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Trichogenia maxillaris Braasch & Soldan 1988
No full textTrichogenia maxillaris Braasch & Soldán Trichogenia maxillaris Braasch & Soldán, 1988: 120 (orig.) Heptagenia maxillaris; (comb.) Kluge, 2004: 173 The larva of this species was adequately described by Braasch and Soldán (1988). The adults are unknown. Diagnosis: Trichogenia maxillaris can be differentiated from congeners by the following combination of characteristics: i) supracoxal sclerites blunt; ii) absence of comb setae on apical margin of maxillae; iii) hypopharynx apically divergent (Fig. 11); iv) ventral surface of terminal segment of labial palp with long row of setae extending to posteromedial margin (Fig. 12). Material Examined: THAILAND: 1 larva, Chiengmai Prov, small stream and waterfalls, Doi Sutep, W of Chiengmai, 19 -XI- 1964, WL and JG Peters [PERC]; VIETNAM: 1 larva, Vihm Phú: Tam Dao Hill Stn, lower waterfall of stream flowing through town, 11 -V- 1996, B Hubley and DC Darling, ROM 961029 [ROM]; 1 larva, Gai Lai: An Keh Dist, 10 km NW Buoenloy, small stream 1km on trail past Cha R, 1 °rainforest edge, 27 -VI- 1996, B Hubley, ROM 961099 [ROM].Published as part of Webb, J. M., Braasch, D. & Mccaferty, W. P., 2006, Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae), pp. 55-68 in Zootaxa 1335 on pages 62-63, DOI: 10.5281/zenodo.17428
Trichogenia nasuta Webb & Braasch & Mccaferty 2006, N. COMB.
No full textTrichogenia nasuta (Ulmer) N. COMB. Heptagenia nasuta Ulmer, 1939: 567 (orig.) Ulmer (1939) described Heptagenia nasuta from adult males and females. The larva is unknown.Published as part of Webb, J. M., Braasch, D. & Mccaferty, W. P., 2006, Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae), pp. 55-68 in Zootaxa 1335 on page 65, DOI: 10.5281/zenodo.17428
Trichogenia Braasch & Soldan, N.SP.
No full textTrichogenia Braasch & Soldán As indicated above, the larva Ulmer (1939: 675) described as C. spectabilis is actually an unnamed species of Trichogenia [= Trichogenia ulmeri N.SP.]. Wang and McCafferty (2004) considered Ulmer's C ompsoneuria to belong to a different genus than Trichogenia based on differences in mouthpart morphology. However, examination of a new species described below and T. ulmeri shows that these two species are fundamentally the same as T. maxillaris Braasch and Soldán, 1988 and differ in only a few characters. Therefore, we place them all in Trichogenia. Adult males have not been associated with any larvae of Trichogenia. However, it is highly probable that Heptagenia nasuta Ulmer, 1939, known only from adults from Indonesia, is Trichogenia (Braasch and Soldán 1988) based on the following: i) the only confirmed larvae of the subfamily Heptageniinae known from southeast Asia are Trichogenia; ii) H. nasuta is the only adult of the subfamily Heptageniinae known from southeast Asia; iii) wings dissected from wingpads of mature Trichogenia larvae have a slight violet tinge, similar to H. nasuta (Braasch and Soldán 1988). For these reasons we provisionally transfer H. nasuta to Trichogenia [= Trichogenia nasuta (Ulmer) N. COMB.]. Diagnosis: Larvae of Trichogenia can be differentiated from those of all other genera of Heptageniidae by the following combination of characters: i) ventral side of maxillae with setae in a row; ii) tergum with branched robust setae and many long, fine setae; iii) lamellae of gills 1 reduced; iv) lamellae of at least gills 2-4 long, narrow, and pointed apically; v) apex of femora without dorsal projection. At this time we do not have material available to provide characters for separating adult males of Trichogenia from other Heptageniinae genera. The long pair of ventral spines on the penes shown in Figures 156 and 157 by Ulmer (1939: 569) in combination with the violet coloration in the wings may be diagnostic. Species included: Trichogenia maxillaris, T. nasuta N. COMB., T. hubleyi N.SP., T. ulmeri N.SP.Published as part of Webb, J. M., Braasch, D. & Mccaferty, W. P., 2006, Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae), pp. 55-68 in Zootaxa 1335 on pages 60-61, DOI: 10.5281/zenodo.17428
Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae)
No full textWebb, J. M., Braasch, D., Mccaferty, W. P. (2006): Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae). Zootaxa 1335: 55-68, DOI: 10.5281/zenodo.17428
Compsoneuria (Siamoneuria) kovaci Braasch 2005
No full textCompsoneuria (Siamoneuria) kovaci Braasch, 2005 Some revisional notes on this species are advisable relating to an unnecessary synonymization by Webb and McCafferty (2008). According to the well defined outlines of the genus Compsoneuria by the authors, larval diagnosis of Compsoneuria (Siamoneuria) is in contradiction to that of Compsoneuria, namely in lacking the combination of “long, sharply pointed supracoxal spurs (as in Fig. 79), black spotting on the head capsule and femora, and narrow, apically pointed glossae (Fig. 68)”. Compsoneuria (Siamoneuria) is characterized in male adults by having a hammer-like ecdyonuroid penis head with dorsally well developed lateral sclerites and by absence of ventral spines. Moreover, the larvae are without any features to join them with true Ecdyonurini, for example setate caudal filaments or fimbriate maxillae. Considering also some larval characters discussed by Braasch (2005), we conclude that Siamoneuria cannot belong in Compsoneuria but probably deserves its own status, a morphospecies possibly independent of other known genera. Further studies on eggs and other larval details are needed for a better assessment of its rightful placement within Heptageniidae (see in Braasch, 2006 c, p. 49–50, Figs. 1–21). For now, we see it as a species INCERTAE SEDIS. Larval characters of Compsoneuria kovaci incertae sedis are: a) unthickened margin of forehead, also in Electrogena Zurwerra & Tomka, 1985 and Thamnodontus Kluge, 2004 (Webb & McCafferty 2008); b) long slender antennae; close standing of labial glossae rounded; c) special gill set: gills III + IV notched and gills V + VI angled; gill VII narrow lanceolate; d) caudal filaments 1.3 x longer and 1.2 x broader than terminal filament; e) tibiae each with 1 row of long, fine setae; f) rounded supracoxal sclerites (also in Electrogena and Thamnodontus); g) larvae without median abdominal ridge which is in Notacanthurus Tshernova, 1974 (Kluge 2004) and in Darthus vadorus Webb &.McCafferty, 2007. Adults can be excluded from Thamnodontus and Notacanthurus by lacking anteriorly expanded head capsules and from Electrogena by absence of “teardrop shaped penes with partially fused apical and lateral sclerites” (Webb & McCafferty 2008).Published as part of Braasch, Dietrich & Boonsoong, Boonsatien, 2010, A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia, pp. 1-26 in Zootaxa 2610 on page 13, DOI: 10.5281/zenodo.19779
Trichogenia ulmeri Braasch & Webb N., N.SP.
No full textTrichogenia ulmeri Braasch & Webb N.SP. Compsoneuria spectabilis Eaton (larva nec adult); (misid.) Ulmer, 1939: 675 Larval Description: see Ulmer (1939). Adult: unknown. Etymology: The specific epithet is in honor of the late Georg Ulmer. Diagnosis: Trichogenia ulmeri can be differentiated from congeners by the following combination of characteristics: i) anterior surface of femora with both long and short robust setae (Fig. 13); ii) presence of comb setae on apical margin of maxillae; iii)long supracoxal spines on the meso- and metathorax; iii) abdominal terga with sparse branched, robust setae, most occurring near posterior margin. Material Examined: HOLOTYPE: 1 larva, Indonesia, Sumatra, Ranau, Urwaldbach, R 25 c, 29 -I- 1929, Thienemann [ZMUH]. PARATYPES: INDONESIA: 1 larva, same data as holotype [ZMUH]; 2 larvae, Indonesia, E. Kalimantan, Long Tua, Tua cr, 3 ° 10 'N 115 ° 47 'E, 440m, 7 -IV- 1994, B Hubley, IIS 940541 [ROM].Published as part of Webb, J. M., Braasch, D. & Mccaferty, W. P., 2006, Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae), pp. 55-68 in Zootaxa 1335 on page 66, DOI: 10.5281/zenodo.17428
FIGURE 2–5. Trichogenia hubleyi. 2 in Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae)
No full textFIGURE 2–5. Trichogenia hubleyi. 2, labrum: a. dorsal b. ventral; 3, left mandible; 4, right mandible; 5, hypopharynx: a. dorsal b. ventral.Published as part of Webb, J. M., Braasch, D. & Mccaferty, W. P., 2006, Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae), pp. 55-68 in Zootaxa 1335 on page 63, DOI: 10.5281/zenodo.17428
FIGURE 1 in Supplementary characters of Bursaphelenchus lini Braasch, 2004 (Nematoda: Parasitaphelenchidae) and remarks on this nematode
No full textFIGURE 1. Scanning electron micrographs of Bursaphelenchus lini: A. Head region, lateral view; B. Enface view of female head; C. Lateral field; D. Female tail; E. Vulval region; F.I: Male tail showing papillae (F, G), bursa (H) and protracted spicules (I).Published as part of <i>Braasch, Helen, Brandstetter, Martin & Burgermeister, Wolfgang, 2006, Supplementary characters of Bursaphelenchus lini Braasch, 2004 (Nematoda: Parasitaphelenchidae) and remarks on this nematode, pp. 55-61 in Zootaxa 1141 (1)</i> on page 59, DOI: 10.11646/zootaxa.1141.1.2, <a href="http://zenodo.org/record/10087138">http://zenodo.org/record/10087138</a>
Epeorus inthanonensis Braasch and Boonsoong, sp. nov.
No full textEpeorus inthanonensis Braasch and Boonsoong sp. nov. Description. HOLOTYPE: female La: Body length 9.3 mm, caudal filaments 13.0 mm; (Body length of male La 8.8 mm, caudal filaments 10.2 mm). PLATE XI. FIGURES 69–74. Larva of Epeorus inthanonensis sp. nov. 69, Head capsule. 70, Labrum, dorsal view (left) and ventral view (right). 71, Right mandible. 72, Left maxilla, ventral view. 73, Hypopharynx, dorsal view (left) and ventral view (right). 74, Labium, dorsal view (left) and ventral view (right). PLATE XII. FIGURES 75–82. Larva of Epeorus inthanonensis sp. nov. 75, Foreleg, right, dorsal view. 76, Foreleg femur, bristles on dorsal face. 77, Foreleg, tarsal claw. 78, Abdominal tergum VII, posterior margin. 79, Abdominal terga I–X. 80, Gill I. 81, Gill V. 82, Gill VII. Head (Fig. 69): ellipsoid, width 10.2 mm, without conspicuous markings, but dark brown around ocelli, eyes and antennae sockets, light brown in rest area; antennae very long: 8.3 mm; HI 34: 27 = 1.2. Labrum (Fig. 70): with median notch on anterior margin, with long hairlike setae dorsally and marginally. Mandibles (Fig. 71): lateral margin without setae, incisors weakly serrate; outer incisor longer than inner incisor. Maxillae (Fig. 72): with 3 acute teeth, medially with row of hairlike setae, apical segment of maxillary palp with dense, long, hairlike setae. Hypopharynx (Fig. 73): lingua with anterolateral lobes, superlinguae slightly expanded laterally with row of hairlike setae. Labium (Fig. 74): with narrow basal separation between glossae, glossae subtriangular, paraglossae slightly expanded laterally, apical segment of labial palp with heavy armature. Thorax: Pronotum light brown without clear marks; mesonotum somewhat darker brown with each a paramedian pair of darker spots anteriorly and a pair of inside curved streaks posteriorly. Femora yellowbrown without distinctively visible patches, dorsally with median spot (Fig. 75), dorsal surface of femora with scattered stout short bristles (Fig. 76), claws with 2 small denticles (Fig. 77). Abdomen: Prospective figures of adult stage is suggested by pigmentation of terga (Fig. 79); sterna light without any markings. Microstructure of terga characteristic as in segment VI (Fig. 78); along midline of terga II–VII a pair of small pointed tubercles; hind margins of terga with blunt bristles (Fig. 78). Gill plates I, III and VII as in Figs. 80–82. Etymology: Denomination refers to collection site on the highest mountain of Thailand: Doi Inthanon. Diagnosis: The new species most resembles Nepalese Epeorus rhithralis Braasch, 1980 and somewhat less Epeorus bifurcatus Braasch & Soldán, 1979. The latter is best distinguished by small submedian, acute pairs of dorsal spines and larger gills I than gill III (Nguyen & Bae 2004 a); in the new species the first gill is, with exception of the seventh gill, smaller than gills II–VI, while in E. rhithralis the first gill is much larger than all other gills. Further differences can be seen in shape of head, hypopharynx, gill set, femur spot and posterior margin of tergum VII (Braasch 1980: p. 63, Figs. 4 b; h; j; k; l; e and d). E. rhithralis as well as E. inthanonensis have paired tubercles on terga more rounded than in E. bifurcatus. E. inthanonensis has relatively the smallest first gill. In Vietnam specimens of E. bifurcatus, the hind margins of terga show blunt bristles densely rowed, whereas those of the two other species are pointed. E. inthanonensis has pairs of rounded, submedian tubercles on terga III–IX, densely provided with small conical bristles, forming a median groove between them, moderately scattered with tiny setae; obliquely running rim of the groove bears scarcely little setae directed mediodistally. The Hong Kong species Epeorus sagittatus Tong & Dudgeon, 2003 has, deviating from other Epeorus species of the region, the terga III–VIII each with a pair of rounded brown stripes submedially. Discussion. The new species is among those derived from Himalayan species, having spread southeastward by following foothills of the Himalayas to Thailand, including species such as Epeorus (Belovius) unicornutus, Epeorus bifurcatus, or still other species of Epeorini Wang and McCafferty, 2004. In contrast to these obviously unchanged species with large geographic ranges between Nepal and Thailand or even Vietnam, E. inthanonensis seems to be a close related species to Nepalese E. rhithralis with a large first gill, while it is strongly reduced in E. inthanonensis. Material examined: HOLOTYPE: female La, NW Thailand, Chiangmai Province, small mountain stream of Doi Inthanon, 18 º 31 ' 33.35 "N, 98 º 29 ' 57.20 "E, alt.> 2200 m; 19.II. 2004 (DB) in 70 % alcohol; PARATYPES: 3 La from the same place (DB); 3 La, small river from mountain region near Pai, 19 ° 28 ’07.28’’N, 98 ° 17 ’ 42.82 ’’E, alt. 829 m; 11.IV.03 (DB). All findings are bottom samples. Types deposition: HOLOTYPE: female La; PARATYPES: 1 La (MNHU), 2 La (SMF Eph); 2 La (ZMKU); 1 La (DBP).Published as part of Braasch, Dietrich & Boonsoong, Boonsatien, 2010, A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia, pp. 1-26 in Zootaxa 2610 on pages 17-19, DOI: 10.5281/zenodo.19779
Trichogenia hubleyi Webb & Braasch & Mccaferty 2006, N.SP.
No full textTrichogenia hubleyi Webb & McCafferty N.SP. Larval description: Head capsule subrectangular (Fig. 1), with numerous fine and sparse branched, robust setae dorsally. Labrum (Fig. 2) much wider than long, dorsally with numerous long, fine setae, ventrally with median row of short, robust setae and many long, fine setae laterally. Mandibles (Figs 3 and 4) with dense patch of setae on lateral margin at base of outer canines, lateral margin with row of long setae on apical half and patch of setae dorsally on basal portion. Hypopharynx (Fig. 5) with lateral margins of superlingua subparallel; lingua with median U-shaped notch bordered by pair of apicolateral processes. Maxillae (Fig 6) ventrally with row of fimbriate setae, dorsally with lateral row of long, fine setae; apical margin of galealaciniae with nine comb setae; first segment of palp with numerous long, fine setae; second segment with long, fine setae laterally, bare medially; third segment small, sharp. Glossae broadly rounded apically, numerous robust setae ventrally; dorsal surface of last segment of palps with row of fine setae stopping at medial margin (Fig 7). Thorax dorsally with numerous fine setae, sparse ginko-shaped robust setae; pronotum with pair of small, spicule-covered protuberances; meso- and metathoracic supracoxal sclerites long, sharp, projecting dorsally (Fig. 8). Forefemora dorsally with row of long, fine setae and numerous long, robust setae; anterior surface with spatulate setae and long, fine setae; ventral margin with numerous short, robust setae and short, fine setae. Foretibiae with sparse row of fine setae on posterior margin; lateral ridge with row of spatulate setae; anterior margin with numerous short, robust setae; entire surface of tibiae covered with fine setae. Foretarsi covered with fine setae, laterally with several spatulate setae. Mid- and hindlegs similar to forelegs but with more numerous robust setae; femora with anterior face with robust setae all of approximately same size and shape (Fig. 9). Abdomen dorsally with numerous fine setae and branched robust setae (Fig. 10); posterior margins of terga with sharp spines. Gills 1–7 with numerous long fibrils, lamellae with long, fine setae; lamellae of gills 1 small; lamellae of gills 2–4 long and narrow; lamellae of gills 5–7 rounded, with short apical point. Caudal filaments with short spines at intersegmental margins; median caudal filament with long, fine setae on both sides of joints; cerci with long, fine setae medially. Adult: Unknown. Etymology: The specific epithet is in honor of Brad Hubley of the Royal Ontario Museum, Canada, who has provided us with numerous southeast Asian specimens. Diagnosis: Trichogenia hubleyi can be differentiated from congeners by the following combination of characteristics: i) well-developed comb setae on the anterior margin of the galealaciniae; ii) long supracoxal spines on the meso- and metathorax; iii) anterior surface of femora with setae all of same size; iv) abdominal terga with numerous branched, robust setae scattered over entire surface. Material examined: HOLOTYPE: 1 larva, Indonesia, Celebes, Sulawesi Utara Province, swift clear stream nr S end of L Mara, 10 -IX- 1985, JT & DA Polhemus [PERC] PARATYPES: 17 larvae (parts of one slide-mounted), same data as holotype [PERC].Published as part of Webb, J. M., Braasch, D. & Mccaferty, W. P., 2006, Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae), pp. 55-68 in Zootaxa 1335 on pages 61-62, DOI: 10.5281/zenodo.17428
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