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Agroeca maghrebensis Bosmans, 1999, n. sp.
No full textAgroeca maghrebensis n. sp. (Figs 5-8) Agroeca annulipes: Simon 1932: 972 (descr. male, non female). Type material. Holotype male from Algeria, Wilaya of Bouira, Ighrem, 490m in pitfalls among Tamarisk along Oued Sahel, 17.XI.1986, R. Bosmans leg.; deposited in IRSNB. Diagnosis: Males of this species are easily distinguished by the dorsal angularity on the male palpal tibia, females by the epigyne with median depression with parallel margins and x-shaped pattern of copulation ducts. Remarks: The male Figured by Simon (1932) as the male of Agroeca annulipes, Sim on 1878 apparently belongs to the species described here. Description: Measurements: Male: Total length 4.0; carapace 2.06 long, 1.68 wide. Female: Total length 4.6-6.4; carapace 2.32-2.84 long, 1.72-2.40 wide. Colour and spination as in Agroeca annulipes. (Figs. 5-6): Tibial apophysis straight, with distinct dorsal angularity; median apophysis hooked, distal part strongly pointed; wide basal part of embolus visible in anteromedian part of bulb, gradually narrowing, curved in antero-dorsal direction. Epigyne (Fig. 7): Longer than wide; with large median depression, posteriorly truncate, anterior half with narrow median septum; depression often filled with concretions (“copulation plugs”); x-shaped pattern of touching copulation ducts visible in transparency. Vulva (Fig. 8): Copulation ducts semi-circular, touching in middle; spermathecae not wider than copulation ducts. Material examined and citations: Algeria - Aïn Temouchent: between El Malah and El Ghella, along Rio Salado, 80m, 1 female in Atriplex litter, 24.IV.1984, R. Bosmans leg. (CRB). - Batna: Massif de l’Aures, Monts de Belezma, Col Telmet, 1820m, 2 males in pitfalls in Cedrus forest, 26.II.1988, R. B osm ans leg. (CRB). - Blida: Atlas Blideen, Meftah, 480m, 4 females in pitfalls in Quercus suber forest, 11.XII.1986, R. Bosmans leg. (CRB). - Bordj Bou Arreridj: Sidi Embarek, 900m, 1 female, stones in cultivated land, 27.II.1990, R. Bosmans leg. (CRB). - Bouira: E. Bechloul, Oued Zaiane, 400m, 1 female in tamarisk litter, 28.IV.1988, R. Bosmans leg. (CRB); Col de Dirah, 900m, 2 females, pitfalls in rough grassland along Oued Djenane, 1.V.1988, R. Bosmans leg. (CRB). - Jijel: Ziama Mansouria, 15m, 7 females in flooded orchard, 24.XI.1989, R. B osm ans leg. (2 females in MNHNP, 2 females in IRSNB, 3 females in CRB). - Laghouat: Laghouat, Oud M’zi, 750m, 1 female in Phragmites litter, 21.V.1990, R. Bosmans leg. (CRB). M.3’sila: 5 km E. Mansourah, 800m, 1 female, stones in dry Pinus halepensis forest, 16.X.1987, R. Bosmans leg. (CRB). - Oran: N. Misserghin, 200m, 1 female in flooded orchard, 25.IV. 1984, R. Bosmans leg. (CRB). - Setif:: D jebel Babor, 1850m, 1 female in mixed forest, 20.VI 1986; N. Magra, Rasfa, 1 female in herbs along an oued. 15.lV. 1990, R. Bosmans leg. (CRB). Skikda: B ouchata, 400m, 1 fem ale, stones in grassland, 12.III.1990, R. Bosmans leg. (CRB). - Souk Ahras: 2 km S. Souk Ahras, 660m, 1 female, stones in P- halepensis forest, 10.II.1988, R. Bosmans leu. (CRB).- Tlemcen: S. Tlemcen, forêt de Tal Terny, 1300m, 1 female, pitfalls in Quercus ilex forest, 24.V.1990, R. Bosmans leg. (CRB). Morocco - Messia - Melilla, 1 female, sub Mesiotelus sp. (MNHNP 19.700). Tunisia - Nabeul: Djebel El Haouaria, 1 female in maquis, 13.IV.1993, K. De Smet leg. (CRB). Distribution: Algeria, Morocco, Tunisia. Not as coastal as A. annulipes. Ecology: Adult in winter. Males were collected from November to February, females from November to May. Agroeca inopina O.P.- Cambridge, 1886 (Figs 9-12) Agroeca inopina O.P.- Cambridge 1886: 71 (descr. male, female); Simon 1932: 972; Grimm 1986: 29. Diagnosis: Besides the copulatory organs, this species is easily distinguished from both other Agroeca species occurring in the Maghreb by its yellowish brown cephalothorax and the uniform legs with no annulations. Description: Measurements: Male: Total length 5.0-5.4; carapace 2.34-2.44 long, 1.82-1.92 wide. Female: Total length 5.3- 7.6; carapace 2.56-2.64 long, 2.02-2.16 wide. Colour: Carapace yellowish brown with narrow black margin and anastomosing, radiating dark brown striae; legs yellowish brown, not annulated; abdomen with central reddish brown band, laterally darker. Palp (Figs. 9-10): Tibial apophysis relatively short; median apophysis large, terminally with elongated hook. Epigyne (Fig. 11): As long as wide, with large anteromedian depression with triangular antero-median septum; depression often filled with secretions (“ copulation plugs”). Vulva (Fig. 12): Copulation ducts c-shaped, widely touching in middle; spermathecae not wider than copulation ducts. Material examined: - Blida: Djebel Mouzaia, 1200m, 1 male 1 female stones in grassland around lake, 27.I.1990, R. Bosmans leg. (CRB); idem, 1250m, 1 male in pitfalls in Quercus suber forest, 27.I.1990, R. Bosmans leg. (CRB). - Tizi Ouzou: Aït Ouabane, 1850m, 2 males, stones in montane grassland, 15.XI.1985, R. Bosmans leg. (CRB); forêt d’Akfadou, 1200m, 1 female in Quercus faginea litter, 18.V.1988 and 910m, 1 female in Quercus faginea litter, 24.III.1988, R. Bosmans leg. (CRB); Tala Guilef, 1600m, 1 male in pitfalls in dense Cedrus forest, 16.III.1990, R. Bosmans leg. (CRB); Yakouren, 800m, 1 female in pitfalls in Quercus faginea forest, 22.IV. 1982, 1 female, 4.XII.1986 and 1 female, 25.1.1990, R. Bosmans leg. (CRB). - Tiemcen: S.E. Tiemcen, forêt d’Hafir, 1350m, 1 female in Quercus ilex litter, 23.V.1990, R. Bosmans leg. (CRB). Distribution: Portugal, Spain, France, S. England, Austria, S.W. Finland, and cited here for the first time from Africa from Algeria, where it appears to be a species of Agroeca inopina O.P.- Cambridge, 1886 (Figs 9-12) Agroeca inopina O.P.- Cambridge 1886: 71 (descr. male, female); Simon 1932: 972; Grimm 1986: 29. Diagnosis: Besides the copulatory organs, this species is easily distinguished from both other Agroeca species occurring in the Maghreb by its yellowish brown cephalothorax and the uniform legs with no annulations. Description: Measurements: Male: Total length 5.0-5.4; carapace 2.34-2.44 long, 1.82-1.92 wide. Female: Total length 5.3- 7.6; carapace 2.56-2.64 long, 2.02-2.16 wide. Colour: Carapace yellowish brown with narrow black margin and anastomosing, radiating dark brown striae; legs yellowish brown, not annulated; abdomen with central reddish brown band, laterally darker. Palp (Figs. 9-10): Tibial apophysis relatively short; median apophysis large, terminally with elongated hook. Epigyne (Fig. 11): As long as wide, with large anteromedian depression with triangular antero-median septum; depression often filled with secretions (“ copulation plugs”). Vulva (Fig. 12): Copulation ducts c-shaped, widely touching in middle; spermathecae not wider than copulation ducts. Material examined: - Blida: Djebel Mouzaia, 1200m, 1 male 1 female stones in grassland around lake, 27.I.1990, R. Bosmans leg. (CRB); idem, 1250m, 1 male in pitfalls in Quercus suber forest, 27.I.1990, R. Bosmans leg. (CRB). - Tizi Ouzou: Aït Ouabane, 1850m, 2 males, stones in montane grassland, 15.XI.1985, R. Bosmans leg. (CRB); forêt d’Akfadou, 1200m, 1 female in Quercus faginea litter, 18.V.1988 and 910m, 1 female in Quercus faginea litter, 24.III.1988, R. Bosmans leg. (CRB); Tala Guilef, 1600m, 1 male in pitfalls in dense Cedrus forest, 16.III.1990, R. Bosmans leg. (CRB); Yakouren, 800m, 1 female in pitfalls in Quercus faginea forest, 22.IV. 1982, 1 female, 4.XII.1986 and 1 female, 25.1.1990, R. Bosmans leg. (CRB). - Tiemcen: S.E. Tiemcen, forêt d’Hafir, 1350m, 1 female in Quercus ilex litter, 23.V.1990, R. Bosmans leg. (CRB). Distribution: Portugal, Spain, France, S. England, Austria, S.W. Finland, and cited here for the first time from Africa from Algeria, where it appears to be a species ofPublished as part of Bosmans, Robert, 1999, The genera Agroeca, Agraecina, Apostenus and Scotina in the Maghreb countries (Araneae: Liocranidae), pp. 25-34 in Bulletin Di L' Institut Royal Des Sciences Naturelles De Belgique Entomologie 69 on pages 27-29, DOI: 10.5281/zenodo.82621
Agroeca maghrebensis Bosmans, 1999, n. sp.
No full text<p>Agroeca maghrebensis n. sp.</p> <p>(Figs 5-8)</p> <p>Agroeca annulipes: Simon 1932: 972 (descr. male, non female).</p> <p>Type material. Holotype male from Algeria, Wilaya of Bouira, Ighrem, 490m in pitfalls among Tamarisk along Oued Sahel, 17.XI.1986, R. Bosmans leg.; deposited in IRSNB.</p> <p>Diagnosis: Males of this species are easily distinguished by the dorsal angularity on the male palpal tibia, females by the epigyne with median depression with parallel margins and x-shaped pattern of copulation ducts.</p> <p>Remarks: The male Figured by Simon (1932) as the male of Agroeca annulipes, Sim on 1878 apparently belongs to the species described here.</p> <p>Description:</p> <p>Measurements: Male: Total length 4.0; carapace 2.06 long, 1.68 wide. Female: Total length 4.6-6.4; carapace 2.32-2.84 long, 1.72-2.40 wide.</p> <p>Colour and spination as in Agroeca annulipes. (Figs. 5-6): Tibial apophysis straight, with distinct dorsal angularity; median apophysis hooked, distal part strongly pointed; wide basal part of embolus visible in anteromedian part of bulb, gradually narrowing, curved in antero-dorsal direction.</p> <p>Epigyne (Fig. 7): Longer than wide; with large median depression, posteriorly truncate, anterior half with narrow median septum; depression often filled with concretions (“copulation plugs”); x-shaped pattern of touching copulation ducts visible in transparency.</p> <p>Vulva (Fig. 8): Copulation ducts semi-circular, touching in middle; spermathecae not wider than copulation ducts.</p> <p>Material examined and citations:</p> <p>Algeria - Aïn Temouchent: between El Malah and El Ghella, along Rio Salado, 80m, 1 female in Atriplex litter, 24.IV.1984, R. Bosmans leg. (CRB).</p> <p>- Batna: Massif de l’Aures, Monts de Belezma, Col Telmet, 1820m, 2 males in pitfalls in Cedrus forest, 26.II.1988, R. B osm ans leg. (CRB).</p> <p>- Blida: Atlas Blideen, Meftah, 480m, 4 females in pitfalls in Quercus suber forest, 11.XII.1986, R. Bosmans leg. (CRB).</p> <p>- Bordj Bou Arreridj: Sidi Embarek, 900m, 1 female, stones in cultivated land, 27.II.1990, R. Bosmans leg. (CRB).</p> <p>- Bouira: E. Bechloul, Oued Zaiane, 400m, 1 female in tamarisk litter, 28.IV.1988, R. Bosmans leg. (CRB); Col de Dirah, 900m, 2 females, pitfalls in rough grassland along Oued Djenane, 1.V.1988, R. Bosmans leg. (CRB).</p> <p>- Jijel: Ziama Mansouria, 15m, 7 females in flooded orchard, 24.XI.1989, R. B osm ans leg. (2 females in MNHNP, 2 females in IRSNB, 3 females in CRB).</p> <p>- Laghouat: Laghouat, Oud M’zi, 750m, 1 female in Phragmites litter, 21.V.1990, R. Bosmans leg. (CRB).</p> <p>M.3’sila: 5 km E. Mansourah, 800m, 1 female, stones in dry Pinus halepensis forest, 16.X.1987, R. Bosmans leg. (CRB).</p> <p>- Oran: N. Misserghin, 200m, 1 female in flooded orchard, 25.IV. 1984, R. Bosmans leg. (CRB).</p> <p>- Setif:: D jebel Babor, 1850m, 1 female in mixed forest, 20.VI 1986; N. Magra, Rasfa, 1 female in herbs along an oued. 15.lV. 1990, R. Bosmans leg. (CRB).</p> <p> <b>Skikda: B ouchata, 400m, 1 fem ale, stones in grassland, 12.III.1990, R. Bosmans leg. (CRB).</b></p> <p>- Souk Ahras: 2 km S. Souk Ahras, 660m, 1 female, stones in P- halepensis forest, 10.II.1988, R. Bosmans leu. (CRB).-</p> <p>Tlemcen: S. Tlemcen, forêt de Tal Terny, 1300m, 1 female, pitfalls in Quercus ilex forest, 24.V.1990, R. Bosmans leg. (CRB).</p> <p> <b>Morocco - Messia - Melilla</b>, 1 <b>female, sub Mesiotelus sp. (MNHNP 19.700).</b></p> <p> <b>Tunisia -</b> <b>Nabeul: Djebel El Haouaria</b>, 1 <b>female in maquis,</b> <b>13.IV.1993, K. De Smet leg. (CRB).</b></p> <p> <b>Distribution: Algeria, Morocco, Tunisia. Not as coastal as</b> <b>A. annulipes.</b></p> <p>Ecology: Adult in winter. Males were collected from November to February, females from November to May.</p> <p>Agroeca inopina O.P.- Cambridge, 1886</p> <p>(Figs 9-12)</p> <p>Agroeca inopina O.P.- Cambridge 1886: 71 (descr. male, female); Simon 1932: 972; Grimm 1986: 29.</p> <p>Diagnosis: Besides the copulatory organs, this species is easily distinguished from both other Agroeca species occurring in the Maghreb by its yellowish brown cephalothorax and the uniform legs with no annulations.</p> <p>Description:</p> <p>Measurements: Male: Total length 5.0-5.4; carapace 2.34-2.44 long, 1.82-1.92 wide. Female: Total length 5.3- 7.6; carapace 2.56-2.64 long, 2.02-2.16 wide.</p> <p>Colour: Carapace yellowish brown with narrow black margin and anastomosing, radiating dark brown striae; legs yellowish brown, not annulated; abdomen with central reddish brown band, laterally darker.</p> <p>Palp (Figs. 9-10): Tibial apophysis relatively short; median apophysis large, terminally with elongated hook.</p> <p>Epigyne (Fig. 11): As long as wide, with large anteromedian depression with triangular antero-median septum; depression often filled with secretions (“ copulation plugs”).</p> <p>Vulva (Fig. 12): Copulation ducts c-shaped, widely touching in middle; spermathecae not wider than copulation ducts.</p> Material examined: - Blida: Djebel Mouzaia, 1200m, 1 male 1 female stones in grassland around lake, 27.I.1990, R. Bosmans leg. (CRB); idem, 1250m, 1 male in pitfalls in Quercus suber forest, 27.I.1990, R. Bosmans leg. (CRB). <p>- Tizi Ouzou: Aït Ouabane, 1850m, 2 males, stones in montane grassland, 15.XI.1985, R. Bosmans leg. (CRB); forêt d’Akfadou, 1200m, 1 female in Quercus faginea litter, 18.V.1988 and 910m, 1 female in Quercus faginea litter, 24.III.1988, R. Bosmans leg. (CRB); Tala Guilef, 1600m, 1 male in pitfalls in dense Cedrus forest, 16.III.1990, R. Bosmans leg. (CRB); Yakouren, 800m, 1 female in pitfalls in Quercus faginea forest, 22.IV. 1982, 1 female, 4.XII.1986 and 1 female, 25.1.1990, R. Bosmans leg. (CRB).</p> <p>- Tiemcen: S.E. Tiemcen, forêt d’Hafir, 1350m, 1 female in Quercus ilex litter, 23.V.1990, R. Bosmans leg. (CRB).</p> Distribution: Portugal, Spain, France, S. England, Austria, S.W. Finland, and cited here for the first time from Africa from Algeria, where it appears to be a species of <p>Agroeca inopina O.P.- Cambridge, 1886</p> <p>(Figs 9-12)</p> <p>Agroeca inopina O.P.- Cambridge 1886: 71 (descr. male, female); Simon 1932: 972; Grimm 1986: 29.</p> <p>Diagnosis: Besides the copulatory organs, this species is easily distinguished from both other Agroeca species occurring in the Maghreb by its yellowish brown cephalothorax and the uniform legs with no annulations.</p> <p>Description:</p> <p>Measurements: Male: Total length 5.0-5.4; carapace 2.34-2.44 long, 1.82-1.92 wide. Female: Total length 5.3- 7.6; carapace 2.56-2.64 long, 2.02-2.16 wide.</p> <p>Colour: Carapace yellowish brown with narrow black margin and anastomosing, radiating dark brown striae; legs yellowish brown, not annulated; abdomen with central reddish brown band, laterally darker.</p> <p>Palp (Figs. 9-10): Tibial apophysis relatively short; median apophysis large, terminally with elongated hook.</p> <p>Epigyne (Fig. 11): As long as wide, with large anteromedian depression with triangular antero-median septum; depression often filled with secretions (“ copulation plugs”).</p> <p>Vulva (Fig. 12): Copulation ducts c-shaped, widely touching in middle; spermathecae not wider than copulation ducts.</p> Material examined: - Blida: Djebel Mouzaia, 1200m, 1 male 1 female stones in grassland around lake, 27.I.1990, R. Bosmans leg. (CRB); idem, 1250m, 1 male in pitfalls in Quercus suber forest, 27.I.1990, R. Bosmans leg. (CRB). <p>- Tizi Ouzou: Aït Ouabane, 1850m, 2 males, stones in montane grassland, 15.XI.1985, R. Bosmans leg. (CRB); forêt d’Akfadou, 1200m, 1 female in Quercus faginea litter, 18.V.1988 and 910m, 1 female in Quercus faginea litter, 24.III.1988, R. Bosmans leg. (CRB); Tala Guilef, 1600m, 1 male in pitfalls in dense Cedrus forest, 16.III.1990, R. Bosmans leg. (CRB); Yakouren, 800m, 1 female in pitfalls in Quercus faginea forest, 22.IV. 1982, 1 female, 4.XII.1986 and 1 female, 25.1.1990, R. Bosmans leg. (CRB).</p> <p>- Tiemcen: S.E. Tiemcen, forêt d’Hafir, 1350m, 1 female in Quercus ilex litter, 23.V.1990, R. Bosmans leg. (CRB).</p> Distribution: Portugal, Spain, France, S. England, Austria, S.W. Finland, and cited here for the first time from Africa from Algeria, where it appears to be a species ofPublished as part of <i>Bosmans, Robert, 1999, The genera Agroeca, Agraecina, Apostenus and Scotina in the Maghreb countries (Araneae: Liocranidae), pp. 25-34 in Bulletin Di L' Institut Royal Des Sciences Naturelles De Belgique Entomologie 69</i> on pages 27-29, DOI: <a href="http://zenodo.org/record/826217">10.5281/zenodo.826217</a>
Zodarion jeanclaudeledouxi Bosmans & Benhalima 2020, sp. nov.
No full textZodarion jeanclaudeledouxi Bosmans & Benhalima sp. nov. Figures 63, 78–81 Etymology. The species is dedicated to the late Jean-Claude Ledoux (†), a well-known French arachnologist and author of many papers on Mediterranean spiders. Diagnosis. This species is recognized by the wide basal part of the tibial apophysis, suddenly narrowing into the distal part, a character shared with Z. mostafai sp. nov.. Both species differ from other members of this group by the larger embolar tooth, which is curved in the prolateral direction in Z. jeanclaudeledouxi sp. nov., while smaller and curved in the anterior direction in Z. mostafai sp. nov.. Description. Measurements: Male (n=1): total length 2.6; carapace 1.39 long, 1.01 wide. Colour (Fig. 63): as in Z. ericorum sp. nov.. Eyes: AM=1 (0.1); AL=0.62; PM=0.63; PL=0.75; a=0.5; b=d=0.25; c=1.62; MOQ: AW=0.81PW; L= 0.88PW. Male palp (Figs 78–81): tibial apophysis elongated, proximal part wide, rectangular, 3 times longer than distal part, suddenly narrowing into distal part, curved in retrolateral direction, terminaly rounded; median apophysis with wide proximal part and somewhat smaller, curved distal part; tip of embolus distinctly curved in antero-prolateral direction. Female: Unknown. Types. Holotype Ƌ from MOROCCO: Béni Mellal-Khénifra, Khénifra, Ouaoumana, 32°42’03’’N, 05°50’27’’W, 800 m a.s.l., leg. R. Bosmans, 17.IV.2012 (stones in wasteland) (RBINS). Distribution. Only known from the type locality (Map 3).Published as part of Benhalima, Souâd & Bosmans, Robert, 2020, Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae), pp. 93-114 in Zootaxa 4899 (1) on page 104, DOI: 10.11646/zootaxa.4899.1.5, http://zenodo.org/record/440070
Haplodrassus crassipes Bosmans & Kherbouche-Abrous & Benhalima & Hervé 2018, comb. n.
No full textHaplodrassus crassipes (Lucas, 1846), comb. n. Figs 1‒9, 18‒23, map 1 Drassus crassipes Lucas, 1846: 217, pl. 13 fig. 4 (♂♀). Drassus macellinus: Pavesi 1884: 467; Simon 1885: 39 (misidentifications). Drassodes macellinus: Simon 1908: 52; Simon 1914: 139; Denis 1937: 1035 (misidentifications). Haplodrassus macellinus: Di Franco 1994: 206 (misidentification). Haplodrassus macellinus hebes: Denis 1948: 146 (misidentification). Types. Lectotype ♂ (MNHN AR14576), paralectotype ♀ (MNHN AR14577) of Drassus crassipes, by present designation, from Algeria, La Calle (= El Kala); MNHN, Coll. Lucas; examined. Remarks. Type material of Drassus crassipes was discovered in the collection of the MNHN and could be compared with our material. It appears to be one of the most abundant Haplodrassus species in the Maghreb, in the past confused with H. macellinus. Diagnosis. Haplodrassus crassipes is the largest Haplodrassus species in North Africa, with males up to 10 mm and females up to 16 mm. Small specimens overlap however with other species, so size cannot be used as a sole diagnostic character. It differs from all other species in males by the S-shaped embolic apophysis with nearly parallel margins, terminally with a small dorsal tooth (Figs 4, 6, 19‒20). Females are most close to H. securifer sp. n. but differ by the epigyne being longer than wide (Figs 7, 22), as wide as long in H. securifer sp. n. (Figs. 145, 165). Description. Measurements: Male 5 (n=20): Total length 6.1‒10.3; carapace 2.8‒5.9 long, 2.2‒4.4 wide. Female (n=20): Total length 10.2 ‒ 16.4; carapace 3.4 ‒6.4 long, 2.2‒4.6 wide. Colour: Carapace brown to dark reddish brown, cephalic part infuscate; chelicerae dark brown; legs yellowish brown to dark reddish brown; abdomen dark brownish to dark greyish, venter pale. Spinulation: General pattern, but ventral spines on MtI and II sometimes missing and without prolateral spines on TiIV in all specimens examined. Male palp (Figs 3‒6, 18‒21): Tibial apophysis nearly twice as long as its diameter, distinctly constricted basally, distal part strongly asymmetric, with truncate dorsal margin; embolic apophysis S-shaped, with nearly parallel margins, terminally with a small dorsal nipple. Epigyne (Figs 7‒9, 22‒23): Distinctly longer than wide (1.3x); hood slightly narrower than areola, posterior margin nearly straight, with strongly recurved lateral parts; pro-fovea relatively large, as long as fovea, laterally with some longitudinal folds; fovea wider than long, elevated in the middle, antero-laterally with obtuse tooth, in most specimens hidden by concretions (compare Figs 7 & 8); areola oval, wider than long; receptacles large, touching; lateral pockets wide, semi-circular. Citations. ALGERIA, Annaba, Monts de l’Edough (36°55'38” E7°43'11”) (Di Franco 1994, as H. macellinus), El Tarf, El Kala (N36°53’44” E8°26’36”) (Lucas 1846), Jijel, Djébel Daya (N36°35’00” E6°11’00”) (Denis 1937, as D. macellinus, Souk-Ahras, Souk-Ahras (N36°17’11” E7°57’04”) (Simon 1885, as D. macellinus). TUNISIA, Beja, Aïn Draham (N36°46’43” E8°41’14”) (Simon 1908, as D. macellinus), Gabes, Gabes (N33°52’53” E10°05’54”) (Simon 1885, as D. macellinus), Tunis, Hammam-el-Lif (Simon 1885, as D. macellinus), Tunis, Porto Farina (N37°10’10” E10°11’26”) (Pavesi 1884, as D. macellinus). Further material examined. “ Algeria: Saïda! Djebel Maadid! Ouarsenis! Bône! (= Annaba) (E. Simon leg.), Theniet-el-Had (L. Bedel leg.). Tunisia ”, 1♂ 12♀, labelled D. macellinus, Coll. Simon 2916 (MNHN AR9216). ALGERIA, Alger, Bab Ezzouar (N36°42’45” E3°10’54”), 25m, 1♀, pitfalls in wasteland at University Campus, 29.IV.1987, 1♀, 15.III.1988, R. Bosmans leg. (CRB), 4♂ 6♀, 7.VII.2008 ‒ 22.V.2009 (COA), Aïn Defla, Miliana, Djébel Zaccar (N36°20’0” E2°13’00”), 1000m, 1♂, stones in Quecus ilex forest, 23.IV.1989, R. Bosmans leg. (CRB), Blida, Atlas Blidéen, Chréa, 1300m, 10♂, pitfalls in Cedrus atlantica forest, 27.VI.1989, R. Bosmans leg. (CRB), Chréa, Djébel Ferroukha, Ghellaï (N36°28’10” E2°56’18”), 1350m, 1♂, sieving litter in Cedrus atlantica plantation, 20.X.1987, R. Bosmans leg. (CRB), Chréa, Pic E. Abdelkader (N36°24’14” E2°50’6”), 1520m, 1♂, pitfalls in Cedrus atlantica forest, 27.V.1989, R. Bosmans leg. (CRB), S. Meftah, Djébel Zerouela (N36°36’0” E3°15’00”), 1♂, stones in grassland, 7.IV.1987, R. Bosmans leg. (CRB), Bouira, Er Rich (N36°25’28” E3°51’ 19’), 450m, 1♂, under stones, 15.IV.1990, R. Bosmans leg. (CRB), Massif du Djurdjura, Tikjda (N36°26'54” E4°7'44”), 1510m, 1♀, pitfalls in Cedrus atlantica forest, 11.VI.1988 and 1♂, 24.IV.1982, R. Bosmans leg. (CRB), Massif du Djurdjura, Tizi Boussouil (N36°27'37” E4°9'57”), 1720m, 4♂ 1♀, pitfalls in stony and spiny grassland, VIII.1990, O. Abrous leg. (COA, CRB), Massif du Djurdjura, Col de Tizi 'n Kouillal (N36°28'26” E4°14'9”), 1520m, 1♂, pitfalls in Amphelodesmus grassland, III.1990, R. Bosmans leg. (CRB), Saharidj, 650m, 1♂, pitfalls in dense Pistacea lentisca maquis, 1.VI.1988, R. Bosmans leg. (CRB), Boumerdes, between Larba and Col des 2 Bassins (N36°31'33” E3°13'30”), 800m, 5♀, stones in open Quercus ilex forest, 21.V.1987, R. Bosmans leg. (CRB), between Toulmout and Keddara (N36°37'17" E3°27'46”), 500m, 2♀, stony grassland near Ifri Nziri cave, 27.IV.1989, R. Bosmans leg. (CRB), Chlef, Forêt de Tacheta, 850m, 2♂, pitfalls in Quercus faginea forest, 29.IX.1989, R. Bosmans leg. (CRB), Sétif, Djébel Babor, N. slope (N31°30'35” E5°29'15”), 1600m, 1♂, stones in open Cedrus atlantica forest, 21.IV.1982, R. Bosmans leg. (CRB), Monts du Hodna, Djébel Bouthaleb (N35°43’25” E5°21’8”), 1450m, 1♀, pitfalls in Cedrus atlantica forest, 20.IV.1989, R. Bosmans leg. (CRB), Tissemsilt, 5 km N. Théniet-el-Had (N35°53'38” E2°3'11”), 880m, 1♀, litter in irrigated garden, 20.IV.1984, R. Bosmans leg. (CRB), Tizi Ouzou, Chabet-el-Ameur (N36°37'17” E3°43'46”), 250m, 2♂, stones bordering fields, 20.IV.1990 (CRB), Mazer, 5 km E. Tigzirt (N36°53'38” E4°2'5”), 50m, 1♂, litter in Pistacea lentisca maquis, 27.IV.1990, R. Bosmans leg. (CRB), 5 km E. Tizi Ouzou, (N36°42'26” E4°10'20”), 180m, 1♂, herbs on damp slope, 25.I.1990, R. Bosmans leg. (CRB), forêt de Mizrana (N6°52' 5' E4°4'5”), 300m, 1♀, stones in Quercus suber forest, 26.I.1990, R. Bosmans leg. (CRB), Massif du Djurdjura, Aït Ouabane (N36°28'1” E4°17'1”), 1♂, 1410m, pitfalls in Cedrus atlantica forest, 1.VI.1988, R. Bosmans leg. (CRB), Massif du Djurdjura, Tala Guilef (N36°28’36” E3°59'48”), 1420m, 29♂ 11♀, pitfalls in grassland, XI.1992 ‒ XI.1993, O. Abrous leg. (COA, CRB), Tirourda (N36°28'33” E4°20'9”), 1300m, 1♀, herbs at entrance of Maccabée cave (Ifri Megath), 27.IV.1989, R. Bosmans leg. (CRB), Tizi Ouzou (N36°42'42” E4°2'37”), 195m, 1♂, stones near hotel Amraoua, 25.I.1990, R. Bosmans leg. (CRB), Tlemcen, Mansourah, plateau de Lalla Setti (N36°4'52” E4°27'42”), 975m, 5♂ 3♀, litter in Pinus halepensis forest, 6.V.1984, R. Bosmans leg. (CRB). MOROCCO, Fès-Meknès, N. Timahdite (N33°14'11” W5°3'31”), 1800m, 1♂ 1♀, stones in grassland, 7.II.1996, R. Bosmans leg. (CRB), Marrakech-Safi, Marakech, Amizmiz (N31°13’26” W8°15’2”), 1♀, 7.II.1962, J. Malhomme leg. (ISRM). SPAIN, Melilla, 1♀, as Drassodes ariasi Simon nomen nudum, Arias leg., Coll. Simon 24784 (MNHN AR15691). Ecology. The species was mostly collected in forests at higher altitudes in the humid and subhumid climate zones of the Maghreb. Males were collected from September to June, females from January to June. Distribution. Tell Atlas in Algeria, Rif and Middle Atlas in Morocco (map 1). MAP 1. Distribution of Haplodrassus crassipes (Lucas).Published as part of Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), pp. 1-67 in Zootaxa 4451 (1) on pages 8-12, DOI: 10.11646/zootaxa.4451.1.1, http://zenodo.org/record/144508
Haplodrassus rufipes Bosmans & Kherbouche-Abrous & Benhalima & Hervé 2018, comb. n.
No full textHaplodrassus rufipes (Lucas, 1846), comb. n. Figs 126‒135, 140‒144, map 6 Drassus rufipes Lucas, 1846: 215, pl. 13, fig. 2 (♀). Drassus corticalis Lucas, 1846: 216, pl. 13, fig. 3 (♀), syn. n. Drassus severus C. L. Koch, 1843: 126, fig. 838 (not Drassus severus C. L. Koch, 1839 from Greece; misidentification of material from Italy). Drassus similis L. Koch, 1866: 103, pl. 4, fig. 67 (♂♀, from Italy), syn. n. Haplodrassus gridellii Caporiacco, 1949: 117 (♀), syn. n. Types. Holotype ♀ of Drassus rufipes Lucas from Algeria, Constantine (N36°21’54” E6°36’53”) (MNHN AR14588); MNHN, Coll. Lucas; examined. Type series of Drassus corticalis Lucas from Algeria, La Calle (= El Kala) 1♂ (MNHN AR14571), 1♂ (MNHN AR14752), 1♂ (MNHN AR14753), 1♀ (MNHN AR14754) 4♂ (MNHN AR14755), Coll. Lucas; examined. Syntypes ♂ and ♀ of Drassus similis from Italy, Sicilia; not examined. Diagnosis. Medium to large species. Males are recognised by the terminal and subterminal denticles of the embolic apophysis and the basal tooth of the embolus (Figs 129, 131, 140, 142). Females are recognised by a combination of characters, the wide hood, the fovea wider than long and the denticulate margin (Figs 132, 143), but this character is only visible in virgin females (compare Figs 132 and 134). Remarks on synonymy. C. L. Koch (1839) described the female of Drassus severus from Nafplio in Greece, and somewhat later (1843), also the male. The descriptions are accompanied by drawings of the general habitus of male and female, but not of the copulatory organs. On the drawings, spines on tibiae I and II are clearly visible and these are always absent in Haplodrassus species. Hence, the transfer to Haplodrassus cannot be justified. Furthermore, Simon (1914) states that the male and female were incorrectly matched: “C. Koch a plus tard (Ar. X, p. 126, fig. 838) attribué au D. severus un mâle très different, beaucoup plus voisin de D. lapidosus ”. C. L. Koch (1843) writes indeed: “Die taster ziemlich wie bei D. lapidicola ”. The taxonomic position of Drassus severus is retained in the genus Drassodes and declared a nomen dubium. The holotype female of Drassus rufipes Lucas, 1846 was examined and appears to be a subadult specimen with the epigyne completely developed under the integument. Abundant material of Drassus corticalis Lucas, 1846 was also found it the MNHN and this is considered to be the type series as well. All female specimens are identical to Drassus rufipes and by page priority D. corticalis becomes a junior synonym. C. L. Koch’s son (L. Koch 1866) described Drassus similis from Italy and was the first to present drawings of the male palp. Again, this is identical to the one of D. corticalis and D. similis also becomes a junior synonym of H. rufipes. Later, several authors cited H. severus from different localities (Simon 1885, 1899, 1909, 1914; Di Franco 1994), and these are all attributed to H. rufipes. The distribution of the species is visualised on map 8 and the type locality of Drassus severus in Greece falls out of the actually known distribution area of H. rufipes, another evidence this is another species. Without examining the type material, Levy (2004) synonymised Haplodrassus gridellii Caporiacco, 1949 with H. pugnans (Simon, 1880). However: (1) according to Caporiacco’s description of H. gridellii, the type specimen is 10.6 mm long, whereas H. pugnans has a maximal length of 9 mm (Levy 2004; Kamura 2007) (2) The type locality of H. gridellii in Libya falls out of the distribution area of H. pugnans (Sinai to Japan) (3) H. pugnans is not present in our abundant material from the Maghreb. There are two large Haplodrassus species in the Maghreb: Haplodrassus severus and H. signifer (table 1). Since Caporiacco (1949) states H. gridellii is related to H. signifer, and he never found H. severus, H. gridellii is considered a junior synonym of that species. Description. Measurements: Male (n=20): Total length 6.1‒8.9 mm; carapace 2.7‒3.9 long; 2.3‒2.7 wide. Female (n=20): Total length 7.2‒10 mm; carapace 2.4‒4.4 long, 2.2‒3.3 wide. Colour: Carapace dark reddish brown, cephalic part darkened; chelicerae dark brown, legs reddish brown; abdomen grey to dark grey, posterior part often with traces of 4‒5 paler chevrons. Spinulation: General pattern, but variable in tibiae and metatarsi III and IV. Male palp (Figs 128‒131, 140‒142): Tibial apophysis robust, somewhat dilating to the rounded top (Figs 130, 141); embolic apophysis elongated, curved, terminally pointed and with subterminal denticle (Figs 131, 142); embolus with basal tooth at prolateral margin (fig. 131). Epigyne (Figs 132‒135, 143‒144): Large, somewhat longer than wide; hood and pro-fovea wide, the latter with some transverse folds and narrower in the middle by the protruding hood; fovea as long as wide, antero-lateral margin with strong tooth (fig. 132) but mostly completed filled by a mating plug in mated females (fig. 134); areola deep dark-brown, elongated, anteriorly truncate and with two oblique grooves; lateral pockets semi-circular, with basal chamber (Figs 133, 144). Citations in the Maghreb (all as H. severus). ALGERIA, Batna, Timgad (N35°29’34” E6°28’11”) (Di Franco 1994), Boumerdes, Cap Matifou (N36°48’42” E3°13’23”) (Simon 1899), Constantine, surroundings of Constantine (N36°21’54” E6°36’53”) (type locality of Drassus rufipes Lucas, 1846), Djelfa, Takersane (N34°24’44” E2°54’16”) (Simon 1899), Jijel, Texenna, Mont M’sid Ech Cheta (N36°40’53” E5°48’17”) (Di Franco 1994), M’sila, surrounding of Baniou (N35°24'55” E4°20'39”) (Di Franco 1994), Tizi Ouzou, Massif du Djurdjura, Bou Adenane (N26°30’10” E4°14’57”) (Di Franco 1994), Yakouren (N36°45'44” E4°24'57”) (Di Franco 1994). LYBIA, Shahhat, Wadi Ben Gadir near Cyrene (Caporiacco 1949, as H. gridellii). MOROCCO, without precise locality (Simon 1909), Beni Mellal-Khénifra, Oued Sefrou (N32°33'0" E6°9'0") (Simon 1911), Fès-Meknès, Nzala Beni Ammar (N34°07’24” W5°25’54”) (Di Franco 1994), Tanger-Tétouane-Al Hoceima, Bab Berred (Di Franco 1994), Ketama (Di Franco 1994), road Ketama-Bab Besen (N34°57'53” W4°39'29”) (Di Franco 1994). TUNISIA, Jendouba, Aïn Draham, Col des Ruines (N36°47'30” E8°41'5”) (Di Franco 1994), Aïn Draham (N36’46’43” E8°41’14”) (Simon 1885), Siliana, Maktar (N35°51’2” E9°12’03”) (Di Franco 1994). Further material examined. AFRICA: ALGERIA, Alger, Bab Ezzouar, universtiy campus (N36°42’45” E3°10’54”), 25m, 1♂, pitfalls in rough grassland, 28.IV.1987, R. Bosmans leg. (CRB), idem, 1♂ 3♀, 7.VII.2008 ‒ 22.V.2009 (COA), El Harrach (N36°42’46” E3°8’56”), 25m, 7♂, pitfalls under Juniperus in garden of I.N. A., 7.III‒ 11.IV.1984, R. Bosmans leg. (CRB), Oued Smar (N36°41’21” E3°8’58”), 30m, 1♂, pitfalls in wheat fields, 12.IV.2008, I. Feriel leg. (CRB), Aïn Témouchent, Aïn Tolba (N35°14'56” W1°15'1”), 250m, 1♂, stones in dry grassland, 19.I.1990 (CRB), Annaba, Chetaibi (N37°2'42” E7°21'59”), 810m, 2♀, stones in grassland, 1.III.1990, R. Bosmans leg. (CRB), Hippône N., Edough National park (36'55'38” E7°43'11”), 820m, 1♂, seeving litter in Quercus faginea forest, 23. XI.1989, R. Bosmans leg. (CRB), Batna, Djébel Chélia (N35°21'52” E6°38'43”), 1600m, 1♂, pitfalls in Cedrus atlantica forest, 1.VII.1988, R. Bosmans leg. (CRB), Blida, Atlas Blidéen, Chréa, Pic Abdelkader (N36°24’14” E2°50’6”), 1550m, 2♂, stones in grassland, 15.II.1987, R. Bosmans leg. (CRB), Atlas Blidéen, Djébel Ferroukha, Ghellaï (N36°28’10” E2°56’18”), 1350m, 1♂, pitfalls in Cedrus atlantica plantation, 29.IV.1987, idem, 1600m, 1♂, 27.VI.89, R. Bosmans leg. (CRB), Djébel Mouzaia (36°22’0” E2°41’45”), 1250m, 8♂ 2♀, stones around lake, 14.V.1988, R. Bosmans leg. (CRB), Meurdja (N36°29'59” E3°8'54”), 955m, 1♂, stones in recent Cedrus atlantica plantation, 1.IV.1982, R. Bosmans leg. (CRB), Bordj Bou Arreridj, Sidi Embarek (N36°6'5” E4°54'30”), 900m, 1♂, stones in fields, 27.II.1990, R. Bosmans leg. (CRB), Boumerdes, between Toulmout and Keddara (N36°37'17" E3°27'46”), 500m, 1♂ 1 subadult ♀, stones in grassland, 27.IV.1989, R. Bosmans leg. (CRB), Djelfa, Djébel Sénalba (N34°33'50” E3°2'9”), 1330m, 1♂, pitfalls in Pinus halepensis forest, 28.III.1989, 1♂, 9.IV.1991, N. Bouragba leg. (CRB), Djébel Djellal (N34°23'32” E2°58'42”), 1350‒1450m, 1♂, pitfals in Pinus halepensis plantation, III.1989 ‒ IV.1990, N. Bouragba leg. (CRB), El Tarf, El Kala, west side of lake Tonga (N36°52'35” E8°29'28”), 10m, 1♀, wet grassland along the lake, 27.III.1988, R. Bosmans leg. (CRB), E. El Kala, Lac Melah (N36°52'55” E8°20'37”), 3m, 1♀, litter in Quercus suber forest, 6.IV.1982, R. Bosmans leg. (CRB), Medea, Col de Beni Chicao (N36°12'12” E2°53'0”), 1230m, 1♂, litter in small Quercus ilex forest, 21.XII.1987, R. Bosmans leg. (CRB), Mostagenem, Stidia (N35°48'39” W 03'0”), 25m, 1♂, pitfalls in Pistacea lentisca maquis in dunes, 25.V.1990, R. Bosmans leg. (CRB), M’sila, S. Aïn Oghrab (N35°0’52” E4°6’24”), 650m, 1♂, pitfalls in Pinus halepensis forest, 20.III‒23.VI.1990, R. Bosmans leg. (CRB), between Aumale (= Sour-El Ghozlane) and Bou Saada (N35°20’36” E6°4’46”), Aïn Oghrab (N35°0’52” E4°6’24”), 1♂, E. Simon leg., Coll. Simon 12988 (MNHN AR15702), Saida, Tircine (N34°54'4” E0°33'16”), 900m, 1♀, stones bordering fields, 4.IV.1989, R. Bosmans leg. (CRB), Sétif, Djébel Babor, south slope (N31°30'35” E5°29'15”), 1300m, 1♀, liter in irrigated garden, 19.IV.1982, R. Bosmans leg. (CRB), Skikda, Ben Azouz, 200m, 2♂ 2♀, open Eucalyptus plantation, 2.III.1990, R. Bosmans leg. (CRB), Bouchata, 400m, 2♂, stones in grassland, 12.III.1990, R. Bosmans leg. (CRB), Tébessa, Tébessa, forêt de Bekkaria (N35°21'20” E8°14'40”), 1300m, 1♂, pitfalls in Pinus halepensis forest, 1.V.1989, R. Bosmans leg. (CRB), Tissemsilt, Massif de l'Ouarsenis, Bou Caid (N35°53'2” E1°37'49”), 1400m, 2♀, sieving litter in Cedrus atlantica forest, 24.III.1988, R. Bosmans leg. (CRB), Djébel Ouarsenis, Théniet-el-Had, 1550m (N35°52'15” E1°56'41”), 1♀, stones in grassland, 3.V.1984 (CRB), Tizi Ouzou, Massif du Djurdjura, Tala Guilef (N36°28’36” E3°59'48”), 1400m, 6♂, stones in grassland, 23.V.1984, R. Bosmans leg. (CRB), idem, 1♂, litter in Cedrus atlantica forest, 29.IV.1984, R. Bosmans leg. (CRB), idem, 1420m, 35♂ 27♀, pitfalls in grassland, XI.1992 ‒ XI.1993, O. Abrous leg. (CRB), Djébel Heizer, 1800m, 2♂, stones in montane grassland, 17.IV.1989, R. Bosmans leg. (CRB), Col de Tizi N’Kouilal (N36°28'26” E4°14'9”), 1510m, 1♂, pitfalls in grassland, 1.XII.1991, R. Bosmans leg. (CRB), Tlemcen, N. Sebdou, source de l'Oued Tafna (N34°41’30” W1°18’48”), 1700m, 1♂, under stones, 18.I.1990, R. Bosmans leg. (CRB), S. Tlemcen, Terny plain (N34°48'16” W1°22'1”), 1175m, 1♂, stones in grassland, 18.I.1990, R. Bosmans leg. (CRB). MOROCCO, Béni Mellal-Khénifra, Moulay Bouazza, Oued Tarralha (N33°15’23” W6°7’54”), 1♀, 8.III.1952, J. B. Panouse leg. (ISRM), Fès-Meknès, plaine du Saïss, Aïn Taoujdate (N33°56’8” W5°12’41"), 480m, 4♂ 4♀, pitfalls in wheat fields, 30.IV.1997, S. Boksch leg. (CRB), Azrou NE. (N33°28'00” E5°9'49”), 1400m, stones in Quercus ilex forest, 18.IV.2012, R. Bosmans leg. (CRB), Azrou N. (N33°28'1” W5°12'8”), 1400m, 3♂, stones in grassland, 7.II.1996, R. Bosmans leg. (CRB), road Azrou-Khénifra (N33°12'53” W5°35'52”), 1200m, 1♂ 1 subadult ♀, 18.II.1952, J. B. Panouse leg. (ISRM), N. Timahdite (N33°14'11” W5°3'31”), 1800m, 1♂ 1♀, stones in grassland, 7.II.1996, R. Bosmans leg. (CRB), Rabat-Salé-Kénitra, Oulmès (N33°25’25” W 6°00’ 10), 1♂, 11.III.1955, J. B. Panouse leg. (ISRM), Tanger-Tetouan-Al Hoceima, 10 km E. Bab Berred (N34°59'40” W4°50'43”), 1525m, 1♂ 1 subadult ♀, stones in Quercus faginea forest, 15.V.1984, R. Bosmans leg. (CRB), Issaguen S. (N34°53'25” W4°34'59”), 1450m, 4♂, stones in Cedrus atlantica forest, 15.XII.2013, R. Bosmans leg. (CRB). TUNISIA, Bizerte, Fejja-Aïn Guellal road (N37°2'25” E9°55'17”), 1♂, stones bordering fields, 29.I.2003, R. Bosmans leg. (CRB), Lake Ichgeul (N37°6'48” E9°42'8”), 1♂ 1♀, stones in the plains, 25.I.1995, R. Bosmans & J. Van Keer leg. (CJVK, CRB), 1♂, 29.I.2003, R. Bosmans leg. (CRB), Jendouba, Aïn Sebaa E., Oued Titria (N36°57’00” E8°56’00”), 200m, 6♂, stones in open Quercus suber forest, 28.II.2005, R. Bosmans & J. Van Keer leg.(CJVK, CRB), Esaouani, 300m, 10♂, 6.III.2005, R. Bosmans & J. Van Keer leg. (CJVK, CRB), Chemtou ruins (N36°29'40” E8°34'16”), 250m, 3♂, stones in Roman ruins, 6.III.2005, J. Van Keer leg. (CJVK), Forêt de Feidja (N36°26'27” E8°22'55”), 600m, 1♂, stones in Quercus ilex forest, 5.II.2005, R. Bosmans leg. (CRB), Fernana N. (N36°42'21” E8°40'47”), 450m, 1♂, stones in maquis, 6.III.2005, J. Van Keer leg. (CJVK), Ghardimaou W. (N36°27'27” E8°24'54”), 250m, 1♂, stones in grassland, 5.III.2005, J. Van Keer leg. (CJVK), Kasserine, Haidra S. (N35°32'41” E8°27'27”), 950m, 1♂, stones in pine forest, 4.III.2005, R. Bosmans leg. (CRB), Haidra (N35°36’01” E8°23’53”), roman ruins, 8♂ 3♀, stones in ruins, 4.III.2005, R. Bosmans & J. Van Keer leg. (CJVK, CRB), Thala S. (N35°33'37” E8°40'39”), 950m, 5♂, stones in steppe, 4.III.2005, R. Bosmans & J. Van Keer leg. (CJVK, CRB), Le Kef, Le Kef, bifurcation to Ben Hazen, 1♂, 21.IV.1981, B. Valle leg. (MCSNB), Hammam Mellègue (N36°6'59” E8°29'23” E), 800m, 1♂, stones bordering small Pinus forest, 4.III.2005, R. Bosmans leg. (CRB), Touiref SE (N36°16'44” E8°27'3”), 650m, Oued Mellègue, 3♂, 5.III.2005 (CJVB, CRB), Nabeul, Aïn Tebournok (N36°34'47” E10°28’3”), 100m, 1♂, stones bordering a river, 28.I.2003, R. Bosmans leg. (CRB), Tazerka (N36°38'30” E10°54'34”), 1♂, in litter in salt marsh, 26. I.2 0 0 3, R. Bosmans leg. (CRB), Siliana, Kesra N. (N35°49'27” E9°21'36”), 1050m, 1♂, stones bordering Pinus forest, 27.I.2003, R. Bosmans leg. (CRB), Tunis, La Goulette (N36°9'51” E10°18'8”), 25m, 2♂, stones in Pinus plantation, 30.I.2003, R. Bosmans leg. (CRB), Zaghouan, Djébel Zaghouan (N36°23'21” E10°8'12”), 500m, 1♂, stones in Pinus halepensis, forest, 24.I.1995, R. Bosmans leg. (CRB), Zriba Village (N36°21'52” E10°14'47”), 100m, 2♂ 1 subadult ♀, stones along Oued El Hammam, 24.I.1995, R. Bosmans leg. (CRB). EUROPE: PORTUGAL, Faro, Tavira (N37°6'57” W7°37'37”), 5m, 3♂, stones in salt marsh, 7.II.2006, R. Bosmans leg. (CRB), Santarém, Porto Alto S. (38°45'48” N8°54'18” W), 30m, 2♂, pitfalls in a Quercus suber forest, 14‒22.IV.2013, R. Bosmans leg. (CRB), Setúbal, Alcochete NW (38°45'37” N8°56'6” W), 2m, 1♀, pitfalls in salt marsh, 14‒22.IV.2013, R. Bosmans leg. (CRB), Gambia, Mourisca (N38°32'59” W8°45'28”), 2m, 2♂, pitfalls in salt marsh, 14‒22.IV.2013, R. Bosmans leg. (CRB), Vila Real, Mondim de Basto, road to Nuestra Sierra de Gracia (N41°25'22” W7°55'32”), 480m, 1♂, stones in grassland, 14.XI.2016, R. Bosmans leg. (CRB). SPAIN, Alicante, Aspe, 250m, 1♀, stones in dry maquis, 4.IV.1996, R. Bosmans leg. (CRB), Avila, Monbeltran (40°15‘35” N5°1’2”), 1♂, P. Poot leg. (CRB), Baleares, Mallorca, Embassement de Cuber (N39°46'49” N2°46'58” E), 850m, 4♂ 1♀, stones in grassland, 2.IV.2003, R. Bosmans leg. (CRB), Caceres, Monfragüe (N9°49'31” W5°56'55”), 250m, 1♀, pitfalls near a spring, 15.IV.1994, R. Bosmans leg. (CRB), Cádiz, Algeciras (N36°7’59” W5°27’1’’), 1♂ 5♀, without date or collector (NHML), Tarifa (N36°0’50” W5°36’25”), 2♂ 1♀, III.1983, 2 ♂ 1♀, IV.1990, P. Poot leg. (CRB), Ciudad Real, Laguna del Camino de Villafranca (N39°24'48” W3°48'33”), 1♂, grassland around lake, 13.IV.1998, R. Bosmans leg. (CRB), Granada, Sierra de la Contreviesa, Puerta Camacho (N36°50'6” W3°21'10”), 1230m, 1♀, stones in Pinus forest, 6.IV.1997, R. Bosmans leg. (CRB), Lacallahora (N37°11'3” W3°3'50”), 1200m, 1♀, stones on slopes to castle, 5.IV.1996, R. Bosmans leg. (CRB), Huesca, Ontiñena S. (N41°38'35” E0°5'21”), 300m, 3♀, stones in dry river bed, 1.IV.1996, R. Bosmans leg. (CRB), Jaen, Puerto de las Palomas (N37°57'11” W2°56'17”), 1290m, 3♀, stones in wasteland, 12.IV.1999, R. Bosmans leg. (CRB), Málaga, Alozaina N. (N36°44'53” W4°50'33”), 250m, 1♀, litter in Quercus suber forest, 11.IV.1998, R. Bosmans leg. (CRB), Archidona E. (N37°6'34” W4°20'49”), 2♂, stones in grassland, 18. XII.1997, R. Bosmans leg. (CRB), Archidona E., La Pena (N37°3'43” W4°28'50”), 750m, 1♀, stones in grassland, 11.IV.1998, R. Bosmans leg. (CRB), Benalmadena N. (N37°3'43” W4°28'50”), 1♂, stones and litter near the beach, 20.XII.1997, R. Bosmans leg. (CRB), Casares (N36°26’44” W5°16’13”), 650m, 1♂ 1♀, pitfalls in Quercus suber forest, V.1986, J. De Beir leg. (CRB), Coin (N36°40'37” W4°45'57”), 150m, 2♂, stones along Rio Seco 19.XII.1997, R. Bosmans leg. (CRB), El Torcal de Antequera (N36°57'50” W4°32'36”), 900m, 1♀, stones in grassland, 11.IV.1998, R. Bosmans leg. (CRB), Torrremolinos (36°37’13” W4°29’59”), 1♂ 2♀, IV.1975, P. Hillyard leg. (NHML), Murcia, Cartagena (N37°36’18” W0°59’10”), 1♀, E. Simon leg., Coll. Simon 13414 (MNHN AR15703), Totana E., Sierra de la Tercia (N37°44'29” W1°33'20”), 300m, 2♀, stones in maquis, 4.IV.1996, R. Bosmans leg. (CRB), Rioja, Navarrete (N42°25'41” W2°33'49”), 420m, 1♀, pitfalls in grassland, 7.IV.1992, R. Bosmans leg. (CRB), Salamanca, Ciudad Rodrigo N. E. (N40°38'10” W6°29'12”), 650m, 1♂ 1♀, stones in maquis, 10.IV.1996, R. Bosmans leg. (CRB), Zaragoza, Escatron, Monasterio Rueda (N41°19'23” W0°20'37”), 225m, 1♂, stones in open Pinus forest, 1.IV.1996, R. Bosmans leg. (CRB). FRANCE, mixed unknown localities (‘Gallia Meridionalis’), one of them Alpes-Maritimes, Menton, 1♂ 2♀, as Drassodes macellinus hebes, Coll. Simon 2083 (MNHN AR15707), Bouches-du-Rhône, Graveson (N43°51’4” E4°46’18”), 1♀, 14.IV.1989, P. Poot leg. (CRB), Pyrénées-Orientales, Saint-André, 1♂, in swimming pool, 3.XII.15, G. Melotti leg. (CPO). ITALY, Lazio, Viterbo, Tarquinia, Sant’Agostino, Foce Mignone (N42°15’15” E11°45’23”), 1♂, 8.I.2004, 1♂, 17.III.2004, pitfalls, L. Pavesi & D. Mariotti leg. (MCSNB), Sicilia, Palermo, Terrasini, Riserva Naturale Orientata Capo Rama (N38°8'22” E13°3'38”), 2♂, 17‒27.II, 3♂ 13‒30.III, 2♂ 1♀, 14‒24.IV.2012, pitfalls in palm grove, C. Cusimano leg. (MCSNB). MAP 6. Distribution of Haplodrassus rufipes (Lucas) (closed red circles: verified localties; open red cercles: unverified citations of H. severus from literature; yellow square: type locality of H. severus). Ecology. The species occurs in all kind of habitats at low and high altitudes, even in cultivated land. In the Maghreb it is confined to the humid and subhumid climate zones. Distribution. The species occurs in the north of Algeria, Tunisia and Morocco, and in Portugal, Spain, France and Italy (map 6).Published as part of Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), pp. 1-67 in Zootaxa 4451 (1) on pages 33-39, DOI: 10.11646/zootaxa.4451.1.1, http://zenodo.org/record/144508
Zodarion ericorum Benhalima & Bosmans 2020, sp. nov.
No full textZodarion ericorum Bosmans sp. nov. Figures 61–62, 74–77, 98–101 Etymology. The first author dedicates this species to Eric Gobbers (Gent, Belgium) and Erika Vanden Bergh (Instituut voor Natuur- en Bosonderzoek, Belgium), two Belgian biologists, for helping him collect material during his first trip to Morocco in 1984. Diagnosis. Males are recognized by the large basal knob in the tegulum, which is absent in other species or only poorly developed in Z. jeanclaudeledouxi sp. nov. Females are distinguished by the shape of the transverse oval pit in the epigyne, which is not the same in all other species. Description. Measurements. Male (n=4): total length 2.4–3.8; carapace 1.30–1.57 long, 0.90–1.1 wide. Female (n=7): total length 3.8–5.5; prosoma 1.76–3.4 long, 1.25–2.49 wide. Colour (Figs 61–62): carapace dark reddish brown to black, posterior part sometimes yellowish orange of variable size; coxae pale yellowish, femora dark brown to black, basal part white; tibiae brown to pale brown, with paler stripes laterally; metatarsi yellowish brown, base often infuscated; tarsi yellowish brown; abdomen in male with scutum, dark sepia, dorsally with two triangular spots and one oval postero-median spot whitish, in some specimens triangular spots absent; venter and oblique lateral stripe whitish. Female as male, but abdomen without scutum. Eyes: AM=1 (0.13); AL=0.78; PM=PL=0.67; a=0.67; b=0.23; c=1.56; d=0.33; MOQ: AW=0.88PW; L=0.81PW. Male palp (Figs 74–77): tibial apophysis elongated, proximal part nearly rectangular and straight, distal part curved in postero-dorsal direction, terminally rounded; retrolateral margin of cymbium with indentation; tegulum with rounded knob at its base; median apophysis with broad base, rectangular, distal part having the form a bird’s beak; tip of embolus slender, terminally curved in antero-prolateral direction. Epigyne (Figs 98–101): with large, transverse, oval pit, somewhat more than 2 times as wide as high, posterolaterally acccompied by semi-circular pockets; spermathecae small, separated by more than 3 diameters. Type material. Holotype Ƌ and 1Ƌ 1♀ paratypes from MOROCCO: Beni Mellal-Khénifra, Oued Zem, 33°51’51’’N, 06°34’12’’W, leg. E. Vanden Bergh, 12.V–23.VI.1984 (pitfalls in young Pinus plantation) (RBINS). Other material examined. MOROCCO: Marrakech-Safi, Ben Guerir 10 km N, 32°14’32’’N, 07°56’46’’W, 500 m a.s.l., leg. R. Bosmans, 9.II.1996 (grassland with Asphodelus), 1♀ (CRB); 10 km S Chichaoua, 31°15’49’’N, 08°50’38’’W, leg. R. Bosmans, 9.II.1996 (stones in steppe), 2♀♀ (CRB); Marrakech, Gueliz, 31°41’37’’N, 08°00’21’’W, 445 m a.s.l., leg. R. Bosmans, 7.VII.1999 (grass field in flooded hotel garden), 1Ƌ (CRB); 3 km S Imi-’n-Tanoute, 31°09’35’’N, 08°50’38’W, 1200 m a.s.l., leg. J. Van Keer, 9.II.1996 (stones in olive groove), 1Ƌ 1♀ (CJVK); Ounara E, 31°32’33’’N, 09°30’47’’W, 250 m a.s.l., leg. R. Bosmans, 8.VII.1999 (stones in Arganus steppe), 1♀ (CRB); Rabat-Salé-Kénitra, Forêt de Maâmora, Sidi Amira, 34°03’06’’N, 06°43’11’’W, leg. S. Benhalima, 5.XII.1989 (litter in Quercus suber forest), 1Ƌ 1♀ (CSB); idem, 2 km S Sidi Allal El Bahraoui, 34°02’41’’N, 06°35’25’’W, leg. R. Bosmans, 8.II.1996, 1♀ (CRB); idem, Sidi Allal El Bahraoui, leg. S. Benhalima, 15.IX.1989, 1Ƌ, 2.X.1989, 1Ƌ, 6.XI.1989, 1Ƌ and 27.II.1990, 1Ƌ (CSB); Souss-Massa, between Aoulous and Taliouine, 30°34’41’’N, 08°03’11’’W, 600 m a.s.l., leg. R. Bosmans, 4.II.1996 (stones in Arganus steppe), 1♀ (CRB). Distribution. Central and West Morocco (Map 3).Published as part of Benhalima, Souâd & Bosmans, Robert, 2020, Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae), pp. 93-114 in Zootaxa 4899 (1) on pages 103-104, DOI: 10.11646/zootaxa.4899.1.5, http://zenodo.org/record/440070
Apostenus algericus Bosmans, 1999, n. sp.
No full textApostenus algericus n. sp. (Figs. 29-22) Type material: Holotype male from Algeria, Wilaya of El Bayadh, N.E. Les Arbaouattes, 900m, in Nerium oleander litter, 20.I.1988, R. Bosmans leg.; deposited in IRSNB. Diagnosis: Closely related to Apostenus maroccamis n. sp. differing by the subterminally curved median apophysis and the gradually narrowing embolus. Description: Measurements: Male: Total length 2.3; carapace 1.14 long, 0.86 wide. Colour: Carapace yellowish orange, before fovea, region of striae with some grey; legs uniformly yellowish orange; abdomen brown with some greyish markings. Spination as in preceding species. Palp (Figs. 29-30): Tibia with oblique, pointed apophysis; median apophysis gently curved, subterminally curved in anterior direction; embolus thick, gradually narrowing. Other material examined: None. Distribution: Only known from the type locality in Algeria.Published as part of Bosmans, Robert, 1999, The genera Agroeca, Agraecina, Apostenus and Scotina in the Maghreb countries (Araneae: Liocranidae), pp. 25-34 in Bulletin Di L' Institut Royal Des Sciences Naturelles De Belgique Entomologie 69 on page 33, DOI: 10.5281/zenodo.82621
Zodarion mostafai Benhalima & Bosmans 2020, sp. nov.
No full textZodarion mostafai Benhalima & Bosmans sp. nov. Figures 64–65, 82–85, 102–105 Etymology. The species is with pleasure and great love dedicated to Mostafa Bounouar (Rabat, Morocco), husband of the first author of this paper. Diagnosis. For males, see Z. jeanclaudeledouxi sp. nov.. Females can be distinghuished from the other species of this group by the presence and the shape of a trapezoid knob in the anterior part of the epigyne, which is absent in the other species. Description. Measurements: Male (n=3): total length 3.5–5.6; carapace 1.86–3.00 long, 1.2–2.2 wide. Female (n=2): total length 4.2–5.3; carapace 1.95–2.30 long, 1.25–1.55 wide. Colour (Figs 64–65): as in Z. ericorum sp. nov., but in males the tibiae are completely black and the pale part of legs more yellowish, and in the female only Ti IV with proximal part white. Eyes AM=1 (0.12); AL=PL=0.89; PM=0.63; a=0.63; b=0.2; c=0.16; d=0.32; MOQ: AW=0.93PW; L=0.96PW. Male palp (Figs 82–85): tibial apophysis elongated, proximal part relatively wide compared to other species, nearly rectangular, distal part a short knob curved in retrolateral direction; median apophysis with trapezoid proximal part and small, pointed distal part; tip of embolus short, curved in antero-prolateral direction. Epigyne (Figs 102–105): with anteromedian trapezoid structure; posterior margin of epigyne deeply incised; trapezoid structure covering 2 median pockets; spermathecae small, separated by 5 diameters. Type material. Holotype Ƌ and 1♀ paratypes from MOROCCO: Rabat-Salé-Kénitra, Forêt de Maâmora, Sidi Amira, 34°03’06’’N, 06°43’11’’W, leg. S. Benhalima, 6.XI.1989 (litter in Quercus suber forest) (ISRM). Paratypes: Same data as holotype, 1Ƌ 1♀ (MNHN). Other material examined. MOROCCO: Rabat-Salé-Kénitra, Forêt de Maâmora, Sidi Allal-El Bahraoui, 34°07’36”N, 06°33’22”W, leg. S. Benhalima, 12.II.1988 (under bark of Quercus suber), 1Ƌ 1♀ (CSB); Forêt de Maâmora, 2 km N Sidi Allal-El Bahraoui, 34°02’41’’N, 06°35’25’’W, leg. S. Benhalima, 3.II.1989, 1Ƌ (CSB); idem, leg. R. Bosmans, 8.II.1996 (litter in Q. suber forest), 1♀ (CRB). Distribution. Only known from the Maâmora forest (Map 3).Published as part of Benhalima, Souâd & Bosmans, Robert, 2020, Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae), pp. 93-114 in Zootaxa 4899 (1) on pages 104-105, DOI: 10.11646/zootaxa.4899.1.5, http://zenodo.org/record/440070
Zodarion ogeri Bosmans & Benhalima 2020, sp. nov.
No full textZodarion ogeri Bosmans & Benhalima sp. nov. Figures 66, 86–89 Etymology. The species is dedicated to our friend Pierre Oger (Waret L’Evêque, Belgium), eminent photographer of all species treated in this paper. Diagnosis. Males are recognized by the tibial apophysis that is relatively wide, nearly straight, terminally truncate and with a small incision, while more curved or terminally pointed in the other species. Zodarion jeanclaudeledouxi sp. nov. is closely related but differs by the longer recurved tip of the tibial apophysis. Description. Measurements: Male (n=1): total length 4.1; carapace 3.12 long, 1.23 wide. Colour (Fig. 66): Carapace dark brown to black; femora black; other segments white to pale yellowish, tibiae laterally striked with brown; abdomen black, with 3 dorsal spots and oblique lateral white stripes. Eyes: AM=1 (0.14); AL=PL=0.77; PM=0.55; a=0.73; b=0.36; c=1.55; d=0.27; MOQ: AW=0.97PW; AW=0.72L. Male palp (Figs 86–89): tibial apophysis elongated as in all species of this group, but proximal part nearly rectangular, more than 4 times longer than wide, terminally incised and curved in posterior direction; proximal part of median apophysis with triangular base, distal part a slender, pointed tooth; embolus slender, curved in anterior direction. Female: Unknown. Type material. Holotype Ƌ from MOROCCO: Meknès, Oued Beth, 33°52’41’’N, 05°55’54’’W, 500 m a.s.l., leg. R. Bosmans, 7.II.1996 (stones in Pinus plantation) (RBINS). Distribution. Only known from the type locality in the region of Meknès (Map 3).Published as part of Benhalima, Souâd & Bosmans, Robert, 2020, Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae), pp. 93-114 in Zootaxa 4899 (1) on page 105, DOI: 10.11646/zootaxa.4899.1.5, http://zenodo.org/record/440070
Lepthyphantes ritae Bosmans 1985
No full textLepthyphantes ritae Bosmans, 1985 ; IG 27782; Algerie, Zemmouri; 5 m, 13.IV.82; litter of Pinus halepensis. Lepthyphantes ritae Bosmans, 1985; R. Bosmans det. 1984, IG 27782; Maroc; 10 km, E. Chechaouen; 500 m, Quercus suber forest, 15. V.1984, R. Bosmans leg.Published as part of Barrientos, José Antonio, Brañas, Neus & Mederos, Jorge, 2020, The high complexity of Micronetinae Hull, 1920 (Araneae, Linyphiidae) evidenced through ten new cave-dweller species from the Morocco, pp. 1-29 in Zoosystema 42 (1) on page 4, DOI: 10.5252/zoosystema2020v42a1, http://zenodo.org/record/362705
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