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Platystoma geula Bodner & Freidberg, 2016, n. sp.
No full textPlatystoma geula n. sp. (Figs. 12, 23, 34, 45, 56, 67, 81, 97) Diagnosis. This species generally matches the characteristics of the Platystoma lugubre species group (Hennig, 1945) with the following exceptions: presence of 2–3 supra-alar setae, as opposed to 4; absence of setulae on the scutellum; black ground color of the postpronotum, as opposed to red (differs from all species in the group except P. lugubre corsicanum Séguy). This species was also compared with specimens of similar species, P. t e gu l a r u m Loew and P. gemmationis (Rondani), from which it differs by a much shorter ventral calypter. Description. Head (Figs. 12, 23): Color and vestiture: Head conspicuously pale to dark brown, with yellowish-brown areas. Frons microtrichose mostly medially, at orbits and around bases of orbital setae, with 4 orange to brown patches delimited by inconspicuous transverse band of microtrichia at middle, and by median line of microtrichia. Ocellar triangle inconspicuously microtrichose, delimited by dense yellowish microtrichia. Lunule yellow to pale brown, with yellowish microtrichia, especially medially. Antenna: scape pale to dark brown, pedicel yellow to orange and 1st flagellomere dark brown, all microtrichose (not visible in certain angles). Face mostly blackish, except yellowish to brownish areas lateroventrally; dense gray microtrichia on most of dorsal part and medioventrally, leaving ventral 0.20–0.33 of antennal groove shiny black without microtrichia, and lateroventrally without microtrichia although matte. Clypeus mostly densely microtrichose, with small lateral shiny area (depending on angle of observation). Chaetotaxy: 2 (pairs) orbital setae and 1 (pair) postocellar seta present. Lunule setulose. Postgena and occiput with long yellowish setulae. Ratios: Head: 1.41–1.51; frons anterior: 1.53– 1.83; frons posterior: 1.32–1.50; ocellar triangle/frons: 0.29–0.38; eye: 1.95–2.24; 1st flagellomere: 1.60–2.13; gena/eye: 0.36–0.41; parafacial/gena: 0.29–0.39; posterior orbital/anterior orbital: 2.33–2.50; medial vertical/ lateral vertical: 0.69–1.17; posterior orbital/medial vertical: 0.56–0.81; ocellar/medial vertical: 0.37–0.54; ocellar/ ocellar triangle: 0.83–1.08; postocellar/medial vertical: 0.23–0.35. Thorax: Color and vestiture: Ground color black, with yellowish-gray microtrichia. Scutum with 3 or 5 microtrichose stripes. Postpronotal lobe ground color black. Scutellum medially and laterally microtrichose, with sublateral black spots, sometimes including round black spots around bases of setae. Subscutellum black, with grayish-yellow microtrichia, mostly laterally and near median. Pleura microtrichose as in genus. Chaetotaxy: Presutural supra-alar seta absent. 0–1 postsutural supra-alar seta present. Postpronotal lobe with yellowish setulae anteriorly and black setulae posteriorly. Anepisternum uniformly short setulose. Katepisternum with long curly yellowish-white to orange setulae. Ratios: Scutum: 0.81–0.95; postsutural supra-alar/intra-alar: 0.33–0.50; postalar/intra-alar: 0.46–1.09. Legs: Color and vestiture: Leg type dark. Tarsi: fore and mid tarsi black, with yellow areas on proximal part of tarsomeres; hind metatarsus mostly yellowish, dorsoapically pale brown; 2nd tarsomere of hind tarsus yellowish ventrally and dorsally on proximal half. Other segments as fore and mid tarsi. Coxae anterodorsally microtrichose, posterodorasally shiny black. Chaetotaxy: Coxae and trochanters with mixed yellow and blackish setae and setulae, posterodorsally bare. Femora dorsally with blackish and mostly short setulae, with long and thicker setulae distally. Fore femur ventrally with yellow and blackish setae and setulae. Mid and hind femora, ventrally with thin long yellowish setulae. Tarsi with short, mostly blackish setulae, with shorter orange-brown setulae ventrally. Hind metatarsus anteroventrally with row of predominantly thick black setulae, distinguished from rest of setulae and more conspicuous in male. Wing (Fig. 34): Pattern: Mostly rather pale brown, reticulate, with large hyaline spots along and near costa and mostly smaller spots arranged in 2 longitudinal rows in each of cells r2+3, br and r4+5. Combined area of hyaline spots subequal to combined area of pigmented pattern. Conspicuous, dark, transverse bands or large spots present as follows: 1 band extending from costa at cell co, over crossvein BM-Cu, ending in anterobasal corner of cell cu1; 2 spots extending from costa in proximal 0.5 of pterostigma, ending in posterior margin of cell r1 proximal 0.5; 1 band extending from distal edge of pterostigma, aligned with crossvein R-M, ending medially at cell dm, anterior to vein Cu1; 1 spot extending from posterior part of cell dm, in level between crossveins R-M and DM-Cu, ending near medial of posterior margin of cell cu1; 1 narrow band extending (more or less intermittently) from costa to posterodistal edge of cell cu1, aligned with crossvein DM-Cu; 3 somewhat oblique bands beyond level of crossvein DM-Cu, converging medially towards cell r4+5: Proximal two forming "X" shape; apical band C-shaped. Costal cell hyaline with 4 dark spots, of which two middle most conspicuous. Venation: Veins generally yellowish to dark brown. Axillary sclerites and stem vein pale to dark brown. Costa mostly yellow. Base of costa, from proximal edge to costagial break, blackish. Vein M ends slightly posterior to wing tip. Crossvein DM-Cu slightly sinuous. Dorsal calypter slightly wider and shorter than ventral calypter. Halter with orange base, yellowish stem and pale to dark brown, sometimes apically beige, knob. Ratios: Wing: 2.68–2.91. Abdomen: Structure: Male T5 almost twice as long as T4, T4 about as long as T3. Female T4 about as long as T3 and slightly longer than T5. Color and vestiture: Microtrichia scattered more or less uniformly over tergites, mostly present medially over T2-T5. Male terminalia (Figs. 45, 56, 67): Epandrium rounded laterally, broadened dorsoventrally; ventral concavity inconspicuous, wide and superficial; subepandrial sclerite in lateral view enclosed by epandrium, indistinctly bent but with sharp corner projected dorsally; lateral surstylus broad, posteriorly rounded and apically narrowed; medial surstylus short and ambiguous, although slightly visible in ventral view. Terminal filaments equally long, slightly longer than preglans and glans combined and conspicuously shorter than distiphallus. Female terminalia (Fig. 81): 3 spermathecae present; spermatheca short ovoid and smooth. Ratios: Male: T3/T4: 0.86–1.00; T4/T5: 0.66–0.77; T3+4/T5: 1.13–1.45; epandrium: 0.81–0.97; preglans: 1.25–1.50; glans: 1.45–2.00; preglans/glans: 0.50–0.57; acrophallus/preglans+glans: 1.00–1.41; distiphallus/ preglans+glans: 11.88–13.93. Female: T3/T4: 0.93–1.10; T4/T5: 1.12–1.28; T3+4/T5: 1.93–2.17; aculeus: 6.17– 6.50; cercal unit: 2.33–2.40; cercal unit/aculeus: 0.16–0.18. Measurements (mm). Body length: 6.70–8.50; wing length: 5.60–7.10. Material examined. Holotype: Ƌ, ISRAEL, Nahal 'Iyyon, Ha'Tanur Waterfall, 430m, 33°16.1'N 35°34.5'E, 15.iii.2011, A. Freidberg. Paratypes: same collection data as holotype (23Ƌ, 26♀); additional paratypes (all from Israel; for explanation of the designation of paratypes, see Comments): large specimens: Har Dov, 8.iv.1983, AF (1♀); Nahal ' Iyyon, haTanur (' HaTanur', or ' Tanur'): 22.ii.2002, AF (1Ƌ, 1♀), 15.iii.1975, AF (27Ƌ, 13♀), 15.iii.1975, F. Kaplan (6Ƌ, 5♀), 430m, 33°16.1'N 35°34.5'E, 15.iii.2011, L. Bodner (24Ƌ, 26♀), AF (14Ƌ, 11♀), E. Morgulis (5♀), 21.iii.1974, D. Furth (2♀); Panyas: 33°14.9'N 35°41.7'E, 15.iii.2011, L. Bodner (13Ƌ, 12♀), AF (32Ƌ, 23♀), E. Morgulis (2Ƌ, 1♀), 16.iv.1992, AF (1♀), 205m, 33°15'N 35°42'E, 28.iv.2011, D. Furth (1♀); Sasa, 18.iv.1981, AF (8Ƌ, 5♀); HaGoshrim, 28–29.ii.1977, AF (5Ƌ), 29.ii.1977, F. Kaplan (1♀); Hurfeish, 7.iv.1987, Y. Zvik, on Phlomis viscosa, (1Ƌ, 2♀); Qiryat Shemona, 16.iv.1992, AF (5Ƌ, 2♀); Dan, Bet Ussishkin: 33°14'N 35°39'E, 15.iii.2011, AF (19Ƌ, 24♀), 33°14'N 35°39'E, 15.iii.2011, E. Morgulis (1♀); Avivim: 18.iv.1981, AF (4Ƌ, 2♀), 13.iii.1984, I Yarom (1♀); Emek HaHula, 26.iii.1992, D. Raucher (1Ƌ, 1♀); Nahal Keziv, Monfort (' Monfort', or ' Nahal Keziv'): 400m, 33°02'N 35°14'E, 24.ii.2011, L. Bodner (18Ƌ, 15♀), AF (5Ƌ), 33°02.6'N 35°13.3'E, 4.iii.2010, I. Katz (10Ƌ, 5♀), 33°02.635'N 35°13.271'E, 4.iii.2010, AF (3Ƌ, 4♀), 33°02.7'N 35°13.6'E, 5.iii.2008, AF (8Ƌ, 4♀), L. Friedman (4Ƌ, 4♀), 9.iii.1982, M. Kaplan (1Ƌ), 10.iii.1981, AF (1Ƌ), F. Kaplan (1♀), 33°02'N 35°14'E, 14.iii.2011, AF (1Ƌ, 1♀), E. Morgulis (1♀); Ma'alot-Tarshiha, 474m, 33°00'N 35°15'E, 26.iv.2005, M. Meir (2♀); Ma'alot, 3.v.2000, J. Halperin, on Rhamnus, (1Ƌ); Yir'on, 663m, 33°04.3'N 35°27'E, 14.iii.2011, L. Bodner (1Ƌ, 1♀); Yir'on, 17.iv.2000, AF (7Ƌ); Meron (' Har / Mt. Meron', or ' Meiron'): 31.iii.1958, J. Kugler (1Ƌ, 1♀), 9.iv.1977, AF (1♀), 1000m, 32°59.7'N 35°24.7'E, 14.iv.2011, AF (1Ƌ, 1♀), 1000m, 32°59.7'N 35°24.7'E, 14.iv.2011, L. Friedman (1Ƌ), 1100m, 32°59.7'N 35°24.7'E, 14.iv.2011, L. Bodner (1Ƌ, 2♀), 1000m, 16.iv.1992, AF (1♀), 1120m, 32°59.8'N 35°24.7'E, 19.iv.2010, E. Morgulis (1Ƌ), 1120m, 32°59.8'N 35°24.7'E, 19.iv.2010, AF (1♀), 23.iv.1973, M. Kaplan (1Ƌ), [Meron Reserve], Gat 'En Zeved, 32°59'N 35°25'E, 24.iv.2002, AF (1Ƌ), [Har Meron Reserve], 'En haZaqen, 32°58'N 35°25'E, 24.iv.2002, AF (1Ƌ), 1100m, 24.iv.2003, AF (3♀), [Har Meron Reserve], Camping under Kefar, 600m, 32°58'N 35°26'E, 25.iv.2002, AF (1Ƌ), 30.iv.1981, F. Kaplan (1Ƌ, 1♀), 3.v.1973, AF (1♀), 5.v.1975, AF (1♀), 5.v.1987, F. Kaplan (2Ƌ, 1♀), 6.v.1986, G. Eldar (1Ƌ, 2♀), 14.v.1974, AF (10Ƌ, 2♀), 14.v.1974, F. Nachbar (1Ƌ, 2♀), 17.v.1976, D. Simon (1Ƌ, 1♀), [Jarmaq], 20.v.1964, J. Kugler (2Ƌ), 20.v.1972, AF (2Ƌ, 1♀), 800m, 22.v.1998, AF (1Ƌ), 1120m, 33°00'N 35°25'E, 26.v.2009, AF (1Ƌ); Yaqqir [Yakir], 4.iv.1981, AF (1Ƌ); Zuri'el, 9.iv.1977, AF (1♀); Qiryat Anavim, 25.iv.1955, J. W. Wahrman (2Ƌ, 1♀). Small specimens: Nahal 'Iyyon, Ha'Tanur [Tanur], 17.iii.1973, D. Furth (2♀); N[ahal] Hazbani, 13.iii. [19]84, I. Yarom (1Ƌ, 1♀); HaGoshrim, 4.iv.1993 (2Ƌ, 5♀); Dafna, 18.iii.1973, M. Kaplan (3Ƌ, 4♀); Dan, 22.ii.1979, D. Furth (1Ƌ); Tel Dan, 13.iv. [19]83, I. Yarom (1♀); N[ahal] Sa’ar, 25.ii.1983, Z. Eythan (1Ƌ); Panyas [Banyas], 8.iii.1984, I. Nussbaum (6Ƌ, 1♀); 1 km East of Shomera, 31.iii. [19]68, Pener et al. (1♀); N[ahal] Bezet, 21.iv. [19]83, I. Yarom (1Ƌ); Horshat [Chorshat] Tal, 11.iv.1976, D. Simon (4Ƌ, 1♀); Nahal Keziv, Monfort ('Monfort', or 'Nahal Keziv'): 2.iii.1987, I. Yarom (1Ƌ), AF (1Ƌ), 4.iii.1976, AF (1Ƌ, 4♀), 4.iii.1993, AF (2Ƌ, 2♀), 17.iii.1983, AF (1♀), 18.iii.1973, AF (1♀); Gonen, 15.iii.1975, AF (3Ƌ); Ga`ton, 6.iii.2000, H. Ackerman (2Ƌ); Netu`a, 21.iv. [19]83, I. Yarom (2♀); Avivim, 18.iv.1981 (1♀,); Hurfeish, 7.iv.1987, I. Nussbaum (1♀); Rihaniya: 15.v.1996 (1♀), 22.v.1998 (1Ƌ); Park haYarden: 21.iii.2000, O. Manheim (1♀), H. Ackerman (1♀), 4.iv. [19]83, I. Yarom (6Ƌ, 1♀), 2.iv.1998, AF (1Ƌ); Biq'at Bet Zayda [Bteicha]: 14.iii.1975, AF (2Ƌ, 1♀), 24.iii.1973, M. Kaplan (1Ƌ); Ginnosar, 28.ii.1977, AF (1Ƌ); Kefar Nahum, 17.iii.1981, AF (1Ƌ); Yehudiya Forest Nature Reserve [Qasabiya], 20.iii.1984 (2Ƌ, 1♀), I. Nussbaum, 5.iv.1983 (1♀); Haifa [Haifa]: 4.ii.1969, AF (1Ƌ), 7.ii.1969, AF (1Ƌ, 1♀), 15.ii.1970, AF (1Ƌ, 3♀), 26.ii.1977, AF (1♀), 28.ii.1970, AF (1Ƌ, 1♀), 4.iii.1971, AF (3Ƌ), 7.iii.1969, AF (1♀), [Haifa, Palestine], 11.iii.1944, Bytinski-Salz (1Ƌ); Biq'at Bet Netofa [Palestina, Galilee, Bet Natufa], 1.iii. [19]55, O. Theodor (3Ƌ, 1♀); Nesher: 28.ii. [19]69, AF (1♀), 4.iii.1975, F. Kaplan (1Ƌ, 2♀); Oranim, 27.iii. [19]63,? (1Ƌ, 1♀); Qiryat Tiv`on [Tivon]: 4.iii.1975, F. Kaplan (22Ƌ, 3♀), 7.iii.1981, AF (2Ƌ), 2.iv.1975, F. Kaplan (1♀); Allonim, 3.ii.1942, Bytinski-Salz (1Ƌ, 1♀); Mishmar haEmeq, 4.iv. [19]53, M. Costa (2Ƌ, 2♀); Gesher, 16.iii.1973, D. Furth (1Ƌ); Karmel [Carmel]: 31.i.1970, AF (1Ƌ), 6.ii.1971 (1♀), 14.ii.1976, AF (2Ƌ), 22.ii.1981, AF (1Ƌ, 1♀), 4.iii.1975, F. Kaplan (1♀), 6.iii.1971, AF (25Ƌ, 10♀), 12.iii. [19]86, C. O`Toole (2Ƌ), 21.iii.1973, AF (2Ƌ), 8.iv. [19]67, D. Gerling (1♀); Yagur (or ' Jagur'): 21.ii.1976, AF (2Ƌ), [Jadjur, Haifa, Palestine], 5.iii. [19]34, O. Th[eodor] (1♀), 22.vi.1956, J. Kugler (1Ƌ); Nahal Oren: 32°43’N 35°01'E, 22.ii.2008, AF (5Ƌ), 4.iii.1975, M. Kaplan (2Ƌ, 2♀), 18.iii.1973, D. Furth (1♀), 18.iii.1981, (1♀); Isafiye, 18.iii.1973, D. Furth (1♀); N[ahal] Yoqne’am, 12.iii.1974, D. Furth (1Ƌ); Even Yizhaq (Gal'ed), 22.ii.2002, AF (1Ƌ); Allone`Abba, 14.iv.1999, AF (2Ƌ, 2♀); IAFAT, 2.iii.1968, J. Kugler (1♀); Bet Lehem haGlilit, 10.iii.1997, L. Friedman (3Ƌ, 1♀); Ramat Dawid, 14.iii. [19]81, T. Furman (1Ƌ, 1♀), 23.iii. [19]53, I. Goldman (1Ƌ); Megiddo, 20.ii. [19]84, S. Eliav (2Ƌ); Har Shana, Carmel, 24.iii.1989, J. Kugler (2Ƌ, 1♀); haZorea', 8.iii.1979, D. Furth (1♀); N[ahal] Tut, 2.iii.1979, D. Furth (1Ƌ); Keren haCarmel [Muhraqa], 18.iii.1987, I. Nussbaum (1♀);`En Hod, 27.iii.1975, AF (2Ƌ, 1♀);`En [Ein] Hod, 30.iii.1987, I. Yarom (1Ƌ, 4♀); Zikhron Ya`aqov: 9.ii.1998, I. Yarom (1Ƌ), 24.ii.1968, J. Kugler (3Ƌ, 2♀), 1.iv.1998, AF (1Ƌ, 2♀); Nahal 'Iron [Wadi/W. Ara]: 2.iii.1978, A. Freudberg (2Ƌ, 1♀), 19.iii.1974, D. Furth (1♀); Nahal Tavor, north facing slope, 26.iii.2001, L. Friedman (2Ƌ); Nahal Tavor, 26.iii.2001, L. Friedman (1♀);`En Pelugot, 16.iv.1973, D. Furth (1Ƌ); Hadera, 11.iii.1940, Com. Inst. Ent. (1♀); Upper Nahal Tirza [Upper W. Faria], 19.ii.1974, D. Furth (1Ƌ); Nahal Tirza [W. Faria]: 1.iii.1973, AF (3Ƌ, 1♀), 6.iii.1974, D. Furth (1Ƌ), 11.iii.1973, D. Furth (1Ƌ); Sabastiya, 24.iii.1973, D. Furth (3Ƌ, 3♀); 'Ammiqam, 8.ii.1997, R. Hoffman (1♀); Shekhem: [Shchem], 1.iii.1973, M. Kaplan (2Ƌ), [Nablus], 11.iii.1973, D. Furth (1Ƌ, 1♀); Kokhav Ya'ir: Ya'ar Sappir, 160m, 32°13.9’N 34°59.5'E, 16.ii.2010, I. Katz (10Ƌ, 4♀), L. Friedman (4Ƌ, 7♀), AF (2♀), 32°14’N 34°59.5'E, 21.iii.2011, L. Bodner (16Ƌ, 12♀); Shoham: 2.iii.2004, I. Zonstein (3Ƌ), 17.iii.2010, L. Friedman (1Ƌ), AF (1♀), 32°00’N 34°57'E, 18.iii.2011, L. Bodner (7Ƌ, 4♀), 32°00’N 34°57'E, 23.iii.2007, L. Younger & E. Kaufmann (1Ƌ, 1♀); Bet Nehemya, 6.iv.1984, I. Nussbaum (2Ƌ, 2♀); Park Canada: 13.ii. [20]10, W. Kuslitzky (1♀), 29.iii.1992, AF (1Ƌ, 1♀); Latrun, 10.iii.2004, AF (1♀); Bet Shemesh: 10.iii.2004, AF (3♀), 21.iii.2000, E. Fonio (1Ƌ), 24.iii. [19]83, AF (1Ƌ, 1♀), 30.iii.2004, I. Zonstein (1Ƌ, 1♀); Devira, 31°34’N 34°51'E, 12.iv.2009, R. Haneman (1♀); Bet Lehem, 10.ii.1995, I. Nussbaum (1Ƌ); Talimon, 3.iii.1982, I. Nussbaum (1Ƌ). The holotype (Fig. 97) is double-mounted (minutien pin and polyporus block), in excellent condition, and deposited in SMNHTAU. Most paratypes are deposited at SMNHTAU; some paratypes will be deposited at the museums listed in the Material and Methods section. Distribution. Israel. Distribution in Israel. Mt. Hermon, Golan Heights, Upper and Lower Galilee, Carmel Ridge, Jordan Valley, Yizre'el Valley, Samaria, Foothills of Judea and Judean Hills. Adult activity in Israel. February to April. Etymology. This species is named after Mrs Geula Freidberg, the late mother of the junior co-author. It is a noun in apposition. Comments. This is the largest species of Platystomatidae found in Israel. The numerous specimens studied by us can be fairly equally divided into two size groups based on body length: 6.70–8.50 mm or 4.20–6.20 mm. Our prior assumption that these two size groups may represent two new species is not supported by any other morphological evidence. We have therefore chosen to base our description of the new species on the large specimens only and designated these as the type series. The small specimens are also recorded in the material examined section following the large specimens, although they are not designated as paratypes. Adults were frequently collected on Silybum marianum (Asteraceae), Mandragora autumnalis (Solanaceae), Scandix pecten-veneris, and Smyrnium olusatrum (both Apiaceae).Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on pages 218-221, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
Platystoma trigonum Bodner & Freidberg, 2016, n. sp.
No full textPlatystoma trigonum n. sp. (Figs. 17, 28, 39, 50, 61, 72, 86, 99) Diagnosis. This species is similar to P. subfasciatum described from Bulgaria (Loew, 1862), differing from it by the following two characters, already used in Hennig's key (1945): (1) ventral half of face completely shiny, lacking the median line of microtrichia (which is present in P. subfasciatum); (2) T5 of the male 3–4 times as long as T3 or T4, as opposed to 1.5 times as long in the latter species. The new species differs from Loew's and Hennig's descriptions of P. subfasciatum mentioned above by three additional characters: (1) the wings illustrated by Hennig (1945) and by Hendel (1913) do not match our specimens, as the pattern of P. subfasciatum is composed of denser and more uniformly scattered hyaline spots, as well as two discrete dark bands on posterior half of wing; (2) ocellar triangle as long as, or slightly longer than frons; and (3) ocellar triangle with short black setulae. This latter character is unique among the local Platystoma species and was not mentioned in the earlier descriptions of P. subfasciatum. We believe that studying specimens of P. subfasciatum from the type locality or around it would unlikely reverse our decision and result in a synonym. Description. Head (Figs. 17, 28): Color and vestiture: Head mostly pale to dark brown, sometimes with yellowish-brown areas. Frons microtrichose mostly at orbits and around bases of orbital setae. Ocellar triangle delimited by dense line of yellowish microtrichia. Lunule dark brown to black dorsally, yellow to pale brown ventrally, with yellowish line delimiting scapes and inconspicuous microtrichia medially. Antenna: scape pale to dark brown, pedicel yellowish to orange-brown and 1st flagellomere dark brown, all irregularly microtrichose. Face: dorsal 0.5 brown to blackish with dense gray microtrichia; ventral 0.5 shiny dark brown to black, without microtrichia. Clypeus with 1 (pair) oval dorsomedial spot, 1 (pair) smaller dorsolateral spot, and single band of microtrichia along ventral margin. Palpus brown to black, with yellowish to pale brown base. Chaetotaxy: 2 (pairs) orbital setae present. Postocellar seta absent. Ocellar triangle setulose. Lunule setulose. Postgena with long yellowish setulae. Occiput with similar setulae as postgena, except shorter. Ratios: Head: 1.40–1.56; frons anterior: 1.67–2.00; frons posterior: 1.42–1.83; ocellar triangle/frons: 0.46–0.79; eye: 1.80–2.14; 1st flagellomere: 1.50– 2.14; gena/eye: 0.30–0.38; parafacial/gena: 0.25–0.36; posterior orbital/anterior orbital: 1.33–2.00; medial vertical/ lateral vertical: 0.90–1.10; posterior orbital/medial vertical: 0.54–0.74; ocellar/medial vertical: 0.32–0.42; ocellar/ ocellar triangle: 0.47–0.55. Thorax: Color and vestiture: Ground color dark brown to black, covered with yellowish-gray microtrichia. Scutum with 3 or 5 microtrichose stripes. Postpronotal lobe ground color black. Scutellum microtrichose marginally and in median stripe. Subscutellum metallic black with fine microtrichia. Anepisternum and katepisternum with reticulate pattern of microtrichia, excluding shiny black ventral part of katepisternum. Chaetotaxy: Presutural supra-alar seta either conspicuously short or absent. Postsutural supra-alar seta present. Postpronotal lobe with long yellowish-white to brown setulae. Anepisternum mostly short setulose, with longer setulae posteriorly. Katepisternum mostly moderately long, yellowish-white to pale brown setulose, with longer setulae ventrally. Ratios: Scutum: 0.84–0.98; presutural supra-alar/intra-alar: 0.20–0.36; postsutural supra-alar/ intra-alar: 0.53–0.83; postalar/intra-alar: 0.86–1.22. Legs: Color and vestiture: Leg type dark. Tarsi with yellowish areas as in genus. Fore coxa with fine microtrichia, mid and hind coxae inconspicuously microtrichose. Chaetotaxy: Coxae anterodorsally and ventrally setulose: fore coax mostly with white setulae, mid and hind coxae with yellowish white to dark brown setulae. Trochanters ventrally with dark brown setulae. Femora black setulose: fore femur with posteriorly long and anteriorly short setulae; mid and hind femora mostly short setulose, with long and thick setulae posterodistally. Hind metatarsus and 2nd tarsomere with yellowish setulae ventrally. Wing (Fig. 39): Pattern: Generally non-uniformly reticulate, with small to medium-size sub-hyaline and hyaline spots; combined area of hyaline and subhyaline spots less than combined area of pigmented pattern; larger such spots mostly around periphery of wing; smaller subhyaline spots mostly arranged in two more or less parallel and regular marginal longitudinal rows along cells r2+3, br, r4+5 and dm. Subhyaline spots otherwise irregular or arranged in 1 or several rows. Pattern with more or less discrete transverse dark bands extending from costa to hind margin of wing: 1 band from distal edge of pterostigma, more or less aligned with or slightly beyond level of crossvein R-M and nearly extending to crossvein DM-Cu, ending in the middle of posterior margin of cell cu1; 3 somewhat oblique bands beyond level of crossvein DM-Cu, converging medially towards cell r4+5: Proximal two forming "X" shape; apical band C-shaped. Costal cell subhyaline to hyaline with 4–6 partially merged dark spots. Venation: Veins generally pale brown; axillary sclerites, stem vein and base of costa from proximal edge to costagial break dark brown; rest of costa yellowish to pale brown. Vein M ends slightly posterior to wing tip. Dorsal calypter narrower and shorter than ventral calypter, with conspicuously longer fringes. Halter base and stem yellow, knob dark brown to blackish. Chaetotaxy: Base of costa near costagial break with ventral seta. Ratios: Wing: 2.79–3.00. Abdomen: Structure: Male T4 about as long as T3 and about 0.33 times as long as T5. Female T4 slightly longer than T3 or T5. Color and vestiture: Male abdomen densely and uniformly microtrichose. Female abdomen sparsely and irregularly microtrichose, mostly medially over tergites. Male terminalia (Figs. 50, 61, 72): Epandrium laterally rounded, dorsally planate; ventral concavity deep; subepandrial sclerite conspicuously broadened anteriorly and extended beyond anterior margin of epandrium; lateral surstylus posteriorly and laterally rounded, strongly curved mesally; medial surstylus ambiguous. Terminal filaments unequally long, each conspicuously longer than distiphallus. Female terminalia (Fig. 86): 3 spermathecae present; spermatheca shortovoid with thin spiny protuberances. Ratios: Male: T3/T4: 0.85–1.11; T4/T5: 0.26–0.37; T3+4/T5: 0.51–0.71; epandrium: 0.45–0.62; preglans: 1.17–1.40; glans: 1.64–2.43; preglans/glans: 0.31–0.41; acrophallus/ preglans+glans: 6.13–9.10 / 12.25–14.38 (short/long filament); distiphallus/preglans+glans: 4.50–6.43. Female: T3/T4: 1.00–1.32; T4/T5: 1.06–1.19; T3+4/T5: 2.29–2.53; aculeus: 5.20–9.63; cercal unit: 2.90–3.00; cercal unit/ aculeus: 0.17–0.22. Measurements (mm). Body length: 3.70–6.20; wing length: 3.60–5.20. Material examined. Holotype: Ƌ, ISRAEL, Har Hermon, 1700m, 8.vi.1975, A. Freidberg. Paratypes: same collection data as holotype (16Ƌ, 6♀); additional paratypes (all from Israel, Har Hermon; but sometimes labeled " Mt. Hermon " or " Hermon ", arranged by increasing altitude): 1200m: 24.v. [19]83, I. Nussbaum (1Ƌ, 2♀); 9.vi. [19]83, I. Yarom (3Ƌ); 1300m, 22.v.1973, AF (1Ƌ, 1♀); 1400m: 23.v.1978, D. Simon (1Ƌ); 23.v.2012, AF (1Ƌ); 21.vi.1971, J. Kugler (1Ƌ); 1400–1600m, 24.v. [19]83, AF (1Ƌ); 1400–1650m, 18.v.1976, AF (2Ƌ); 1500m: 6.v. [19]75, D. Simon (1♀); 18.v.1976, D. Simon (1Ƌ, 1♀); 23.v.1978, AF (4Ƌ, 3♀); 20.vi. [19]77, M. Broza (1Ƌ, 1♀); 1515m, 33°17.7’N 35°45.6'E, 17.v.2011, L. Bodner (1♀); 1500–1700m, 24.v.2012, L. Bodner (2♀); 1600m: 6.v.1975, F. Kaplan (1♀); 33°18.3’N 35°46.2'E, 17.v.2011, L. Bodner (1♀); 23.v.1998, AF (1♀); 33°18.1’N 35°46.2'E, 24.v.2010, E. Morgulis (1♀); 6.vi.1973, M. Kaplan (1♀); 9.vi.1975, M. Kaplan (2Ƌ, 1♀); 9.vi. [19]83, I. Yarom (1Ƌ); 12.vi.2003, AF (1Ƌ); 22.vi.1971, J. Kugler (1Ƌ, 1♀); 27.vi.1977, J. Kugler (1♀); 6.vii.1987, I. Nussbaum (1Ƌ); 7.vii.1987, AF (1Ƌ), Yoram Zvik (1Ƌ); 8.vii.1987, I. Nussbaum (1Ƌ); 9.vii.1987, I. Nussbaum (1Ƌ, 1♀), F. Kaplan (2Ƌ); 1650m: 6.v.1975, AF (1Ƌ, 2♀), J. Kugler (1Ƌ); 22.v.1973, AF (3Ƌ, 1♀); 9.vi.1975, AF (2Ƌ); 16.vi.1969, J. Kugler (1Ƌ); 22.vi.1973, AF (3Ƌ); 1700m: 9.vi. [19]83, AF (1Ƌ, 2♀); 9.vi. [19]83, I. Nussbaum (3Ƌ); 10.vi. [19]83, I. Nussbaum (5Ƌ, 7♀); 16.vi.1977, AF (11Ƌ, 8♀); 16.vi. [19]77, Z. Feler (2Ƌ); 21.vi.1982, AF (1♀); 22.vi.1973, AF (5Ƌ, 2♀); 28.vi.1971, J. Kugler (2Ƌ, 3♀); 1750m: 8.vi.1975, M. Kaplan (1Ƌ); 22.vi.1973, M. Kaplan (1Ƌ); 1800m: 11.vi.1976, AF (1Ƌ); 11.vi.2003, L. Friedman (1Ƌ); 12.vi.1996, AF (1♀); 1900m, 1.vii.1986, AF (1Ƌ); 2000m: 20.v.1986, AF (2Ƌ, 1♀); 30.v.1979, M. Kaplan (1Ƌ); 2.vi.1985, I. Nussbaum (1♀); 9.vi. [19]83, AF (2♀); 12.vi.1996, AF (1Ƌ, 1♀); 21.vi.1982, AF (1Ƌ); 28.vi.1971, J. Kugler (1♀); 13.viii.1973, AF (1♀); (No altitude) 19.v. [19]81, I. Yarom (1Ƌ). The holotype (Fig. 99) is double-mounted (minutien pin and polyporus block), in excellent condition, and deposited at the SMNHTAU. Most paratypes are deposited at the SMNHTAU; some paratypes will be deposited at the museums listed in the Material and Methods section. Distribution. Israel. Distribution in Israel. Mt. Hermon. Adult activity in Israel. May to July. Etymology. We name this species " trigonum " (Latin: "triangular harp"), due to its unique ocellar triangle. Comments. As we were unable to obtain comparative material of Platystoma subfasciatum (which appears to be the most similar species), the validity of this new species is based only on comparisons with published descriptions.Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on pages 229-231, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
Platystoma elizabethae Bodner & Freidberg, 2016, n. sp.
No full textPlatystoma elizabethae n. sp. (Figs. 11, 22, 33, 44, 55, 66, 80, 96) Diagnosis. This species bears some resemblance to Platystoma malatiense Hennig, except for the following characters: ventral half of face shiny brown, without microtrichia; clypeus microtrichose only marginally; T5 of the male 7–14 times as long as T4 (significantly longer than in all other congeners). Description. Head (Figs. 11, 22): Color and vestiture: Head mostly pale to dark brown, with orange-brown areas. Frons sparsely and irregularly microtrichose, with denser microtrichia at orbits and around bases of orbital and vertical setae. Ocellar triangle delimited by thin yellowish line of microtrichia. Lunule orange to dark brown, sometimes dorsomedially microtrichose, with yellowish lines delimiting scapes. Antenna brown to blackish, densely microtrichose (less pronounced in certain angles); pedicel sometimes yellowish or pale brown. Face mostly brown to blackish, with yellowish to orange-brown areas ventrally; dorsal 0.5 mostly grayish microtrichose with small, shiny brown to dark brown area medially; ventral 0.5, dorsally shiny brown to blackish, ventrally matte, and mostly yellowish-orange except blackish areas laterally. Clypeus with conspicuous microtrichose spots (sometimes forming narrow marginal bands): 1 (pair) dorsomedially, 1 (pair) dorsolaterally, 1 (pair) ventrolaterally, and 1 broad spot ventromedially. Palpus brown to dark brown with yellowish base. Chaetotaxy: 1–2 (pairs) orbital setae present; anterior seta, if present, short and setula-like. Postocellar seta absent. Lunule densely setulose. Gena with 2–3 setae and long blackish setulae. Postgena with thin, long, yellowish setulae; occiput with similar but shorter setulae. Ratios: Head: 1.37–1.50; frons anterior: 1.29–1.50; frons posterior: 1.10–1.32; ocellar triangle/frons: 0.27– 0.38; eye: 2.00–2.36; 1st flagellomere: 1.50–2.00; gena/eye: 0.26–0.35; parafacial/gena: 0.17–0.25; posterior orbital/anterior orbital: 1.79–2.75; medial vertical/lateral vertical: 1.06–1.27; posterior orbital/medial vertical: 0.50–0.67; ocellar/medial vertical: 0.41–0.57; ocellar/ocellar triangle: 1.07–1.42. Thorax: Color and vestiture: Ground color mostly dark brown to black, sometimes with reddish-brown areas, with yellowish-gray microtrichia. Scutum with 5 microtrichose stripes. Postpronotal lobe ground color reddishbrown to dark brown. Posterior corner of postpronotal lobe and median articulation of anepisternum and katepisternum, shiny orange to brown. Scutellum mostly microtrichose, except black spots around bases of setae. Subscutellum dark brown to blackish, grayish microtrichose. Anepisternum with homogenous reticulate pattern of microtrichia. Katepisternum mostly shiny black, except irregularly microtrichose dorsally. Chaetotaxy: Postpronotal lobe with 1 seta and with long, yellowish-white to pale brown setulae. Scapullar setae, if present, thin and setula-like. Presutural supra-alar seta and postsutural supra-alar seta present (presutural seta sometimes setulalike). Anepisternum with short yellowish to pale brown setulae anterodorsally, and long yellowish to pale brown setulae posteriorly and ventrally. Katepisternum with long yellowish to orange setulae. Ratios: Scutum: 0.85–0.94; presutural supra-alar/intra-alar: 0.19–0.34; postsutural supra-alar/intra-alar: 0.40–0.68; postalar/intra-alar: 0.92– 1.12. Legs: Color and vestiture: Leg type dark. Tarsi with yellowish areas as in genus. Coxae dorsally with fine microtrichia. Femora inconspicuously microtrichose. Chaetotaxy: Coxae anterodorsally setulose. Trochanters ventrally with brown setulae. Femora pale brown to blackish setulose: fore femur anterodorsally with short setulae and posteroventrally with long setulae; mid femur and hind femur mostly short setulose, except long setulae posterodistally and ventrally. Hind metatarsus and 2nd tarsomere with yellowish setulae ventrally. Wing (Fig. 33): Pattern: Generally non-uniformly reticulate, with small to medium-size sub-hyaline and hyaline spots; combined area of hyaline and subhyaline spots less than combined area of pigmented pattern; larger such spots mostly around periphery of wing; smaller subhyaline spots mostly arranged in two parallel marginal rows along cells r2+3, br, r4+5 and dm; subhyaline spots otherwise irregular or arranged in 1 or several rows. Discrete transverse dark bands present as follows: 1 band from middle of costal cell to anterobasal margin of cell cu1; 1 band from distal tip of costal cell to end of vein A1+Cu2, fading posteriorly; 1 band from costa between subcostal vein and R1 to middle of vein Cu1; 1 broad band aligned with crossvein R-M, extending from costa to posterior margin of cell cu1; 3 somewhat oblique bands beyond level of crossvein DM-Cu, converging medially towards cell r4+5: Proximal two forming "X" shape; apical band C-shaped. Costal cell subhyaline to hyaline with 3–4 large dark spots. Venation: Veins and sclerites mostly dark brown, except yellowish to pale brown costa, humeral, subcostal and stem vein. Vein M ends slightly posterior to wing tip. Crossvein DM-Cu slightly sinuous, without stump vein. Calypters subequal; dorsal calypter with longer fringes than ventral calypter. Halter base and knob brown, stem yellowish-brown. Chaetotaxy: Costa ventrally with seta and long whitish setulae, and with conspicuously long setulae basally. Ratios: Wing: 2.56–2.85. Abdomen: Structure: Male T4 either shorter, or about as long as T3, less than 0.20 times as long as T5. Female T4 longer than T3 and about as long as T5. Color and vestiture: T1+2 densely and irregularly microtrichose; T3 mostly metallic black with sparse microtrichia; T 4 in male medially microtrichose, in female uniformly and sparsely microtrichose; T 5 in male microtrichose mostly or only in distal 0.5, in female uniformly and more densely microtrichose. Male terminalia (Figs. 44, 55, 66): Epandrium in lateral view dorsoventrally narrowed, posterior margin concave; ventral concavity wide deep; subepandrial sclerite conspicuously elongated and almost completely exposed (in lateral view), anteriorly extended beyond margin of epandrium and posteroventrally with conspicuously long narrow bridges; lateral surstylus short, ventrally broad, anteriorly concaved (in lateral view), posteriorly and apically rounded; medial surstylus broad and ambiguous; prensisetae strongly curved mesally. Terminal filaments equally long, and conspicuously longer than distiphallus. Female terminalia (Fig. 80): 2 spermathecae present; spermatheca short ovoid, with small protuberances. Ratios: Male: T3/T4: 0.50–1.17; T4/T5: 0.06–0.14; T3+4/T5: 0.10–0.30; epandrium: 0.47–0.52; preglans: 1.25–1.40; glans: 1.89–2.31; preglans/glans: 0.33–0.50; acrophallus/preglans+glans: 12.56–14.40; distiphallus/preglans+glans: 5.35–6.47. Female: T3/T4: 1.25–1.36; T4/T5: 1.00–1.14; T3+4/T5: 2.25–2.67; aculeus: 5.00–7.20; cercal unit: 2.00–3.50; cercal unit/aculeus: 0.17–0.19. Measurements (mm). Body length: 3.60–4.70; wing length: 3.80–4.60. Material examined. Holotype: Ƌ, ISRAEL, Nahal Keziv, Monfort, 10.iii.1981, A. Freidberg. Paratypes: same collection data as holotype (18Ƌ, 23♀); additional paratypes (all from Israel; for explanation of the variant locality names on the labels, see Comments): Nahal Keziv, Montfort, 400m, 33°02'N 35°14'E, 24.ii.2011, L. Bodner (6Ƌ, 2♀), AF (3Ƌ, 3♀); 24.ii–14.iii.2011, L. Bodner (7Ƌ, 8♀); 2.iii.1987, AF (3Ƌ, 8♀), A. Shlagman (2Ƌ, 7♀), I. Yarom (3Ƌ, 1♀); 4.iii.1976, AF (21Ƌ, 12♀), M. Kaplan (3Ƌ, 4♀); 4.iii.1993, AF (1Ƌ, 1♀); 400m, 33°02'N 35°14'E, 4.iii.2010, AF (1♀); 33°02.635'N 35°14.271'E, 4.iii.2010, AF (1Ƌ); 5.iii.1978, M. Kaplan (1♀); 5.iii.1978, AF (1Ƌ); 33°02.7'N 35°14.6'E, 5.iii.2008, AF (2♀); 9.iii.1982, AF (5Ƌ, 2♀); 10.iii.1981, AF (19Ƌ, 23♀), F. Kaplan (4Ƌ, 4♀), T. Furman (3Ƌ, 2♀); 14.iii.1985, AF (5♀); 33°02'N 35°14'E, 14.iii.2011, AF (11Ƌ, 13♀), D. Furth (2Ƌ, 2♀); 400m, 33°02'N 35°14'E, 14.iii.2011, E. Morgulis (1Ƌ, 14♀), L. Bodner (4Ƌ, 9♀); 17.iii.1983, AF (11Ƌ, 4♀); 18.iii.1973, D. Furth (1♀); 4.iv.1976, AF (1♀); 10.iv.1993, AF (7Ƌ, 2♀). The holotype (Fig. 96) is double-mounted (minutien pin and polyporus block), in excellent condition, and deposited at the SMNHTAU. Most paratypes are deposited at the SMNHTAU; some paratypes will be deposited at the museums listed in the Material and Methods section. Distribution. Israel. Distribution in Israel. Upper Galilee: Lower Nahal Keziv (Montfort). Adult activity in Israel. February to April. Etymology. This species is named after our colleague, Elizabeth Morgulis, who has contributed to the senior author's research in many ways. It is an adjective. Comments. This species has been collected to date only at the Monfort crusader castle and within a radius of about one kilometer from it, constituting one of the most beautiful sites in Israel. The locality names originally used and cited on the labels, i.e., “Montfort”, “Monfort”, “Nahal Keziv”, “Nahal Keziv, Monfort” and “N. Keziv” (Nahal, in Hebrew, meaning river or stream) more or less refer to the same site.Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on pages 216-218, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel
No full textBodner, L., Freidberg, A. (2016): Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel. Zootaxa 4171 (2): 201-245, DOI: http://doi.org/10.11646/zootaxa.4171.2.
FIGURES 74–76. Ovipositor and cercal unit. 74. Platystoma torridum, ovipositor. 75. Platystoma elizabethae, cercal unit. 76 in Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel
No full textFIGURES 74–76. Ovipositor and cercal unit. 74. Platystoma torridum, ovipositor. 75. Platystoma elizabethae, cercal unit. 76. Rivellia israelica, cercal unit.Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on page 240, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
Rivellia israelica Bodner & Freidberg, 2016, n. sp.
No full textRivellia israelica n. sp. (Figs. 18, 29, 40, 51, 62, 73, 76, 87, 100) Diagnosis. This species is diagnosed both in the key to the Platystomatidae of Israel and in the generic diagnosis above. It was not found to be conspecific with any of the Palaearctic species keyed by Hennig (1945). However, we did not continue comparing our specimens with specimens or descriptions of the remaining 130 species around the world. See also the explanatory comment under the genus title above. Description. Head (Figs. 18, 29): Structure: First flagellomere truncate and more or less angular apically, often distinctly pointed dorsoapically; arista with short dense rays, nearly plumose, total width including rays about 0.25–0.33 times height of 1st flagellomere. Color and vestiture: Head yellowish-brown. Conspicuous narrow stripes of silvery-white microtrichia present only around eye, especially noticeable along posteroventral margin of eye. Ocellar triangle blackish, subshiny, not microtrichose. Lunule yellow. Antenna mostly yellowish but tip blackish, and dark coloration often extended more proximally, with fine microtrichia. Face yellowish, dorsal half with dense grayish microtrichia. Clypeus shiny brownish, without microtrichia. Palpus yellow. Proboscis shiny brownish (sometimes discolored). Chaetotaxy: Frons densely setulose, bare around ocellar triangle. 2 (pairs) orbital setae and 0–1 (pair) of postocellar seta present. Lunule setulose. Usually 1 conspicuous genal seta present, among several sparse, thin, moderately long, straight brownish setulae. Occiput sparsely setulose. Ratios: Head: 1.20–1.37; frons anterior: 1.03–1.24; frons posterior: 0.91–1.09; ocellar triangle/frons: 0.28–0.44; eye: 1.65–1.89; 1st flagellomere: 2.84–3.97; gena/eye: 0.15–0.19; parafacial/gena: 0.15–0.32; posterior orbital/anterior orbital: 0.49–0.81; medial vertical/lateral vertical: 0.89–1.00; posterior orbital/medial vertical: 0.40–0.72; ocellar/medial vertical: 0.48–0.59; ocellar/ocellar triangle: 0.80–1.26. Thorax: Color and vestiture: Ground color usually black and yellow, greatly varied among the specimens; in dark specimens most of mesonotum from neck to scutellum, but not postpronotum and area around it, notopleuron and posterolateral margin of scutum to level of dorsocentral seta blackish; subscutellum, mediotergite and laterotergite also blackish; katepisternum black except dorsally, strongly shiny, almost without microtrichia; laterotergite and small area between base of forecoxa and anterior spiracle with conspicuous small whitish or silvery spots of microtrichia. In pale specimens blackish area either restricted to median region of scutum or attains U-shaped form, with blackish extension from arm of U laterally along transverse suture, and pleura entirely yellow, although mediotergite partly blackish; in dark specimens mesonotum almost entirely blackish, except more or less narrow margin, and mediotergite and ventral pleura blackish, mesonotum covered with thin microtrichia. Mediotergite mostly shiny, laterally microtrichose. Chaetotaxy: Postpronotal lobe with 1 seta and brownish or yellowish setulae. Scutum setae (pairs): 1–2 scapular, 0–2, usually 0 or enlarged setulae, presutural supra-alar, 1 postsutural supra-alar, 1 postalar, 1 intra-alar, 1 dorsocentral and 0–1 acrostichal. 2 notopleural setae present. Scutum uniformly short setulose; setulae usually about 0.2 times as long as adjacent major setae. Scutellum with 2 (pairs) scutellar setae, basal about 0.5 times as long as apical, scutellum also with 10–15 setulae. Pleura: Anepisternum posteriorly uniformly short setulose with 1–2 posterodorsal setae; setulae thin, erect, yellowish. Anepimeron with 1 seta and several fine setulae. Katepisternum with 1–2 setae ventrally, and with sparse, erect, long, yellowish setulae. Ratios: Scutum: 1.10–1.52; postsutural supra-alar/intra-alar: 0.52–0.96; postalar/intra-alar: 0.99–1.25. Legs: Color and vestiture: Leg entirely yellow except hind tibia almost entirely brownish, except base and tip, non-microtrichose. Chaetotaxy: only fore femur posteroventrally with distinct row of setae, with 3–4 distal setae larger and darker. Otherwise, without overt features. Wing (Fig. 40): Pattern: In most species, including the new species from Israel, wing banded, with 4–5 bands variable in dimension, orientation and degree of coalescence. Basal band usually marginal and longitudinal; distal two bands usually joined together along costa; pattern sometimes reduced to single band and small spot, or wing almost entirely hyaline. Venation: Veins generally yellowish to pale brown. Vein M ends slightly posterior to wing tip. Crossvein DM-Cu slightly curved toward base of wing. Calypteres white. Halter microtrichose, with yellowish base and stem and blackish knob. Chaetotaxy: R1 dorsally with dense row of evenly-spaced short black setulae, ventrally bare. R4+5 dorsally with row of short sparse blackish setulae, ventrally with 2 setulae aligned with pterostigma. Tegula usually with short blackish setulae and 2–4 black setae. Ratio: Wing: 2.29–2.98. Abdomen: Structure: Male and female with 5 preabdominal tergites. Female’s T6 absent. Color and vestiture: Ground color black or blackish, subshiny, shagreened. Microtrichia absent completely. Male terminalia (Figs. 51, 62, 73): Epandrium in lateral view ventrally convex and not saddle-shaped, squarish anteriorly, prolonged posteriorly; prolongation considerably longer than squarish part. Lateral surstylus indistinguishable from epandrium (without obvious articulation), apparently fused with it and strongly sclerotized. Subepandrial sclerite not bent ventrally in middle. Medial surstylus hidden by lateral surstylus. Two heavily sclerotized claw-like prensisetae present. Phallus with bulbous preglans and glans, and with 2 equally-long terminal filaments; terminal filaments with swollen base. Female terminalia: Ovipositor mostly as in Platystoma (Fig. 74); cercal unit (Fig. 76) broadened basally; subapical ventral setulae not adjoined. 3 spermathecae present (Fig. 87), 2 connected by common duct broadened into single atrium; 3rd single spermatheca also opens into similar atrium; spermatheca shallow funnel-like, moderately sclerotized, with smooth surface, without ring-shaped base, apical bulge and internal structure. Ratios: Male: T3/T4: 0.83–0.93; T4/T5: 1.14–1.37; T3+4/T5: 2.11–2.54; epandrium: 0.98–1.01; preglans: 0.75–0.88; glans: 1.89–1.97; preglans/glans: 0.28–0.33; acrophallus/preglans+glans: 1.57–1.6; distiphallus/preglans+glans: 4. Female: T3/T4: 0.63–0.84; T4/T5: 1.44–2.81; T3+4/T5: 2.52–4.75; aculeus: 4.39– 5.26; cercal unit: 3.370–3.373; cercal unit/aculeus: 0.25–0.26. Measurements (mm). Body length: 3.45–3.94; wing length: 2.70–3.36. Material examined. Holotype: Ƌ, ISRAEL, Park HaYarden, 20.vi.1982, A. Freidberg, malaise trap. Paratypes: same collection data as holotype (12Ƌ, 92♀); additional paratypes (all from Israel): Panyas (usually as ' Baniass'): 24.iv.1982, AF (3Ƌ, 2♀), F. Kaplan (2♀), I. Yarom (1♀), 5.v.1977, D. Simon (1Ƌ, 2♀), 9.vi.1976, AF (1Ƌ, 1♀), D. Simon (1♀), 10.vi.1976, D. Simon (2♀), 13.vi.1982, AF (2Ƌ, 1♀), 28.vi.1977, AF (1♀), 10.vii.1975, M. Kaplan (2♀), 25.vii.1985, AF (1♀), 30.viii.1981, AF (1♀); Tel Dan: 5.v.1977, AF (6Ƌ, 8♀), D. Simon (1♀), 6.v.1979, D. Furth (1Ƌ, 1♀), 16.v.1971, AF (2Ƌ, 3♀), 27.v.2003, AF (1Ƌ), 18.x.1969, AF (2Ƌ); HaGosherim, 21.iv.1969, AF (1♀); ' Enot ' Enan, 1.v.2006, L. Friedman (2Ƌ); Rosh Pinna, 19.vii.1977, AF (1Ƌ); Park HaYarden: -200[m], 32°54.6'N 35°37.5'E, 21.iii.2010, E. Morgulis (1Ƌ), 32°54.5'N 35°37.5'E, 14.iv.2011, AF (2♀), 7.v.2009, AF (1Ƌ, 1♀), 17.v.1997, AF (1♀), W. Kuslitzky (1♀), - 190m, 32°54.7'N 35°37.6'E, 22.v.2011, E. Morgulis (1♀), 14.vi.1982, AF (3Ƌ, 2♀), 16.vi.1982, AF (6Ƌ, 2♀), 17.vi.1982, AF (1Ƌ), 24.vi.1982, AF (1♀), 25.vii. [19]83, Y. Zvik (1♀), 5.viii.1986, AF (1Ƌ), [P. HaYarden], 7.viii. [19]83, Nussbaum (3Ƌ, 1♀); Biq'at Bet Zayda [Bteicha]: 14.iii.1975, AF (1♀), 11.viii.1970, AF (1Ƌ, 1♀); ' En Gev, 7.v.1997, AF (1♀); Rosh ha'Ayin, 8.vi.1996 (2Ƌ, 1♀). The holotype (Fig. 100) is double-mounted (minutien pin and polyporus block), in excellent condition, and deposited at the SMNHTAU. Most paratypes are deposited at the SMNHTAU; some paratypes will be deposited at the museums listed in the Material and Methods section. Distribution. Israel. Distribution in Israel. Mostly along the Rift Valley—Upper Galilee, Golan Heights and Jordan Valley. A few specimens were collected in the Central Coastal Plain (Rosh ha'Ayin). Etymology. The specific epithet ‘ israelica’ denotes the country from which this species is known and in which it is geographically isolated from other members of the genus. Adult activity in Israel. March to October. FIGURES 63–73. Phallus. 63. Platystoma arcuatum. 64. P. dalia. 65. P. dimidiatum. 66. P. elizabethae. 67. P. geula. 68. P. malatiense. 69. P. nitidiventre. 70. P. pubescens. 71. P. torridum. 72. P. trigonum. 73. Rivellia israelica. FIGURES 77–87. Spermathecae. 77. Platystoma arcuatum. 78. P. dalia. 79. P. dimidiatum. 80. P. elizabethae. 81. P. geula. 82. P. malatiense. 83. P. nitidiventre. 84. P. pubescens. 85. P. torridum. 86. P. trigonum. 87. Rivellia israelica.Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on pages 232-241, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
FIGURES 19–29. Head, lateral view. 19. Platystoma arcuatum. 20. P. dalia. 21. P. dimidiatum. 22. P. elizabethae. 23. P. geula. 24. P. malatiense. 25. P. nitidiventre. 26. P. pubescens. 27. P. torridum. 28. P. trigonum. 29 in Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel
No full textFIGURES 19–29. Head, lateral view. 19. Platystoma arcuatum. 20. P. dalia. 21. P. dimidiatum. 22. P. elizabethae. 23. P. geula. 24. P. malatiense. 25. P. nitidiventre. 26. P. pubescens. 27. P. torridum. 28. P. trigonum. 29. Rivellia israelica.Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on page 235, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
Platystoma arcuatum Loew 1856
No full textPlatystoma arcuatum Loew, 1856 (Figs. 8, 19, 30, 41, 52, 63, 77) Platystoma arcuatum Loew, 1856: 50. Selected references: Hendel, 1913: 97 (monograph of Platystoma); Hennig, 1945: 27 (palaearctic Platystomatidae). Diagnosis. This species differs from all other congeners by the combination of small size (usually 2.5–3.50 mm), the presence of a nearly complete hyaline transverse band composed of hyaline spots over crossvein DM-Cu, and the T5 of the male, which is 3–4 times as long as either T4 or T3. Redescription. Head (Figs. 8, 19): Color and vestiture: Head mostly brown to black, with pale brown areas. Frons mostly translucent microtrichose, with dense silvery-white microtrichia at orbits and around bases of orbital and vertical setae. Ocellar triangle delimited by thin grayish line of microtrichia. Lunule brown to black, with grayish median line of microtrichia. Antenna brown to blackish, densely microtrichose (less pronounced in certain angles); pedicel sometimes yellowish anterodorsally. Face dark brown to black; dorsal 0.5 grayish microtrichose; ventral 0.5 shiny, except thin median line of grayish microtrichia. Clypeus shiny black with either microtrichose spots on corners, or narrow microtrichose band ventrally. Palpus dark brown to black, with pale brown base. Chaetotaxy: 2 (pairs) orbital setae and 1 (pair) postocellar seta present. Lunule setulose. Gena with 1–3 setae, and with long dark brown setulae. Postgena with long pale brown setulae. Occiput with short pale brown setulae. Ratios: Head: 1.46–1.56; frons anterior: 1.17–1.44; frons posterior: 1.20–1.39; ocellar triangle/frons: 0.27–0.36; eye: 1.90–2.38; 1st flagellomere: 1.67–2.50; gena/eye 0.24–0.29; parafacial/gena: 0.20–0.28; posterior orbital/ anterior orbital: 1.35–2.67; medial vertical/ lateral vertical: 1.00–1.31; posterior orbital/medial vertical: 0.41–0.76; ocellar/medial vertical: 0.32–0.64; ocellar/ocellar triangle: 1.08–1.92; postocellar/medial vertical: 0.30–0.48. Thorax: Color and vestiture: Ground color mostly black, inconspicuously grayish microtrichose. Scutum with 3 or 5 weak microtrichose stripes. Postpronotal lobe ground color dark brown to black. Scutellum usually matte black, sometimes inconspicuously microtrichose marginally. Subscutellum dark brown, without microtrichia. Anepisternum with sparsely reticulate pattern of microtrichia. Katepisternum mostly shiny dark brown to black, with inconspicuously microtrichose margins. Chaetotaxy: Postpronotal lobe with 1 seta and moderately long brown setulae. Presutural supra-alar seta and postsutural supra-alar seta present. Anepisternum mostly short dark brown setulose, with paler, slightly longer setulae posteriorly. Katepisternum mostly moderately long dark brown setulose, with longer setulae ventrally. Ratios: Scutum: 0.85–0.94; presutural supra-alar/intra-alar: 0.31–0.52; postsutural supra-alar/intra-alar: 0.44–0.73; postalar/intra-alar: 1.00–1.33. Legs: Color and vestiture: Leg type dark. Tarsi with following yellowish areas: fore metatarsus proximally, mid and hind metatarsi ventrally, and 2nd tarsomere of hind tarsus ventrally. Coxae inconspicuously microtrichose. Trochanters, femora and tibiae shiny, without microtrichia. Chaetotaxy: Coxae and trochanters black setulose. Femora dark brown setulose, with short setulae dorsally and long setulae ventrally. Fore and hind tibiae with 0–2 spines. All tarsi with dark brown setulae dorsally and yellowish to pale brown setulae ventrally. Wing (Fig. 30): Pattern: Generally dark brown, with small hyaline spots scattered more or less uniformly on proximal 0.5–0.6 of wing, posterior to vein R2+3, and with small to large hyaline spots scattered somewhat irregularly on distal 0.5 of wing; more or less conspicuous, fragmented, sometimes complete, transverse band present, consisting of large hyaline spots extending from costa to hind margin of wing, aligned with crossvein DM- Cu; hyaline spots otherwise scattered mostly on apex and around periphery of wing. Combined area of hyaline spots less than combined area of pigmented pattern. Pattern with discrete transverse dark bands extending from costa to hind margin of wing: 1 band extending from pterostigma, over crossvein R-M, near crossvein DM-Cu, ending in posterodistal margin of cell cu1; 2 somewhat oblique bands beyond level of crossvein DM-Cu: Proximal band wider than medial band, starts at apex of cell r1; 1 spot or narrow band over wing apex. Costal cell mostly dark brown, with thin subhyaline spots. Venation: Veins and sclerites dark brown. Vein M ends slightly anterior to wing tip. Crossvein DM-Cu without stump vein. Calypteres pale brown with dark brown margins and yellowish-white fringes; dorsal calypter slightly narrower and shorter than ventral calypter, with conspicuously longer fringes. Halter base brown, stem beige to pale brown, knob dark brown. Chaetotaxy: Base of costa near costagial break, with long ventral seta. Ratios: Wing: 2.41–2.67. Abdomen: Structure: Male T4 about as long as T3 and less than 0.33 times as long as T5. Female T4 slightly shorter than T3 and about as long as T5. Color and vestiture: Abdomen almost completely shiny black, with slight microtrichia (sometimes more in female). Male terminalia (Figs. 41, 52, 63): Epandrium laterally broad; ventral concavity with posterior depression; subepandrial sclerite anteriorly broadened and extended beyond anterior margin of epandrium; lateral surstylus in ventral view finger-like, posteriorly rounded and curved mesally, with pointed apex; medial surstylus ambiguous. Terminal filaments equally long and slightly longer than distiphallus. Female terminalia (Fig. 77): 3 spermathecae present; spermatheca short ovoid, with large protuberances. Ratios: Male: T3/T4: 0.71–1.27; T4/T5: 0.26–0.28; T3+4/T5: 0.48–0.57; epandrium: 0.49–0.55; preglans: 1.17–1.50; glans: 1.64–1.89; preglans/glans: 0.39–0.55; acrophallus/preglans+glans: 8.24–10.17; distiphallus/preglans+glans: 6.47–7.29. Female: T3/T4: 0.89–1.00; T4/T5: 0.93–1.12; T3+4/T5: 1.80–2.12; aculeus: 4.17–5.50; cercal unit: 2.50–3.33; cercal unit/aculeus: 0.20–0.23. Measurements (mm). Body length: 2.40–5.30; wing length: 2.70–3.40. Material examined. ISRAEL: Nahal ' Iyyon, haTanur (' HaTanur', or ' Tanur'): 17.iii.1973, D. Furth (1Ƌ, 1♀), 21.iii.1974, D. Furth (1Ƌ), 13.iv. [19]83, AF (1♀), 430m, 33°16.1'N 35°34.5'E, 15.iii.2011, L. Bodner (19Ƌ, 12♀), AF (5Ƌ, 5♀); Newe Ativ, 26.iv.1964, D. Furth (1♀); Senir [Hazbani], 24.iv.1982, F. Kaplan (1Ƌ); Mezudat Nimrod: [Cala’at Nemrod], 24.iv.1982, AF (1Ƌ), 33°15.2’N 35°43,2'E, 760m, 17.v.2011, L. Bodner (7Ƌ, 1♀), AF (2Ƌ); Dan, Bet Ussishkin, 33°14’N 35°39'E, 15.iii.2011, AF (7Ƌ, 1♀); Tel Dan, 13.iv. [19]83, AF (3Ƌ, 6♀), I. Yarom (1Ƌ, 3♀); Panyas (or ' Banias'): 33°14.9'N 35°41.7'E, 15.iii.2011, L. Bodner (1♀), M. Morgulis (1♀), 205m, 33°15'N 35°42'E, 28.iv.2011, D. Furth (2Ƌ), 4.v.1977, AF (1♀); 24.iv.1982, AF (11Ƌ, 6♀), I. Yarom (1Ƌ), F. Kaplan (1Ƌ, 1♀), 12.iv. [19]83, AF (1Ƌ). I. Yarom (2Ƌ, 1♀), 16.iv.1992, AF (7Ƌ, 4♀), 8.iii.1984, I. Nussbaum (1♀); Nahal Bezet: [Karkara], 31.iii. [19]68, Pener et al. (5Ƌ, 1♀), 6.iv. [19]67, J. Margalit (3Ƌ, 2♀), 6.iv. [19]67, Pener et al. (3Ƌ), 9.iv. [19]68, Pener et al. (1♀), 12.iv. [19]67, Pener et al. (1Ƌ), [N. Bezet], 21.iv. [19]83, I. Yarom (1Ƌ), Y. Zvik (1♀); Hanita, 27.iii.1976, D. Gerling (1Ƌ, 1♀); Nahal Keziv, Monfort (' Monfort', or ' Nahal Keziv'): 14.iii.1985, AF (1Ƌ), 10.iv.1993, AF (1Ƌ, 2♀), 33°02.635'N 35°13.271'E, 4.iii.2010, AF (1♀), 33°02.7'N 35°13.6'E, 5.iii.2008, L. Friedman (2Ƌ), AF (1Ƌ, 1♀), 4.iii.2010, L. Friedman (1Ƌ, 1♀), 33°02.6'N 35°13.3'E, 4.iii.2010, I. Katz (1Ƌ); Avivim, 18.iv.1981, AF (1♀); Sasa, 18.iv.1981, AF (21Ƌ, 7♀); Meron (' Har / Mt. Meron', or ' Meiron'): [Jermak], 16.iv. [19]63, J. Margalit (1Ƌ), 20.iv.1972, AF (4Ƌ, 2♀), 3.v.1973, AF (1♀), 32°59'N 36°24'E, 30.iv.2007, T. Levanony, Malaise trap (2♀), 1100m, 32°59.7'N 36°24.7'E, 14.iv.2011, L. Bodner (1Ƌ); Nahal 'Ammud: 17.iii.1973, D. Furth (2Ƌ, 1♀), 4.iv. [19]83, I. Yarom, (1♀), 31.iii. [19]82, I. Nussbaum (1Ƌ); Zefat, 10.iv.1976, D. Simon (♀); 'Ein el Asad, 18.iv.1984, I. Nussbaum (1Ƌ, 1♀); Golan, Upper Nahal Zavitan, 7.v.1987, A. Shlagman (1Ƌ); Golan, Devora Fall, 23.iv.1973, AF (1♀); Golan, Qazrin, 4.v.1999, AF (1Ƌ); Park haYarden: - 210m, 32°54.5’N 35°37.5'E, 14.iv.2011, AF (1Ƌ, 1♀), 2.iv.1998, AF (2♀), 4.iv. [19]83, AF (1♀), 7.v.1997, AF (1Ƌ); Teverya [Tiberias, Jordan Valley, Palestine], 15.v.1949, D. Theodor (1Ƌ, 2♀); Nahal Tavor, 25–26.iii.2001, L. Friedman (4Ƌ, 2♀); Qiryat Tiv'on, 22.iii.1975, B. Shadmot (1♀), 2.iv.1975, F. Kaplan (7Ƌ, 2♀), 2.iv.1975, M. Kaplan (1♀); Har haKarmel, Horeshat ha'Arba'im, 450m, 32°45.5’N 35°01.6'E, 2.v.2011, AF (1♀); Nahal Oren: 9.iii.2012, AF (7Ƌ, 1♀), 16.iv.2003, AF (1Ƌ), 14.iv.2006, AF (2♀); HaZorea', 8.iii.1979, D. Furth (1Ƌ); Gal'ed, 19.iii. [19]83, AF (1Ƌ); Ma'agan Mikha'el, 32°33’N 34°55.6'E, 26.iii.2009, W. Kuslitzky (1Ƌ); Zikhron Ya'aqov: [Z.Y.], 12.iii.1940, Com. Inst. Ent. Coll. No. 12720 (1Ƌ, 1♀), 1.v.1998, AF (5Ƌ, 1♀); Zomet Elyaqim, 1.iv.2012, L. Bodner (31Ƌ, 6♀); Sebastiya, 24.iii.1973, D. Furth (1Ƌ); Kokhav Ya'ir, 32°19’N 34°59'E, 21.iii.2011, AF (2Ƌ, 1♀), L. Bodner (1Ƌ); Qedumim, 24.iii. 2 0 0 5, L. Friedman (1♀); Yarhiv, 24.iii. [19]83, AF (1Ƌ); Tel Aviv, Abu Kabir, 28.iv.1981, AF (1Ƌ); Bet Dagan, 32°00’N 34°50'E, 18.iv.2008, W. Kuslitzky (2♀); Rantis Junction, near Nahshonim, 13.iv.1999, AF (1♀); Shoham, 18.iii.2011, L. Bodner (1Ƌ); Park Canada, 29.iii.1992, AF (1Ƌ); Bet Shemesh, 30.iii.2004, L. Friedman (2Ƌ, 2♀), I. Zonstein (4♀); Zomet haEla, 4.iv.1999, AF (1Ƌ). Type locality. Beirut. Distribution. Turkey, Lebanon, Syria and Israel. Distribution in Israel. Galilee, Golan Heights, Jordan Valley, Carmel Ridge, Samaria, Central Coastal Plain, Foothills of Judea and Judean Hills. Adult activity in Israel. March to May. Comments. This is the smallest species of Platystomatidae found in Israel, and is probably the smallest Platystoma species anywhere. Adults were frequently collected on Scandix pecten-veneris and Smyrnium olusatrum (Apiaceae) and Euphorbia sp. (Euphorbiaceae).Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on pages 211-213, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
Platystoma malatiense Hennig 1945
No full textPlatystoma malatiense Hennig, 1945 (Figs. 13, 24, 35, 46, 57, 68, 82) Platystoma malatiense Hennig, 1945: 26 (palaearctic Platystomatidae). Diagnosis. This small-to-medium size species bears some resemblance to P. arcuatum and P. trigonum n. sp. in its overall appearance, but is easily distinguished from these and other congeners by the presence of dense white microtrichia on almost the entire facial area and on the clypeus. Redescription. Head (Figs. 13, 24): Color and vestiture: Head pale to dark brown with orange to reddishbrown areas. Frons densely microtrichose at orbits, on and around ocellar triangle and around bases of orbital and vertical setae. Ocellar triangle densely microtrichose. Lunule brown to blackish-brown with grayish microtrichia. Antenna brown to blackish-brown with dense grayish microtrichia; pedicel pale brown, sometimes yellowish dorsally; 1st flagellomere dark brown. Face dark brown to black medially, yellowish to pale brown lateroventrally, mostly densely microtrichose, except black shiny areas dorsolaterally, and yellowish-brown matte areas ventrolaterally. Clypeus densely microtrichose. Chaetotaxy: 2 (pairs) orbital setae present. Postocellar seta absent. Lunule setulose dorsomedially. Postgena with long yellowish setulae. Occiput with similar setulae as postgena, except shorter. Ratios: Head: 1.33–1.46; frons anterior: 1.35–1.56; frons posterior: 1.16–1.33; ocellar triangle/ frons: 0.24–0.33; eye: 1.92–2.17; 1st flagellomere: 1.38–2.00; gena/eye: 0.30–0.35; parafacial/gena: 0.25–0.31; posterior orbital/anterior orbital: 1.73–3.27; medial vertical/lateral vertical: 0.90–1.25; posterior orbital/medial vertical: 0.73–1.06; ocellar/medial vertical: 0.30–0.78; ocellar/ocellar triangle: 0.78–2.15. Thorax: Color and vestiture: Ground color black, with yellowish-gray microtrichia. Scutum with 5 microtrichose stripes. Postpronotal lobe ground color usually black, sometimes reddish. Scutellum microtrichose marginally, sometimes also with medial stripe of microtrichia. Subscutellum metallic dark brown to black, with fine microtrichia. Proepisternum and proepimeron densely white microtrichose. Chaetotaxy: Presutural supra-alar seta and postsutural supra-alar seta present. Postpronotal lobe with thin long yellowish-white setulae. Anepisternum with long, posteriorly brown and ventrally yellowish-white setulae, otherwise, with short yellowishwhite to brown setulae. Katepisternum with yellowish-white setulae. Ratios: Scutum: 0.88–0.97; presutural supraalar/intra-alar: 0.31–0.45; postsutural supra-alar/intra-alar: 0.53–0.80; postalar/intra-alar: 1.04–1.26. Legs: Color and vestiture: Leg type dark. Tarsi with yellowish areas as in genus. Coxae, trochanters and most parts of forelegs with fine microtrichia. Mid and hind femora, either inconspicuously microtrichose or bare. Chaetotaxy: Coxae anteriorly and distally with mixed white and brown setulae. Trochanters with yellowish setulae. Fore femur black setulose, with long setulae ventrally and shorter setulae dorsally. Mid and hind femora with short black setulae dorsally, and with long paler setulae ventrally. Hind femur with long and thick black setulae dorsally and distally. Tibiae short setulose, with blackish setulae dorsally and yellowish setulae ventrally. Hind metatarsus and 2nd tarsomere with yellowish setulae ventrally. Wing (Fig. 35): Pattern: Reticulate, with small to medium-size hyaline spots, scattered more or less uniformly on proximal 0.5 of wing and irregularly on distal 0.5 of wing. Combined area of hyaline spots less than combined area of pigmented pattern. Pattern with discrete transverse dark bands extending from costa to hind margin of wing: 2 bands converging posterior to vein R2+3 and extending as one band over crossvein R-M near to crossvein DM-Cu, ending near middle of posterior margin of cell cu1; 3 somewhat oblique bands beyond level of crossvein DM-Cu, converging medially towards cell r4+5. Costal cell subhyaline to hyaline with 3–5, sometimes partially merged large dark spots. Venation: Veins mostly pale to dark brown; axillary sclerites, stem vein and base of costa from proximal edge to costagial break, dark brown; rest of costa yellowish to pale brown. Vein M ends slightly anterior to wing tip. Crossvein DM-Cu without stump vein. Dorsal calypter narrower and shorter than ventral calypter, with conspicuously longer fringes. Halter base and stem yellowish to pale brown, knob dark brown. Ratios: Wing: 2.68– 3.03. Abdomen: Structure: Male T4 about as long as T3 and about half as long as T5. Female T4 slightly shorter than T3 and slightly longer than T5. Color and vestiture: Male abdomen more or less uniformly microtrichose. Female abdomen irregularly microtrichose, with microtrichia scattered sparsely over T3 and T4, and more densely over T5. Male terminalia (Figs. 46, 57, 68): Epandrium broad; ventral concavity wide and superficial; subepandrial sclerite laterally enclosed by epandrium; lateral surstylus in ventral view finger-like, rounded laterally and slightly curved mesally, with pointed apex; medial surstylus rectangle, rather distinguishable (mostly visible in ventral view); cerci conspicuously elongated. Terminal filaments equally long, and distinctly shorter than distiphallus. Female terminalia (Fig. 82): 3 spermathecae present; spermatheca short ovoid and smooth. Ratios: Male: T3/T4: 1.00–1.08; T4/T5: 0.57–0.71; T3+4/T5: 1.17–1.43; epandrium: 0.63–0.77; preglans: 1.22–1.50; glans: 1.82–2.25; preglans/glans: 0.38–0.50; acrophallus/preglans+glans: 3.12–4.33; distiphallus/preglans+glans: 9.88–10.09. Female: T3/T4: 1.05–1.20; T4/T5: 1.08–1.18; T3+4/T5: 2.38–2.44; aculeus: 4.39–6.07; cercal unit: 2.71–3.20; cercal unit/aculeus: 0.18–0.22. Measurements (mm). Body length: 3.20–5.30; wing length: 2.80–3.90. Material examined. ISRAEL: Har Hermon (or ' Mt. Hermon'): 1300m, 23.v.1998, AF (1♀); 1400m, 24.v.2012, AF (3♀); 1400m, 15.vi.2011, AF (1♀); 1400–1600m, 24.v. [19]83, AF (1Ƌ, 1♀); 1500m, 17.v.2009, L. Friedman (3♀), AF (2♀); 1515m, 33°17.7’N 35°45.6'E, 17.v.2011, L. Bodner (25Ƌ, 22♀), AF (10Ƌ, 4♀); 1500– 1700m, 24.v.2012, L. Bodner (7Ƌ, 7♀); 1600m, 4.v.1999, AF (1Ƌ); 33°18.3’N 35°46.2'E, 17.v.2011, AF (1Ƌ, 7♀), L. Bodner (2Ƌ, 1♀); 1600m, 18.v.2009, AF (3Ƌ, 3♀), L. Friedman (2Ƌ, 4♀); 1700m, 17.v.2009, AF (6♀), L. Friedman (1Ƌ, 4♀), W. Kuslitzky (1♀); 1800m, 33°18.1’N 35°46.4'E, 7.v.2010, AF (1♀); Nahal ‘Iyyon, HaTanur Waterfall, 430m, 33°16.1’N 35°34.5'E, 15.iii.2011, AF (15Ƌ, 22♀), L. Bodner (15Ƌ, 18♀), E. Morgulis (1Ƌ, 2♀); Panyas [Banias], 12.iv. [19]83, AF (1Ƌ); Avivim, 18.iv.1981, AF (43Ƌ, 26♀); Yir'on, 17.iv.2000, AF (31Ƌ, 6♀); Sasa, 18.iv.1981, AF (1♀); Rihaniya [Rihanyia], 16.iv.1992, AF (1Ƌ, 1♀); Meron (' Har / Mt. Meron', or ' Meiron'): 1100m, 9.iv.1977, AF (3♀), 1100m, 32°59.7’N 35°24.7'E, 14.iv.2011, L. Bodner (14Ƌ, 15♀), 1000m, 32°59.7’N 35°24.7'E, 14.iv.2011, L. Friedman (5Ƌ, 5♀), AF (1Ƌ, 3♀), E. Morgulis (1Ƌ, 1♀), 1100m, 17.iv.2000, AF (4Ƌ, 3♀), H. Ackerman (1Ƌ), 17.iv.2012, L. Friedman (1Ƌ, 4♀), 20.iv.1972, AF (5Ƌ, 2♀), 1100m, 27.iv. [19]84, I. Yarom (1Ƌ, 1♀), 900m, 27.iv. [19]84, AF (1Ƌ), 1100m, 30.iv.1981, F. Kaplan (1Ƌ); 3.v. [19]83, Y. Zvik, (1Ƌ), 14.v.1974, AF (1♀), 1100m, 28.v.1981, AF (1♀), 800m, 28.v.2003, L. Friedman (1Ƌ), [Har Meron Reserve], 'En Zeved, 32°59'N 35°26'E, 24.iv.2002, AF (1Ƌ, 3♀), L. Friedman (1Ƌ), Nahal 'Ofaim, 1100m, 20.v.2013, L. Friedman (1♀); ' En-el-Asad, 18.iv.1984, I. Nussbaum (2Ƌ, 1♀); Emeq [k] Peqi'in, 3.iv.1978, D. Furth (1♀); Har Addir, 980m, 20.v.2013, L. Friedman, on Q[uercus]. boissieri, Q. calliprinos, (1Ƌ); Dalton, 25.iv.1974, D. Furth (2Ƌ); Qiryat ' Anavim: 25.iv.1955, HEBR. UNIV., J.W. Wahrman (3Ƌ, 1♀), [Jerusalem, Palestine, Qiryat ' Anavim], 25.iv. [19]55, O. Theodor (1Ƌ, 6♀); Yerushalayim, Bet haKerem [Beth Hakerem, Jerusalem, Palestine], 25.iv. [19]55, O. Theodor (1Ƌ, 1♀). Type locality. Malatia (Turkey). Distribution. Turkey, Armenia, Iraq and Israel. Distribution in Israel. Upper Galilee, Mt. Hermon, Golan Heights and Judean Hills. Adult activity in Israel. April to May.Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on pages 221-222, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
Platystoma malatiense Hennig 1945
No full textPlatystoma malatiense Hennig, 1945 (Figs. 13, 24, 35, 46, 57, 68, 82) Platystoma malatiense Hennig, 1945: 26 (palaearctic Platystomatidae). Diagnosis. This small-to-medium size species bears some resemblance to P. arcuatum and P. trigonum n. sp. in its overall appearance, but is easily distinguished from these and other congeners by the presence of dense white microtrichia on almost the entire facial area and on the clypeus. Redescription. Head (Figs. 13, 24): Color and vestiture: Head pale to dark brown with orange to reddishbrown areas. Frons densely microtrichose at orbits, on and around ocellar triangle and around bases of orbital and vertical setae. Ocellar triangle densely microtrichose. Lunule brown to blackish-brown with grayish microtrichia. Antenna brown to blackish-brown with dense grayish microtrichia; pedicel pale brown, sometimes yellowish dorsally; 1st flagellomere dark brown. Face dark brown to black medially, yellowish to pale brown lateroventrally, mostly densely microtrichose, except black shiny areas dorsolaterally, and yellowish-brown matte areas ventrolaterally. Clypeus densely microtrichose. Chaetotaxy: 2 (pairs) orbital setae present. Postocellar seta absent. Lunule setulose dorsomedially. Postgena with long yellowish setulae. Occiput with similar setulae as postgena, except shorter. Ratios: Head: 1.33–1.46; frons anterior: 1.35–1.56; frons posterior: 1.16–1.33; ocellar triangle/ frons: 0.24–0.33; eye: 1.92–2.17; 1st flagellomere: 1.38–2.00; gena/eye: 0.30–0.35; parafacial/gena: 0.25–0.31; posterior orbital/anterior orbital: 1.73–3.27; medial vertical/lateral vertical: 0.90–1.25; posterior orbital/medial vertical: 0.73–1.06; ocellar/medial vertical: 0.30–0.78; ocellar/ocellar triangle: 0.78–2.15. Thorax: Color and vestiture: Ground color black, with yellowish-gray microtrichia. Scutum with 5 microtrichose stripes. Postpronotal lobe ground color usually black, sometimes reddish. Scutellum microtrichose marginally, sometimes also with medial stripe of microtrichia. Subscutellum metallic dark brown to black, with fine microtrichia. Proepisternum and proepimeron densely white microtrichose. Chaetotaxy: Presutural supra-alar seta and postsutural supra-alar seta present. Postpronotal lobe with thin long yellowish-white setulae. Anepisternum with long, posteriorly brown and ventrally yellowish-white setulae, otherwise, with short yellowishwhite to brown setulae. Katepisternum with yellowish-white setulae. Ratios: Scutum: 0.88–0.97; presutural supraalar/intra-alar: 0.31–0.45; postsutural supra-alar/intra-alar: 0.53–0.80; postalar/intra-alar: 1.04–1.26. Legs: Color and vestiture: Leg type dark. Tarsi with yellowish areas as in genus. Coxae, trochanters and most parts of forelegs with fine microtrichia. Mid and hind femora, either inconspicuously microtrichose or bare. Chaetotaxy: Coxae anteriorly and distally with mixed white and brown setulae. Trochanters with yellowish setulae. Fore femur black setulose, with long setulae ventrally and shorter setulae dorsally. Mid and hind femora with short black setulae dorsally, and with long paler setulae ventrally. Hind femur with long and thick black setulae dorsally and distally. Tibiae short setulose, with blackish setulae dorsally and yellowish setulae ventrally. Hind metatarsus and 2nd tarsomere with yellowish setulae ventrally. Wing (Fig. 35): Pattern: Reticulate, with small to medium-size hyaline spots, scattered more or less uniformly on proximal 0.5 of wing and irregularly on distal 0.5 of wing. Combined area of hyaline spots less than combined area of pigmented pattern. Pattern with discrete transverse dark bands extending from costa to hind margin of wing: 2 bands converging posterior to vein R2+3 and extending as one band over crossvein R-M near to crossvein DM-Cu, ending near middle of posterior margin of cell cu1; 3 somewhat oblique bands beyond level of crossvein DM-Cu, converging medially towards cell r4+5. Costal cell subhyaline to hyaline with 3–5, sometimes partially merged large dark spots. Venation: Veins mostly pale to dark brown; axillary sclerites, stem vein and base of costa from proximal edge to costagial break, dark brown; rest of costa yellowish to pale brown. Vein M ends slightly anterior to wing tip. Crossvein DM-Cu without stump vein. Dorsal calypter narrower and shorter than ventral calypter, with conspicuously longer fringes. Halter base and stem yellowish to pale brown, knob dark brown. Ratios: Wing: 2.68– 3.03. Abdomen: Structure: Male T4 about as long as T3 and about half as long as T5. Female T4 slightly shorter than T3 and slightly longer than T5. Color and vestiture: Male abdomen more or less uniformly microtrichose. Female abdomen irregularly microtrichose, with microtrichia scattered sparsely over T3 and T4, and more densely over T5. Male terminalia (Figs. 46, 57, 68): Epandrium broad; ventral concavity wide and superficial; subepandrial sclerite laterally enclosed by epandrium; lateral surstylus in ventral view finger-like, rounded laterally and slightly curved mesally, with pointed apex; medial surstylus rectangle, rather distinguishable (mostly visible in ventral view); cerci conspicuously elongated. Terminal filaments equally long, and distinctly shorter than distiphallus. Female terminalia (Fig. 82): 3 spermathecae present; spermatheca short ovoid and smooth. Ratios: Male: T3/T4: 1.00–1.08; T4/T5: 0.57–0.71; T3+4/T5: 1.17–1.43; epandrium: 0.63–0.77; preglans: 1.22–1.50; glans: 1.82–2.25; preglans/glans: 0.38–0.50; acrophallus/preglans+glans: 3.12–4.33; distiphallus/preglans+glans: 9.88–10.09. Female: T3/T4: 1.05–1.20; T4/T5: 1.08–1.18; T3+4/T5: 2.38–2.44; aculeus: 4.39–6.07; cercal unit: 2.71–3.20; cercal unit/aculeus: 0.18–0.22. Measurements (mm). Body length: 3.20–5.30; wing length: 2.80–3.90. Material examined. ISRAEL: Har Hermon (or ' Mt. Hermon'): 1300m, 23.v.1998, AF (1♀); 1400m, 24.v.2012, AF (3♀); 1400m, 15.vi.2011, AF (1♀); 1400–1600m, 24.v. [19]83, AF (1Ƌ, 1♀); 1500m, 17.v.2009, L. Friedman (3♀), AF (2♀); 1515m, 33°17.7’N 35°45.6'E, 17.v.2011, L. Bodner (25Ƌ, 22♀), AF (10Ƌ, 4♀); 1500– 1700m, 24.v.2012, L. Bodner (7Ƌ, 7♀); 1600m, 4.v.1999, AF (1Ƌ); 33°18.3’N 35°46.2'E, 17.v.2011, AF (1Ƌ, 7♀), L. Bodner (2Ƌ, 1♀); 1600m, 18.v.2009, AF (3Ƌ, 3♀), L. Friedman (2Ƌ, 4♀); 1700m, 17.v.2009, AF (6♀), L. Friedman (1Ƌ, 4♀), W. Kuslitzky (1♀); 1800m, 33°18.1’N 35°46.4'E, 7.v.2010, AF (1♀); Nahal ‘Iyyon, HaTanur Waterfall, 430m, 33°16.1’N 35°34.5'E, 15.iii.2011, AF (15Ƌ, 22♀), L. Bodner (15Ƌ, 18♀), E. Morgulis (1Ƌ, 2♀); Panyas [Banias], 12.iv. [19]83, AF (1Ƌ); Avivim, 18.iv.1981, AF (43Ƌ, 26♀); Yir'on, 17.iv.2000, AF (31Ƌ, 6♀); Sasa, 18.iv.1981, AF (1♀); Rihaniya [Rihanyia], 16.iv.1992, AF (1Ƌ, 1♀); Meron (' Har / Mt. Meron', or ' Meiron'): 1100m, 9.iv.1977, AF (3♀), 1100m, 32°59.7’N 35°24.7'E, 14.iv.2011, L. Bodner (14Ƌ, 15♀), 1000m, 32°59.7’N 35°24.7'E, 14.iv.2011, L. Friedman (5Ƌ, 5♀), AF (1Ƌ, 3♀), E. Morgulis (1Ƌ, 1♀), 1100m, 17.iv.2000, AF (4Ƌ, 3♀), H. Ackerman (1Ƌ), 17.iv.2012, L. Friedman (1Ƌ, 4♀), 20.iv.1972, AF (5Ƌ, 2♀), 1100m, 27.iv. [19]84, I. Yarom (1Ƌ, 1♀), 900m, 27.iv. [19]84, AF (1Ƌ), 1100m, 30.iv.1981, F. Kaplan (1Ƌ); 3.v. [19]83, Y. Zvik, (1Ƌ), 14.v.1974, AF (1♀), 1100m, 28.v.1981, AF (1♀), 800m, 28.v.2003, L. Friedman (1Ƌ), [Har Meron Reserve], 'En Zeved, 32°59'N 35°26'E, 24.iv.2002, AF (1Ƌ, 3♀), L. Friedman (1Ƌ), Nahal 'Ofaim, 1100m, 20.v.2013, L. Friedman (1♀); ' En-el-Asad, 18.iv.1984, I. Nussbaum (2Ƌ, 1♀); Emeq [k] Peqi'in, 3.iv.1978, D. Furth (1♀); Har Addir, 980m, 20.v.2013, L. Friedman, on Q[uercus]. boissieri, Q. calliprinos, (1Ƌ); Dalton, 25.iv.1974, D. Furth (2Ƌ); Qiryat ' Anavim: 25.iv.1955, HEBR. UNIV., J.W. Wahrman (3Ƌ, 1♀), [Jerusalem, Palestine, Qiryat ' Anavim], 25.iv. [19]55, O. Theodor (1Ƌ, 6♀); Yerushalayim, Bet haKerem [Beth Hakerem, Jerusalem, Palestine], 25.iv. [19]55, O. Theodor (1Ƌ, 1♀). Type locality. Malatia (Turkey). Distribution. Turkey, Armenia, Iraq and Israel. Distribution in Israel. Upper Galilee, Mt. Hermon, Golan Heights and Judean Hills. Adult activity in Israel. April to May.Published as part of Bodner, L. & Freidberg, A., 2016, Taxonomy and immature stages of the Platystomatidae (Diptera: Tephritoidea) of Israel, pp. 201-245 in Zootaxa 4171 (2) on pages 221-222, DOI: 10.11646/zootaxa.4171.2.1, http://zenodo.org/record/15886
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