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    Figs 21-23 in New genera and species of Afrotropical Ancylolomiini Ragonot, 1889 (Lepidoptera: Pyralidae sensu lato: Crambinae)

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    Figs 21-23. Female genitalia. (21) Diploschistis stygiocrena, DRC. (22) Afroromieuxia bernardlandryi, paratype. (23) A. aarviki, holotype.Published as part of Bassi, Graziano, 2021, New genera and species of Afrotropical Ancylolomiini Ragonot, 1889 (Lepidoptera: Pyralidae sensu lato: Crambinae), pp. 477-486 in Revue suisse de Zoologie 128 (2) on page 484, DOI: 10.35929/RSZ.0058, http://zenodo.org/record/564016

    Figs 9-15 in New genera and species of Afrotropical Ancylolomiini Ragonot, 1889 (Lepidoptera: Pyralidae sensu lato: Crambinae)

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    Figs 9-15. Habitat and male genitalia. (9) Habitat of Afrocharltona oblongissima (photo G. Bassi, Dec., 2010). (10) A. oblongissima, holotype. (11) A. oblongissima, paratype, Botswana, Maun, uncus and gnathos in lateral view. (12) Afroromieuxia bernardlandryi, holotype. (13) Afrocharltona katanga, holotype. (14) Afroromieuxia bernardlandryi, paratype, uncus, gnathos and costal process in lateral view. (15) Diploschistis stygiocrena, DRC.Published as part of Bassi, Graziano, 2021, New genera and species of Afrotropical Ancylolomiini Ragonot, 1889 (Lepidoptera: Pyralidae sensu lato: Crambinae), pp. 477-486 in Revue suisse de Zoologie 128 (2) on page 480, DOI: 10.35929/RSZ.0058, http://zenodo.org/record/564016

    Zovax vangoghi Bleszynski. Moreover, I 1965

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    Zovax vangoghi Błeszyński, 1965 Zovax vangoghi Błeszyński, 1965b: 7. HOLOTYPE: ZSM; ♀; Sudan, Ed Damer, Hudeiba, 23. VII. 1962, Leg. R. Remane; GS 4062 SB. PARATYPES: BMNH, ZSM, USNM; 27 33, 1♀♀; same data as holotype. DISTRIBUTION: Known only from the type locality.Published as part of Bassi, Graziano, 2013, Notes on some Old World Prionapterygini Landry, 1995 (Lepidoptera: Pyraloidea, Crambidae, Crambinae), with descriptions of new species, pp. 131-160 in Revue suisse de Zoologie 120 (1) on pages 133-134, DOI: 10.5281/zenodo.611854

    Zovax Bleszynski. The 1962

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    Zovax Błeszyński, 1962 Zovax Błeszyński, 1962: 130; type species: Prionapteryx whiteheadii Wollaston, 1879, by original designation. DIAGNOSIS: Small to medium sized species with well-developed ocelli and frons with corneous point. Forewing with or without hook. Male genitalia with uncus and gnathos well-developed; valvae without basal costal process. Female genitalia with dorsal sclerite in subrectangular papillae anales; apophyses posteriores basally bulged; corpus bursae with lateral pouch. DIFFERENTIAL DIAGNOSIS: The most closely related genus seems to be Mesolia Ragonot. The male genitalia of Mesolia have flat and sclerotized coremata (Fig. 15), and the gnathos shorter than uncus. The female genitalia of Zovax are most similar to Mesolia, differing by the presence of a dorsal sclerite in the papillae anales, the apophyses posteriores basally bulged, and the corpus bursae with a moderate lateral pouch. REMARKS: Further investigations are needed to fully understand the phylogenetic relationships among the species of this genus. Coremata are very delicate structures, rarely considered in slide mounting. In absence of fresh material I cannot confirm the presence of coremata in Z. whiteheadii and in Z. vangoghi Błeszyński. Moreover, I could not yet study females of these two species, and they are very important for generic relationships.Published as part of Bassi, Graziano, 2013, Notes on some Old World Prionapterygini Landry, 1995 (Lepidoptera: Pyraloidea, Crambidae, Crambinae), with descriptions of new species, pp. 131-160 in Revue suisse de Zoologie 120 (1) on page 133, DOI: 10.5281/zenodo.611854

    Glaucocharis maculosa Bassi & Mey 2011

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    Glaucocharis maculosa Bassi & Mey, 2011 Glaucocharis maculosa Bassi & Mey, 2011: 241, 242, figs 294-296, pl. 37 fig.1. HOLOTYPE: TMSA; 3: [RSA] Buffelspoort, 15. XII.[19]24, A.J. T. Janse legit; (not dissected). PARATYPES: – NAMIBIA. – CB; 1♀; Abachaus, S [outh] W[est] A[frica], Jan [uary] [19]’45, G. Hobohm legit; GS 4023 GB. – MFNB and CB; 633, 3♀♀, Otavi Mts., 21.II.2007, J. Deckert legit, GS 5189 and 5199 GB. – MFNB; 433, 10♀♀; Waterberg N. P., Okatjikona, LF, 14-18.II.2008, W. Mey legit. – MFNB; 1♀; Namibia, Outjo, 4.II.2009, W. Mey legit. – RSA. – TMSA; 13; same label of the holotype. – TMSA and CB; 13, 2♀♀, Pretoria, 10.X. [19]’17, 14.X.’16 and 9.2. [19]’13, A.J. T. Janse legit. – CB; 13, 1♀; Modderpoort, 18. XII. [19]’24, A.J. T. Janse legit; GS 3660 GB. – TMSA; 1♀; Nylstroom, 21. XII.25, A.J. T. Janse legit; GS 4075 GB. – TMSA; 1♀; Pret. [oria] North, 24.X.1924, C. J. Swiestra legit. ETYMOLOGY: The name was derived from maculosus-a (Latin), spotted, referring to the coloration of the forewings of this species. REMARKS: This is the correct type series and etymology.Published as part of Bassi, Graziano, 2013, Notes on some Old World Prionapterygini Landry, 1995 (Lepidoptera: Pyraloidea, Crambidae, Crambinae), with descriptions of new species, pp. 131-160 in Revue suisse de Zoologie 120 (1) on page 158, DOI: 10.5281/zenodo.611854

    Crambus proteus Bassi & Mey 2011

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    Crambus proteus Bassi & Mey, 2011 Crambus proteus Bassi & Mey, 2011: 242, figs 297-299, pl. 36, fig. 8. HOLOTYPE: TMSA; 3; [RSA, Western Cape] Knysna, C [ape] P[rovince], Garden of Eden, 16-20.I.1955, A. J. T. Janse legit (not dissected). PARATYPES: RSA, Western Cape. – CB; 13, 2♀♀; same data as holotype. – SAM; 2♀♀; Cape Town, Table Mt., II.1919, K. Barnard legit. – TMSA; 333, 3♀♀; Cape Town, III.1912, Lord Gladstone legit. – TMSA and CB; 13, 2♀♀; Cape Town, IV.1912, Lord Gladstone legit; GS 3757 GB. – TMSA; 13; Swellendam, 9.III.1980, Scoble & Kroon legit; GS 3578 GB. – TMSA; 1 3; Hogsback (32 27 CA), 24-25.II.1978, M. J. Scoble legit. – TMSA; 8 ♀♀; Deepwalls For. Res. nr Knysna, 17-21.II.1978, M. J. Scoble legit. – TMSA; 13; Tokai, 11.III.1956, CGC. Dickson legit; GS 3771 GB. – CB; 1♀; Kogelberg, Nature Reserve, 14- 21.III.1983, Kroon & Molekane legit; GS 3753 GB. – MFNB and CB; 1133, 5♀♀; Stellenbosch, Assegaibosch N. R., 25.III.2009, LF, W. Mey legit; GS 5193 and 5210 GB. – RSA, Eastern Cape. – Kroon Collection; 1♀; Zingcuka Forest, Mt. Fletcher Dist., 28.XI.1976, D.M. Kroon legit. REMARKS: This is the correct type series. Crambus proteus has two closely related species (Bassi, 2012): C. attis Bassi and C. rossinii Bassi. They are characterized by their similar external appearance and genitalia of both sexes, C. proteus usually differing in the more ochreous tinge of the dark scales in forewings, in male genitalia the gnathos is only slightly longer than the uncus, the tegumen is S-shaped, the costal process of the valva is stronger and longer, the tip of the phallus is longer, and in female genitalia the lateral processes of the sterigma are pointed as opposed to rounded in C. attis and elongate in C. rossinii.Published as part of Bassi, Graziano, 2013, Notes on some Old World Prionapterygini Landry, 1995 (Lepidoptera: Pyraloidea, Crambidae, Crambinae), with descriptions of new species, pp. 131-160 in Revue suisse de Zoologie 120 (1) on pages 158-159, DOI: 10.5281/zenodo.611854

    Afrocharltona katanga Bassi 2021, sp. n.

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    Afrocharltona katanga sp. n. Figs 8, 13 Material examined Holotype: Male;[DRC], H[au]t Katanga, Tshinkolobwe, 5.x. [19]30, J. Romieux [legit], GS 5970 GB, MHNG. Etymology: The species is named after the Province of the Democratic Republic of the Congo where the holotype was collected and is treated as a noun in apposition. Diagnosis: The forewing with pale yellow ground colour flushed with pink grey and the brown hindwing with yellow, pink and grey suffusion distinguish A. katanga from A. oblongissima. The male genitalia are similar to those of A. oblongissima but the valva and apex of the costal arm are shorter and the phallus has two cornuti in the vesica. Description (Fig. 8): Wingspan 36 mm. Labial palpus 2.5 x eye diameter, chestnut brown, apically paler. Maxillary palpus basally chestnut brown, apically paler. Antenna thickened, brown with costa paler. Frons rounded, slightly produced, off-white. Vertex and patagia pale yellow. Tegulae and thorax pale yellow, sprinkled with brown. Forewing and hindwing as in Fig. 8. Forewing sprinkled with brown, more intensely basally; suboval brown dot after cell; fringes bright ivory yellow. Underside pale yellow, suffused with brown dorsally in hindwing. Abdomen pale yellow. Legs yellow; tibial spurs asymmetrical, first pair longer than apical pair. Male genitalia. (Fig. 13). Uncus as long as gnathos, strongly bulged dorsally and with apex blunt and notched. Gnathos with pointed and slightly upcurved apex. Tegumen almost twice as long as uncus, narrow. Vinculum stout, subtriangular. Pseudosaccus narrow, concave. Juxta suboval, with subtriangular protrusions medioventrally. Valva extremely elongated, narrowing apically, with rounded cucullus; costal arm slightly longer than valva, strongly sclerotized, with apical pointed thorn. Phallus with bent phallobase: vesica with two light sclerotized cornuti and minute scobinations. Female unknown. Biology: Unknown. Distribution: Presently known only from the Haut- Katanga province in the DRC.Published as part of Bassi, Graziano, 2021, New genera and species of Afrotropical Ancylolomiini Ragonot, 1889 (Lepidoptera: Pyralidae sensu lato: Crambinae), pp. 477-486 in Revue suisse de Zoologie 128 (2) on page 481, DOI: 10.35929/RSZ.0058, http://zenodo.org/record/564016

    Afroromieuxia ustjuzhanini Bassi 2021, sp. n.

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    Afroromieuxia ustjuzhanini sp. n. Figs 4, 16, 20 Material examined Holotype: Female; East Africa, Zambia, Northern Zambia Prov[ince], Mutinondo Wilderness, 12°27’S 31°17’E, 01.i.2011, Kovtunovich & Ustjuzhanin [legunt], GS 5330 GB, 58550 Collezione Bassi, RCGB. Etymology: The species is named after one of its collectors, Petr Ustjuzhanin (Altai State University, Barnaul, Russia), well-known Pterophoridae and Alucitidae specialist, with many thanks for his friendship. Diagnosis: For differences from A. aarviki and A. bernardlandryi externally and in the female genitalia, see the diagnosis under A. aarviki. Description (Fig. 4): Wingspan 29 mm. Labial palpus 4 x eye diameter, bronze brown bordered white; inner side white. Maxillary palpus basally bronze brown, apically white. Antenna filiform, with scape white, then bronze brown. Frons rounded, slightly produced, offwhite with edge bronze brown. Vertex white suffused with pale yellow laterally and with brown line medially. Patagia white with lateral edge dark brown. Tegulae dark brown with external border white. Thorax brown, off-white distally. Forewing and hindwing as in Fig. 4 and under diagnosis of A. aarviki. Forewing fringes bright silvery grey with medial band white. Underside of forewing brown medially, suffused with white elsewhere; underside of hindwing brown along costa, then white. Abdomen: first segment and anal tuft white, other segments off-white densely suffused with grey brown. Legs bronze brown with inner side white; tibial spurs small, asymmetrical, both pairs of same length. Female genitalia (Fig. 20). Papillae anales produced ventrally, densely covered with setae; lower edge not traceable, fused with ninth abdominal segment. Ninth abdominal segment sclerotized, strongly bulged dorsally. Apophyses posteriores basally only recognizable by a comma-like sclerotization, ending shortly pointed close to bases of apophyses anteriores. Abdominal segment VIII well developed, sclerotized. Apophyses anteriores short, as long as apophyses posteriores. Ostium bursae moderately enlarged, sclerotized, slightly produced. Ductus bursae half the length of corpus bursae, funnelshaped for 2/3, then cylindrical. Ductus seminalis branching off at 2/3 of ductus bursae. Corpus bursae suboval, medially bulged, with a patch of spinulae medio-ventrally. Male unknown. Biology: Unknown. The holotype was attracted to artificial light in open mixed forest (Fig. 16). Distribution: Presently known from Northern Zambia only.Published as part of Bassi, Graziano, 2021, New genera and species of Afrotropical Ancylolomiini Ragonot, 1889 (Lepidoptera: Pyralidae sensu lato: Crambinae), pp. 477-486 in Revue suisse de Zoologie 128 (2) on pages 485-486, DOI: 10.35929/RSZ.0058, http://zenodo.org/record/564016

    Afrocharltona Bassi 2021, gen. n.

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    Afrocharltona gen. n. Figs 7-11, 13, 17-19 Type species: Afrocharltona oblongissima sp. n. Diagnosis: In the adult the slender forewing distinguishes Afrocharltona from Charltona. Some Prionotalis [i.e. P. balia (Tams, 1932)] also have forewing yellow marbled with brown, but their apex of forewing is more pointed. In the male genitalia the combination of modified uncus and the very long and narrow valva represents a distinctive feature. In the female genitalia the lateral extension of medium length branching off in the middle of the ductus bursae and the ductus seminalis originating close to the lateral extension, are unique among related genera except for the prepiella species complex in the inornata group in Ancylolomia (Bassi & Trematerra, 2014). In this complex, however, the lateral extension is always very thin. The holotype and the female paratype from Zambia of A. oblongissima are barcoded, showing intraspecific distance of 1.27% and distance of 9.36% from Charltona tritonella (Hampson, 1898) and 7.74% from Charltona plurivittalis Hampson, 1910. However, the paucity of available barcoded specimens of Ancylolomia and allied genera does not allow a comprehensive DNA comparison, and in the future more molecular markers should be analysed to be conclusive on the relationships to other genera of Ancylolomiini. Etymology: The generic name is a combination of afro (from Africa) and Charltona, the probably closest genus. The gender of the new genus is feminine. Description: Medium to large species, with labial palpi rather short (2.5 - 3 x eye diameter in side view), ocelli and chaetosemata poorly developed, narrow forewing with rounded apex and arched termen. Male genitalia. Uncus slightly longer than gnathos (1.1 x), heavily bulged dorsally and with blunt and wrinkled apex. Gnathos subtriangular, slightly upcurved apically. Tegumen with slender arms, tegumen roof 0.3 of tegumen arm length. Vinculum v-shaped, moderately produced dorsally. Juxta suboval, more or less folded medially. Pseudosaccus a small plate fused with juxta. Valva strongly elongated, tapering towards apex; costal process always developed over cucullus. Phallus stout, 0.6 x valva length, with phallobase downcurved. Female genitalia. Papillae anales subtriangular, rounded ventrally. Apophyses well developed, roughly the length of papillae anales. Ostium concave, with sclerotized edge. Ductus bursae elongated, sclerotized and bearing a narrow lateral extension. Ductus seminalis originating at the middle of ductus bursae. Corpus bursae oval, wrinkled. Biology: Unknown. Distribution: Central Africa and Northern part of Southern Africa.Published as part of Bassi, Graziano, 2021, New genera and species of Afrotropical Ancylolomiini Ragonot, 1889 (Lepidoptera: Pyralidae sensu lato: Crambinae), pp. 477-486 in Revue suisse de Zoologie 128 (2) on pages 477-479, DOI: 10.35929/RSZ.0058, http://zenodo.org/record/564016

    Prionapteryx Stephens 1834

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    Prionapteryx Stephens, 1834 Prionapteryx Stephens, 1834: 316; type species: Prionapteryx nebulifera Stephens, 1834, by monotypy. Alloea Turner, 1947: 37; type species: Alloea xylochroa Turner, 1947, by monotypy. Calarina Walker, 1866 b: 1769, 1770; type species: Calarina albirenella Walker, 1866, by monotypy. Hypotomorpha Rebel, 1892: 252-253; type species: Hypotomorpha lancerotella Rebel, 1892, by original designation. Loxophantis Meyrick, 1935: 570; type species: Loxophantis triplecta Meyrick, 1935, by monotypy. Syn. n. Nuarace Walker, 1863: 188; type species: Nuarace eugraphis Walker, 1863, by monotypy. Pindicitora Walker, 1863: 134; type species: Pindicitora thysbesalis Walker, 1863, by subsequent designation. Platytesia Strand, 1918: 255; type species: Platytes alikangiella Strand, 1918, by monotypy. DIAGNOSIS: Prionapteryx differs from Mesolia and Zovax in the male genitalia with a basal costal process on the valva and without coremata. The basal costal process is shared with Surattha, but the latter differs in having a strongly arched uncus and the phallus with a pointed tip. The female genitalia can be distinguished from those of Mesolia, Zovax and Surattha by the triangular papillae anales, the long, tubular 8th abdominal segment and by the very strongly developed intersegmental membranes between abdominal segments VII and VIII and abdominal segments VIII and IX. REMARKS: The genus Prionapteryx includes 57 species (Nuss et al., 2012) since Błeszyński (1967: 92) synonymized in this genus a number of genera. Some of these species have to be returned to Surattha Walker since this name was revised as valid by Bassi & Mey (2011: 234). On the other hand, Loxophantis has to be considered as a junior synonym of Prionapteryx as shown by the study of both external features and genitalia of the type species. FIGS 18-23 Prionapterygini spp., male genitalia and sclerotizations of abdominal segment VIII, scale bar 0.5 mm. (18) Zovax venus sp. n., paratype GS 5350 GB. (19) Z. venus sp. n., paratype GS 5338 GB. (20) Z. whiteheadii Wollaston, lectotype. (21) Z. vangoghi Błeszyński, paratype 4360 GB, ZSM. (22) Mesolia meyi sp. n., holotype; phallus and juxta from paratype 5321 GB. (23) M. uniformella Janse, paratype, Umvuma, 20.XII.1917, AJT Janse legit, type TMSA 1591, GS 3065 GB.Published as part of Bassi, Graziano, 2013, Notes on some Old World Prionapterygini Landry, 1995 (Lepidoptera: Pyraloidea, Crambidae, Crambinae), with descriptions of new species, pp. 131-160 in Revue suisse de Zoologie 120 (1) on pages 142-143, DOI: 10.5281/zenodo.611854
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