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    Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae)

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    Grehan, John R., Mielke, Carlos G.C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf, Xue, Dayong (2021): Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae). Zootaxa 4920 (3): 339-358, DOI: https://doi.org/10.11646/zootaxa.4920.3.

    FIGURE 30 in Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae)

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    FIGURE 30. Magnificus FW major banding patterns (coloured outlines). Wing veins—blue lines. FW illustrations 30g, h of limited quality. Higher resolution unavailable.Published as part of Grehan, John R., Mielke, Carlos G.C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong, 2021, Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae), pp. 339-358 in Zootaxa 4920 (3) on page 352, DOI: 10.11646/zootaxa.4920.3.2, http://zenodo.org/record/447805

    FIGURES 9–11 in New species of Endoclita (Lepidoptera: Hepialidae) and revived species status of E. kosemponis from Taiwan

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    FIGURES 9–11. Genitalia: 9a, E. meifenga sp. n (HT); 9b, E. kosemponis (LT): dpp—dorsal posterior projection, pvfposterio-ventral flange, mpp—medial posterior projection, mt—medial tooth, avn—anterio-ventral nexus, va—valva; 9c, E. sinensis (vincular margin with medial spine detached with abdomen—ref. Fig. 8c); 9d, E. davidi (HT). 10, external female genitlaia: 10a, E. kosemponis; 10b, E. sinensis. 11, internal female genitalia: 11a, E. kosemponis; 11b, E. sinensis.Published as part of Buchsbaum, Ulf & Grehan, John R., 2019, New species of Endoclita (Lepidoptera: Hepialidae) and revived species status of E. kosemponis from Taiwan, pp. 432-444 in Zootaxa 4551 (4) on page 439, DOI: 10.11646/zootaxa.4551.4.3, http://zenodo.org/record/262304

    A new Endoclita (C. &. R. Felder, 1874) from the High Mountains of Taiwan with notes about its evolutionary origins (Lepidoptera: Hepialidae)

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    Buchsbaum, Ulf, Hsu, Li-Peng, Chen, Ding-Jia, Grehan, John R. (2018): A new Endoclita (C. &. R. Felder, 1874) from the High Mountains of Taiwan with notes about its evolutionary origins (Lepidoptera: Hepialidae). Zootaxa 4521 (3): 441-450, DOI: 10.11646/zootaxa.4521.3.

    FIGURES 12–17. Distribution and diversity. 12 in A new Endoclita (C. &. R. Felder, 1874) from the High Mountains of Taiwan with notes about its evolutionary origins (Lepidoptera: Hepialidae)

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    FIGURES 12–17. Distribution and diversity. 12, type locality of Endoclita atayala sp. n., Miaoli County, Taiwan. 13, Biotope of Endoclita atayala sp. n., Miaoli County, Taiwan. 14, Species numbers of Hepialidae recorded from different countries and regions (data from Nielsen et al. (2000). 15, Regional comparison of Endoclita species numbers by major land units. 16, Grid of species distribution densities for Endoclita. Grid units 15 x 15°. 17, distribution records for E. atayala sp. n. (red circle, Taiwan), E. crenilimbata (blue circles, Ping-fa, Giuzhou and Mupin, Sichuan) and E. nodus (white circles, Yuexi, Anhui, and Lushan, Jiangxi).Published as part of Buchsbaum, Ulf, Hsu, Li-Peng, Chen, Ding-Jia & Grehan, John R., 2018, A new Endoclita (C. &. R. Felder, 1874) from the High Mountains of Taiwan with notes about its evolutionary origins (Lepidoptera: Hepialidae), pp. 441-450 in Zootaxa 4521 (3) on page 445, DOI: 10.11646/zootaxa.4521.3.9, http://zenodo.org/record/261001

    FIGURE 23 in Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae)

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    FIGURE 23. Magnificus abdominal sclerites: (a) M. regius tergite II, (MWM), photo by Nikolai Ignatev, (b) M. bouvieri holotype SII (photo by Joël Minet), (c) Magnificus sp. sternites VII-VIII, photo by Nikolai Ignatev), (d) M. bouvieri holotype sternites VII-VIII, photo by Joël Minet), (e) M. regius sternites VII-VIII, photo by Nikolai Ignatev, (f) M. regius and (g) M. roseus, photos by Ulf Buchsbaum, (h) M. bouvieri holotype abdomen, photo by Joël Minet, (i) Magnificus sp. tergosternal sclerite, (j) Magnificus regius tergosternal sclerite, photos by Nikolai Ignatev, (k) Hepialus humuli tsclerite, (l) Zenophassus schamyl tergosternal sclerite, photos by John Grehan.Published as part of Grehan, John R., Mielke, Carlos G.C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong, 2021, Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae), pp. 339-358 in Zootaxa 4920 (3) on page 350, DOI: 10.11646/zootaxa.4920.3.2, http://zenodo.org/record/447805

    FIGURES 1–9 in Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae)

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    FIGURES 1–9. Adult moths: 1, Magnificus bouvieri male holotype, dorsal (MNHN); 2, M. dirschi male holotype, dorsal (MfN). Photo by Théo Léger; 3, M. miniatus female holotype, (a) dorsal, (b) ventral (IZAS). Photo by Dayong Xu; 4, M. regius: (a) female syntype (from Staudinger 1896, pl. 5, Fig. 11), (b) M. regius rubellus, male lectotype, dorsal (MfN). Photo by Théo Léger, (c) M. regius rubellus (ZSBS), Photo by Ulf Buchsbaum, (d) M. regius rubellus (MWM). Photo Nikolai Ignatev; 5, M. roseus: (a) holotype male, dorsal (MNHN). Photo by Joël Minet, (b) male, dorsal (ZSBS), Photo by Ulf Buchsbaum; 6, M. zhiduoensis female holotype, dorsal [cropped image available only] (GAAH). Photo by Dayong Xue; 7, Magnificus sp. female, dorsal (MWM). Photo by Nikolai Ignatev; 8, Hepialus humuli, female, dorsal (Moscow, Russia SKC). Photo by Svyatoslav Knyazev; 9. Zenophassus schamyl, male, dorsal (Apsheronsk environs, Caucasus, Russia SKC). Photo by Svyatoslav Knyazev.Published as part of Grehan, John R., Mielke, Carlos G.C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong, 2021, Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae), pp. 339-358 in Zootaxa 4920 (3) on page 342, DOI: 10.11646/zootaxa.4920.3.2, http://zenodo.org/record/447805

    FIGURES 10–20. Genitalia. 10 in Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae)

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    FIGURES 10–20. Genitalia. 10, Magnificus bouvieri male holotype genitalia, ventral view. Photo by Joël Minet; 11, Magnificus dirschi male holotype genitalia, ventral view. Photo by Théo Léger; 12, Magnificus regius genitalia: a, male syntype, ventral view, dissection ESN 2845(MfN). Photo by Théo Léger; b, male, ventral view (ZSBS). Photo by Ulf Buchsbaum; c, male, ventral (left) and lateral (right) (MWM). Photo by Nikolai Ignatev; 13, Magnificus roseus male genitalia, ventral view, (a) holotype (MNHN). Photo by Joël Minet, (b) (ZSBS). Photo by Ulf Buchsbaum; Fig. 16. Magnificus jiuzhiensis genitalia: (a) holotype male, ventral view (from Yan 2000); 15, 'Hepialiscus sylvina Linnaeus, 1761', male, ventral view (from Chu & Wang 1985b: fig. 31 and Zhu et al. 2004: fig. 120); 16, Magnificus jiuzhiensis, female external posterior view (from Yan 2000); 17, Magnificus miniatus female holotype external genitalia, posterior view. Photo by Dayong Xu; 18, Magnificus regius female external posterior and lateral interior view (MWM). Photo by Nikolai Ignatev; 19, Magnificus zhiduoensis female holotype external posterior view (from Yan 2000); 20, Magnificus sp. female external posterior view (MWM). Photo by Nikolai Ignatev.Published as part of Grehan, John R., Mielke, Carlos G.C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong, 2021, Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae), pp. 339-358 in Zootaxa 4920 (3) on page 347, DOI: 10.11646/zootaxa.4920.3.2, http://zenodo.org/record/447805

    Magnificus bouvieri Grehan & Mielke & Minet & Ignatev & Buchsbaum & Xue 2021, comb. n.

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    Magnificus bouvieri (Oberthür, 1913) comb. n. (Figs 1, 10, 23b, 23d) Hepialus (Phassus) bouvieri (Oberthür 1913: 671) Sthenopis bouvieri (Nielsen et al. 2000: 850) Examined material. Holotype ♂ MNHN: Verbatim label: Chasseurs indigènes des Missionnaires de [local collectors of the missionaries of] Ta-tsien-Loû, 1905 / ESN slide no. 2872/ Type. Original description (Oberthür 1913). “ Phassus bouvieri is close to regius, Str., and roseus, Obthr.; however, Phassus bouvieri does not present any vestige of the pink hue which characterizes regius and roseus. It is a very pretty species whose designs on the upper wings are very complicated and delicate.” Female. Unknown. Etymology. Named in honour of Eugène-Louis Bouvier, Professor of Entomology, MNHN (Oberthür 1913). Flight period. Unknown Remarks. Forewing pattern with oblique white band (orange outline in Fig. 30b) broken by dark brown patch over outer anterior and central discal cells. Very similar to M. zhiduoensis. Male genitalia (Fig. 10) distinguished from all other species (where male is known) by larger and rounded lateral corners and broadly rounded anterior margin of the saccus. Sympatric with S. roseus (Fig. 31).Published as part of Grehan, John R., Mielke, Carlos G. C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong, 2021, Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae), pp. 339-358 in Zootaxa 4920 (3) on page 343, DOI: 10.11646/zootaxa.4920.3.2, http://zenodo.org/record/447805

    FIGURES 24–29. 24–27 in Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae)

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    FIGURES 24–29. 24–27, Male genitalia of some northern Eurasian Hepialidae: (a—ventral view, b—pseudotegumen posteriodorsal view, c, lateral view); 24, Hepialus humuli M153; 25, Zenophassus schamyl M134, JRGC; 26, Triodia sylvina M158, JRGC; 27, Pharmacis spp. (a, b) P. bertrandi M156 a—ventral view, b—posterio-dorsal view, (c) P. cantabricus, ventral view (photos 24–26 by John Grehan, 27a-b by Carlos Mielke, 27c by Axel Kallies); 28, Sthenopis pretiosus M179: (a) ventral, (b) lateral (apex of pseudoteguminal arm behind valva), JRGC; 29, Sternum II in some northern Eurasian and North American Hepialidae, (a) Hepialus humuli M153, (b) Zenophassus schamyl M132, (c) Triodia sylvina M158, (d) Pharmacis bertrandi M156, (e) Sthenopis pretiosus M179, (f) Sthenopis purpurascens M119. Photos by John Grehan.Published as part of Grehan, John R., Mielke, Carlos G.C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong, 2021, Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae), pp. 339-358 in Zootaxa 4920 (3) on page 348, DOI: 10.11646/zootaxa.4920.3.2, http://zenodo.org/record/447805
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