BOLD Sample ID SV1062, 2 in NHMO; BOLD Sample ID SV1064, in SDEI); ‘Svalbard, Colesbukta’ [= Svalbard, Spitsbergen, Nordenskiöld Land, bay Colesbukta on the southern side of Isfjorden] 78.1120°N, 15.0290°E, 11.VII.2012, T. Ekrem, E. Stur & G.E.E. Søli, 1 male (NHMO 261328 and BOLD Sample ID SV910, in NHMO); ‘ Svalbard, Hanaskogdalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Hanaskogdalen on the eastern side of Adventfjorden], 78.2830°N, 15.6050°E, 25 m, 12.VII.2012, T. Ekrem, E. Stur & G.E.E. Søli, 4 males (NHMO 261385 and BOLD Sample ID SV967, NHMO 261390 and BOLD Sample ID SV972, 2 in NHMO; BOLD Sample ID SV945, 1 in NTNU-VM; BOLD Sample ID SV974, 1 in SDEI); ‘Svalbard, Krossfjorden, 14. juli bukta’ [= Svalbard, Spitsbergen, bay Fjortende Julibukta (northern side) on the eastern side of Krossfjorden], 79.1284°N, 11.8582°E, 3 m, 18.VII.2013, T. Ekrem, K. Harsaker & G.E.E. Søli, 1 male (NHMO 261046 and BOLD Sample ID SV1137, in NHMO); ‘ Svalbard, Lakselva’ [= Svalbard, Bjørnøya, river Lakselva], 74.4994°N, 18.9776°E, 15 m, 28.VII.2009, T. Ekrem, 1 male (NTNU-VM 50612 and BOLD Sample ID BJ262, in NTNU-VM); same locality but 3.VIII.2009, T. Ekrem, 2 males (NTNU-VM 50606 and BOLD Sample ID BJ256, NTNU-VM 50608 and BOLD Sample ID BJ258, in NTNU-VM); ‘ Svalbard, Lillehoeoekfjorden, Nilspynten’ [Sval- bard, Spitsbergen, Albert I Land, Lillehøkfjorden, E part of Mitrahalvøya, Nilspynten], 79.2660°N, 11.5690°E, 10 m, 18.VII.2013, G.E.E. Søli, 6 males (NHMO 261244 and BOLD Sample ID SV825, NHMO 261245 and BOLD Sample ID SV826, NHMO 261253 and BOLD Sample ID SV834, NHMO 261254 and BOLD Sample ID SV835, 4 in NHMO; BOLD Sample ID SV828, in SDEI; BOLD Sample ID SV 831 in MZH); ‘ Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2170°N, 15.6180°E, 16.VII.2012, G.E.E. Søli, T. Ekrem & E. Stur, 2 males (NHMO 261508 and BOLD Sample ID SV1263, NHMO 261175 and BOLD Sample ID SV1266, in NHMO); ‘Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2088°N, 15.5889°E, 14.VII.2012, G.E.E. Søli, 2 males (NHMO 261508 and BOLD Sample ID SV-NHMO91, in NHMO; BOLD Sample ID SV-NHMO94, in SDEI); ‘ Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2130°N, 15.6040°E, 14.VII.2012, E. Stur, 2 males (NHMO 261191 and BOLD Sample ID SV1282, in NHMO; BOLD Sample ID SV1283, in MZH); Svalbard, NW part of Spitsbergen, southern coast of Kongsfjord, W of Ny Ålesund, yellow pan trap, 2–15.VII.1974, Stephan, 2 males (in NHMO, SDEI); SWEDEN, Lapland, Abisko, Naturvetenskapliga Station, meadow at the station, sweep-net, 28.VI.1988, M. von Tschirnhaus, 4 males (2 in NHMO, 2 in SDEI). BIN. BOLD:ABA5288. Discussion. The species was described from one female from Novaya Zemlya by Holmgren (1883) and later described again based on some males and females from Greenland as Sciara glacialis (Lundbeck 1898) and from Spitsbergen as Sciara permutata (Lundbeck 1900). The species was redescribed with a list of literature, synonyms and combinations by Menzel & Mohrig (2000). Trichocoelina vitticollis (Holmgren) can be distinguished from all other species of the genus by its very large, setose intergonocoxal lobe of hypopygium and in having the gonostylus with a group of slender, oblique medial megasetae. See also under Trichocoelina tecta sp. n.Published as part of Vilkamaa, Pekka & Menzel, Frank, 2019, Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species, pp. 1-67 in Zootaxa 4665 (1) on pages 47-50, DOI: 10.11646/zootaxa.4665.1.1, http://zenodo.org/record/400055
Trichocoelina ithyspina Vilkamaa & Menzel 2019, sp. n.
Trichocoelina ithyspina sp. n. Figs 8 A, 9 B, 18 C Material studied. Holotype male. NORWAY, Hedmark, Stor-Elvdal, river Atna Solbakken, 61.746°N, 10.746°E, 380 m, Malaise trap, 30.VI–7.VII.2008, E. Stur (NTNU-VM 201775, BOLD Sample ID TRD-Sci029, in NTNU- VM). Description. Male. Head. Face and antenna concolorous brown but pedicellus yellow, maxillary palpus pale yellowish. Eye bridge, face and clypeus poorly visible in the specimen studied. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 seta, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.9x as long as wide, the neck shorter than broad, the longest setae longer than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 9 setae. Proepisternum with 11 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.3 mm. Width/length 0.45. Anal lobe weak. Veins distinct. c/w 0.65. R 1 /R 0.80. stM shorter than fork of M. r-m longer than bM, bM non-setose, r-m with 1 seta. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large indistinct patch in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, moderately long and fine. Hypopygium (Fig. 8 A). Brown, as abdomen. Intergonocoxal area moderately long, with long, apically bifid setose lobe (Fig. 18 C). Gonocoxa moderately broad, longer than gonostylus, medial margin basally smoothly curved, with rather long but sparse setosity. Gonostylus (Fig. 9 B) long, apically truncate, strongly impressed; with normal setosity, with a rather long apical tooth, with three megasetae subapically near the tooth and two more basally, megasetae long and slender, with distinct basal bodies; with 1 well-differentiated whiplash seta on ventromedial margin. Tegmen broader than long, laterally roundish, apically straight, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. BOLD:ACX5966. Discussion. The gonostylus shown in Figs 8 A and 9 B was slightly distorted in preparation. Trichocoelina ithyspina sp. n. resembles most T. jukkai sp. n. For discussion, see under T. dispansa sp. n. Etymology. The name is derived from the Greek word ithys, straight, and the Latin word spina, spine, referring to the straight megasetae of the gonostylus.Published as part of Vilkamaa, Pekka & Menzel, Frank, 2019, Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species, pp. 1-67 in Zootaxa 4665 (1) on page 29, DOI: 10.11646/zootaxa.4665.1.1, http://zenodo.org/record/400055
Lycoriella excavata Menzel & Vilkamaa 2021, sp. n.
Lycoriella excavata sp. n. Figs 2B, 3D, 12B Material studied. Holotype male. USA, Colorado, Laramer Co., Pinewood Springs, 40.2677°N, 105.3678°W, Malaise trap, 26.VII–7.IX.2014, S. & B. Fitzgerald (in MZH, http://id.luomus.fi/GE.1811). Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 1–2 facets wide. Face with 27 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 3 setae, with deep dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.8x as long as wide, neck slightly longer than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 9 setae. Proepisternum with 10 setae. Scutellum with 2 moderately long and some short and fine setae. Wing. Fumose. Length 1.9 mm. Width/length 0.40. Anal lobe small. Veins distinct, except for stM. c/w 0.65. R1/R 0.75. stM longer than fork of M. bM as long as r-m, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Pale brown; setae pale. Fore tibial organ forming large patch of setae in demarcated depression. Fore tibial spur longer than apical width of tibia. Abdomen. Pale brown; setae pale, short and weak. Hypopygium (Fig. 2B). Brown, like abdomen. Intergonocoxal area moderately long, with semicircular setose lobe. Gonocoxa narrow and long, longer than gonostylus, with sparse, rather short setosity. Gonostylus (Fig. 3D) widest subapically, slightly narrowed towards apex, deeply excavated dorsally; densely setose apically, with long and narrow apical tooth, with 1 subapical and 7–8 medial megasetae on ventral margin of apical half of gonostylus; with long whip-lash seta in middle. Tegmen (Fig. 12B) longer than wide, slightly notched apically, weakly sclerotized laterally, with area of minute teeth apicoventrally, with area of small aedeagal teeth. Aedeagal apodeme moderately long. BIN. Unknown. Discussion. Lycoriella excavata sp. n. with its subapically widened, lobe-shaped gonostyli resembles Lycoriella piristylata Vilkamaa, Hippa & Heller, 2013, which has (in contrast to the original description) also a palpal sensory pit and therefore belongs to Lycoriella Frey. In contrast to Lycoriella excavata sp. n., in L. piristylata a long whiplash seta is missing, the intercoxal lobe is small (or absent in some specimens), the apical tooth is significantly shorter or completely reduced and the tegmen is short and trapezoidal (Vilkamaa et al. 2013: fig. 3A–C). Lycoriella excavata sp. n. (Fig. 2B) also differs from this species in its ventrally enlarged lobe-shaped gonostylus with a very deep dorsal excavation, all medial megasetae of the gonostylus on its ventral margin, and its tegmen with apicolateral teeth. Apicolateral teeth of the tegmen are currently known only in Lycoriella taimyrensis described here (Fig. 13E), in some species of angustostylata species group in Bradysia Winnertz, 1867 (Menzel & Heller 2005; Mohrig 2016), as well in Odontosciara Rübsaamen, 1908 (Mohrig 2003). Etymology. The name is a Latin adjective, excavata, meaning hollowed, and refers to the strongly hollowed gonostylus.Published as part of Menzel, Frank & Vilkamaa, Pekka, 2021, New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region, pp. 501-530 in Zootaxa 5072 (6) on pages 509-510, DOI: 10.11646/zootaxa.5072.6.1, http://zenodo.org/record/575154
Abstract
This chapter on longan and litchi (Litchi chinensis) covers the following: effects of soil type, soil pH, salinity and mycorrhizas on fertilizer requirement; factors affecting leaf composition; leaf and soil analyses, and orchard surveys for fertilizer requirement determination; N, P, K, Ca, Mg, Zn, Fe, Cu, B and Mn nutrition (concentration and uptake, effects on productivity, and deficiency symptoms); fertilizer programme for young orchards in Australia; and fertigation
Abstract
This chapter on photosynthesis and productivity in litchi (Litchi chinensis) and longan covers the following: relationship between photosynthesis and plant development; effects of the environment on photosynthesis; assimilate production and distribution; relationship between fruit growth and leaf expansion; relationship between yield and assimilate supply; relationship between fruit size and yield; relationship between yield and tree size; effects of light and temperature on growth and productivity; effects of planting density and pruning on productivity; and practical aspects of pruning. Tabulated data on the productivity of litchi and longan in China and Thailand, and on the effects of leaf position and stage of development on leaf fresh weight and area, chlorophyll content, net carbon dioxide assimilation, and stomatal conductance in litchi (cv. Bengal) in Queensland, Australia, are presented
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