7,001 research outputs found
Influence of ac ageing on space charge dynamics in LDPE
Polymeric materials have been widely used as insulation in power industry due to their excellent electrical properties. However, these properties deteriorate in time irreversibly when the material is subjected to electric stress. Although space charge is believed to play an important role in ac ageing, exact mechanisms are poorly understood due to very limited experimental data. In the present work efforts have been made to investigate the influence of ac ageing on space charge dynamics in low-density polyethylene (LDPE). LDPE films with 200mm were aged at 50 kV/mm at 50 Hz for various times at ambient temperature. Space charge dynamics in the samples prior to and after ageing were monitored using the pulsed electroacoustic (PEA) technique under dc electric stress. The results indicate that there is a significant amount of homocharge accumulation in the unaged sample due to charge injection. These injected charges are the captured by the deep traps originated from the interface between crystalline and amorphous regions in LDPE. Ageing under ac condition does not necessarily lead to an increase in amount of charge in the bulk but leads to an increase in mobility of charge carriers. Chemical analysis by infrared spectroscope (FTIR) reveals there are chemical changes taken place in the bulk of the material after ac ageing. It is believed that the chemical changes introduce shallow traps which promote the movement of charge carriers in the bulk. Consequently, the injected charges spread across the sample
Arganthomyza bivittata Rohacek & Barber 2013
Arganthomyza bivittata Roháček & Barber, 2013 (Figs 107, 109, 126, 128–145) Arganthomyza bivittata Roháček & Barber, 2013: 21. Type material. HOLOTYPE: ♂, “CAN:ON: SSMarie, Base-line Rd., 22.vi.2005, KNBarber, sweeps, Aster, Rubus, Equisetum, Carex, ferns, under aspen 46°31.40'N 84°24.40'W ” and “ HOLOTYPUS ♂, Arganthomyza bivittata sp.n., J. Roháček & K. N. Barber det. 2011” [red label] (DEBU, intact, see Fig. 107). PARATYPES: 173 ♂♂ 156 ♀♀ (AMNH, CASC, CNCI, DEBU, INHS, LACM, LEMQ, RBCM, SMOC, USNM) (details in ROHÁĆEK & BARBER 2013). Other material examined (not included in type series). 1 ♂ 4 ♀♀ (CNCI, LEMQ, RBCM, damaged, see ROHÁĆEK & BARBER 2013). Additional records. CANADA: ONTARIO: Fergus nr. Guelph, Grand River, riverside vegetation, 30.vii.1994, 2 ♂♂, J. Roháček leg. (SMOC, 1 ♂ genit prep.); Moosonee, 51°14.75'N 80°40.33'W, sweeps, mostly Rubus, Ribes, under Populus, 9.vii.2014, 1 ♀; Moosonee, 51°14.79'N 80°40.35'W, sweeps, mostly Solidago, Calamagrostis, 9.vii.2014, 1 ♀; Moosonee, 51°16.33'N 80°39.11'W, sweeps, mostly Rubus, Impatiens, under Salix, Alnus, 10.vii.2014, 4 ♂♂ 1 ♀, 11.vii.2014, 2 ♀♀; Moosonee, 51°16.54'N 80°39.00'W, sweeps, Equisetum, Rubus, Cornus, graminoids, edge of wet forest trail, 10.vii.2014, 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[te.] Marie, Baseline Rd., 46°31.40'N 84°24.40'W, lab-reared from 1 ♀ [with following neld collection data] sweeps, Aster [Doellingeria], Rubus, Equisetum, Carex, ferns, under aspen, 26.vi.2005, [plus additional rearing data], 3 ♂♂ 2 ♀♀ [each with empty puparium in gelatin capsule], K. N. Barber leg. (CNCI); S[ault] S[te.] Marie, Birchwood Pk., mixed forest, 27.vii.1986, 1 ♀; S[ault] S[te.] Marie, Bristol Pl[ace] Pk., 46°30.77'N 84°16.66'W, sweeps, Impatiens, Clematis, Equisetum, Rubus, ferns, Phalaris, 27.vii.2009, 1 ♂, both K. N. Barber leg. (both INHS). QUEBEC: Lac Roddic, 16 km S Maniwaki, 22.vi.1991, 1 ♀, M. Barták leg. (MBPC); Roddic Lake, pan traps, 1–15.viii.1982, 2 ♂♂, L. Huggert leg. (MZLU, 1 ♂ genit. prep.). Material of Arganthomyza cf. bivittata (questionable status). CANADA: NOVA SCOTIA: Lockeport, 25.vii.1958, 1 ♀, J. R. Vockeroth leg. (CNCI, genit. prep.). Diagnosis. Male 2.12–2.94 mm, female 2.77–3.73 mm. Largely yellow, with some brown pattern on head, thorax and abdomen (Figs 107, 109), sparsely light grey microtomentose and relatively shining. Except for dorsal pale stripes, occiput darkened in dorsal half extending medially onto ocellar triangle and laterally around bases of vertical setae; dark notal stripes in dorsocentral lines usually well developed and extending onto scutellum; apical tarsomeres and epandrium dark brown. pvt setae unusually (uniquely) long. Scutellum uniquely with small setulae in addition to marginal setae. Frontal triangle short and very narrow, reaching only midpoint of frons. Mid and hind basitarsus without short thickened setae. T1 and T2 dorsally separate, only laterally fused. Wing hyaline (Fig. 126). Male genitalia (see Figs 128–135 for details). Genitalia generally resembling those of A. acuticuspis and A. duplex. Epandrium (Figs 128, 129) brown, slightly higher than long. Gonostylus (Figs 128–130) yellow, nat, in sublateral (widest extension) view subtriangular and hence most similar to that of A. acuticuspis, but broader, with less acute tip, and more bent medially (cf. Fig. 129). Female postabdomen and genitalia (see Figs 136–145 for details). Postabdomen (Figs 136–138) at 6th segment wider than in A. acuticuspis and A. duplex. T7 and S7 completely fused to form ring-shaped tergosternum T7+S7 (Figs 137, 138); anteroventrally with long, dark, nnely sinuate transverse ledge-like band, spiracles situated posterior to its lateral ends (contrasting with its two nearest relatives). Ventral receptacle (Fig. 141) elongate, slender, similar to that of above species but its proximal part wider. Spermathecae (1+1) broadly ovoid (Figs 140, 143), subequal in size, with dark transversely striated (striae usually interrupted) surface except for very basal part carrying a few (4–5) short, blunt and curved spines on bulges around duct insertion; duct without distinct cervix. Discussion. Arganthomyza bivittata belongs to the A. duplex group (see above) and is the sister species of A. duplex as demonstrated by both ROHÁĆEK & BARBER (2013) and ROHÁĆEK & TÓTHOVÁ (2014). Arganthomyza bivittata can be easily distinguished from congeners by the largely yellow body (mesonotum with only a pair of narrow longitudinal brown vittae which can sometimes be absent, particularly in females; abdomen with yellow preabdominal terga and sterna), unusually long pvt setae, long medial rows of ac microsetae, uniquely setose scutellum with several microsetulae in addition to 2 usual pairs of long sc setae, and densely setose abdominal sclerites. An unusually dark female specimen was reported and discussed by ROHÁĆEK & BARBER (2013) and listed separately above. No similar specimens have yet been found. Biology. The habitat of this species can be generally described as mesic mixed or deciduous forest with relatively dense undergrowth of various components in Ontario (see Biology under A. duplex). It is still not clear which components of this undergrowth are being used by the larvae (see account of rearing attempt below). This eastern habitat can be contrasted with some of the western Canadian collection sites where open areas dominated by grasses yielded specimens (Alberta: Dunvegan and Cadomin). Adults have been collected from 14 June (Quebec: Hull) to 22 September (Ontario: Sault Ste. Marie). The attempt at rearing this species was based on a single female ovipositing from 28 July to 4 September 2011 at 25°C. A choice of oviposition substrate was represented by nve species of plants on which a total of 39 eggs was laid (Dryopteris carthusiana – 18, Equisetum arvense – 10, Thalictrum sp. – 6, Doellingeria umbellata – 4, and Carex sp. – 1). The stems of Thalictrum sp. and D. umbellata were judged to be too dry and coarse and were not being used by the larvae so subsequent larval maintenance relied on the other three species as a mixture. Anecdotally, the rachises of D. carthusiana were judged to be more acceptable as a rearing substrate. They were presented after being split lengthwise to provide more ready access for the larvae but in so doing provided tight crevices within a nbrous matrix not offered by the other two plants. The overwintering period imposed on the surviving 24 larvae at 4°C from 17 December 2011 to 26 March 2012 eventually produced a total of nine puparia yielding nve adults distributed across the three plants as: D. carthusiana – 5 larvae yielding 2 puparia, E. arvense – 2 yielding 2, and Carex sp. – 2 yielding 1. The pupariation periods for the three males were 14, 15, and 15 days at 22°C while those for the two females were 14 and 15 days. Distribution. Transcontinental in Canada from British Columbia to Nova Scotia (Alberta, British Columbia, Manitoba, Nova Scotia, Ontario, Quebec, Saskatchewan) with a single record from the United States of America (Michigan: Carp Lake) (ROHÁĆEK & BARBER 2013, see Table 2).Published as part of Roháćek, Jindřich & Barber, Kevin N., 2016, Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera), pp. 1-412 in Acta Entomologica Musei Nationalis Pragae (suppl.) (suppl.) 56 on pages 83-89, DOI: 10.5281/zenodo.427282
A power sharing series power BJT array with isolated low voltage control for AC power control applications
A technique for a continuously variable AC resistance using a series BJT array is presented. This array provides high power dissipation capability and uniform voltage and power distribution across the individual transistors. The array, controlled using a set of optoisolators to maintain the electrical isolation between the control circuits and the power stage, could be used as the basis to develop several useful techniques including a solid state AC regulator with comparable performance to the commonly used ferro-resonant systems; a linear AC electronic load suitable for testing UPS and other power conditioners; and, in other AC power control applications such as switching capacitors in AC resonant circuits
Barber Gamecock Float, 1968 Homecoming Parade 1
Homecoming was held October 26, 1968 at Jacksonville State University. The homecoming slogan was Massacre the Indians. Shown is a float during the homecoming parade. An experienced gamecock barber scalps an Indian. The float reads, Barber Gamecock 23 years experience and Gamecocks -- Scalp the Indians. Martin Hall can be seen in the background.https://digitalcommons.jsu.edu/lib-ac-histimg/31363/thumbnail.jp
Experimental High-Frequency Parameter Identification of AC Electrical Motors
In order to predict conducted electromagnetic interference in inverter-motor drive systems, high-frequency (HF) motor models are requested and the involved parameters have to be available. In previous studies, the authors have presented an accurate HF model for induction motors and they have defined the procedures to identify the model parameters. In this paper, these results are extended to several types and sizes of industrial ac motors such as induction, synchronous reluctance (without interior permanent magnets), and brushless motors. The model parameter-identification procedure has been improved, and it is based on a least-squares data fitting applied to the measured magnitude and phase-frequency-response curves of the phase-to-ground and the phase-to-neutral impedances. The aim of this paper is to provide quick indications to select the suitable values of the HF model parameters, with reference to the size and type of the ac motor, to evaluate the HF voltage and current components in inverted-fed ac motor system
Using the AC Drive Motor as a Transducer for Detecting Electrical and Electromechanical Faults
Condition monitoring of AC motors is a subject area that has received extensive research. Whether this monitoring is carried out on a scheduled basis by engineer intervention, or continuously using an on-line unit, the results of this testing enable preventative maintenance work to be a carried out earlier, before any major failure occurs. Monitoring using vibration analysis is the most common and depending on the plant, can be done once or twice a year. This is usually limited to the condition of motor bearings and is not commonly used to detect failures in the motor electromagnetic systems. Monitoring units that use motor current measurements are also available, but these are less widely-used and usually on major plant motors (>250kW for example) that have a large capital outlay to replace.
The industry drivers – as always – are maximum plant and machinery uptime, with the minimal amount of scheduled maintenance. If maintenance is carried out too regularly, costs rise significantly not only due to the maintenance activity itself, but disruption to production schedules. Maintenance schedules that are too infrequent can result in an unacceptable rise in total failures of plant that are unexpected and may cause a significant amount of production disruption and downtime, especially if this occurs during out-of-hours working time. However, industry now faces another big challenge and one that has had a good share of exposure over the last few years. It is of course, the drive to reduce carbon emissions and with it the amount of energy that a plant itself consumes. What has brought this more to the fore recently is the significant rise in energy costs. Whilst product margins have to remain the same, many companies energy costs have seen a two to three-fold increase in energy budgets in the last few years alone. For industry processes that have a significant amount of fan and pump applications, the manufacturers of low-cost AC inverters have saturated the lower-performance market of inverter drives such that any drive can control these type of fan and pump applications, where accurate speed control is not a major driver.
Unfortunately, this can be a step backwards for end-users of plant that use equipment to monitor motor condition via motor current signals. Additionally, vibration analysis that relies upon ‘base-lining’ motor data when the AC motor is running at base speed may not give accurate readings when the motor is under inverter control and running at a different speed.
For manufacturers of AC inverter drives in this low-end market, it can be difficult to sell a product from one manufacturer over another without the unit having a “USP”, or Unique Selling Proposition. Most decisions taken on inverter equipment purchase at this level are usually in favour of the equipment that costs the least to purchase. Credibility of manufacturers based on product history and perceived reliability cuts little ground with an ever cost-conscious industry.
This is where the research into diagnosis of faults on inverter driven motor systems can provide just this USP for manufacturers. If the incorporation of on-line diagnosis for simple inverter applications can be brought to a typical inverter unit at a reasonable cost, the manufacturer who can offer this gains a unique foothold in the marketplace – a drive that can monitor and signal that the motor it is driving is showing signs of early failure.
It will be sensible to limit this research to simple inverter applications as high-end inverter drives that operate equipment such as high-speed printing presses, rotary shears will be more difficult to model and simulate than a fan or pump application.
It is hoped that a typical inverter drive can relay enough detailed information about the load which it is driving to allow this to be used for abnormal motor load conditions as this will provide a platform on which to extend the research beyond this MSc and into the realms of incorporating such technology into a drive manufacturers equipment. If this can be done without major modification to an inverter, then it may be easier to implement in equipment offered by different manufacturers.
It is quite possible that this technology could be licensed under a name that guarantees the performance of the condition monitoring algorithms and reliability from one drive manufacturer to another
Anthomyza furvifrons Roháćek & Barber 2016, sp. nov.
Anthomyza furvifrons sp. nov. (Figs 518, 519, 524, 527–543) Type material. HOLOTYPE: ♂, “CAN: ON: ~74kmNNE The-ssalon, shore of Mississagi R., 17.vii.2010, KNBarber, sweeps, graminoids, herbs, Equisetum spp. 46°53.94’N 83°16.23’W ” and “ Holotypus ♂ Anthomyza furvifrons sp. n., J. Roháček & K. N. Barber det. 2014” (red). The specimen is in perfect condition, with exposed genitalia and highly visible gonostyli (see Fig. 518) (CNCI, intact). PARATYPES: CANADA: NEW BRUNSWICK: Birch Cove, nr. Chamcook, 4.vii.1965, 1 ♂ 1 ♀ (1 ♂ most of legs missing); Chamcook, 30.vi.1965, 1 ♀, on goldenrod, 7.viii.1957, 1 ♀, Centaurea, 7.viii.1957, 1 ♀, all G. E Shewell leg.; Maces Bay, Hwy 790, 9.vii.1971, 1 ♂ 3 ♀♀ (1 ♀ genit. prep.), B. V. Peterson leg. (all CNCI); Middle Sackville, 45°55.4'N 64°21.4'W, sweep vegetation along old rail line, 19.vii.2002, 1 ♂ 1 ♀, J. Forrest & T. Wheeler leg. (LEMQ 0039175, -40345, 1 ♀ genit. prep.); Musquash, 10. viii.1956, 1 ♀, A. H. Sturtevant leg. (USNM); Sackville, near Mt. Allison Univ., 45°53.9'N 64°22.5'W, sweep old garden in vacant lot, 19.vii.2002, 4 ♀♀, J. Forrest & T. Wheeler leg. (LEMQ 0040333, -36, -39, -41); Woodstock, 8.viii.1956, 1 ♀, A. H. Sturtevant leg. (USNM). NEWFOUNDLAND: Pasadena, sweeping low veg., 24.viii.1984, 1 ♂, low veg. in neld, 1.viii.1984, 1 ♂, L. Hollett leg. (CNCI). ONTARIO: Algonquin Park, 26–30.vii.1955, 2 ♀♀, Sabrosky leg. (USNM); Baptiste Lake, 45°10'N 78°00'W, sweep near lake shore, 25.vii.2000, 1 ♀, J. Forrest leg. (LEMQ 0039493); Bruce Co., Spring Creek at Hwy 6, grass – Typha, 22.vii.2000, 1 ♂, S.A. Marshall leg.(DEBU 00079576); Bruce Peninsula N.P., Dorcas Bay Rd.at Willow Creek, 45°09.4'N 81°34.4'W, sweeps, mostly creekside graminoids, 3.vii.1999, 1 ♀; same locality but Singing Sands, 45°11.6'N 81°34.7'W, sweeps, Calamagrostis canadensis, 29.vii.1997, 1 ♀, sweeps, Agropyron / Calamagrostis, 30.vii.1997, 1 ♂, 5.vii.1998, 3 ♀♀, all K. N. Barber leg. (all DEBU); Cochrane, 49°03.54'N 81°04.41'W, sweeps, grasses on edge of hwy. pulloff, 21.vii.2009, 2 ♀♀; ~ 5 km SE Cochrane, 49°01.16'N 80°57.93'W, sweeps, railside Equisetum spp., graminoids, herbs, 18.vii.2009, 8 ♂♂ 15 ♀♀ (1 ♂ genit. prep.); Dryden, 49°47.1'N 92°48.4'W, sweeps, grasses in low area, 10.vii.1999, 1 ♂ 1 ♀ (1 ♀ left wing missing); Dryden, 49°47.27'N 92°48.62'W, sweeps, mixed graminoids/herbs, 17.vii.2008, 1 ♂, all K. N. Barber leg. (all CNCI); Dubreuilville, 48°21.09'N 84°33.90'W, sweeping undergrowth of Pinus / Populus forest, with Clintonia, Vaccinium, ferns, graminoids, 10.vii.2010, 1 ♂, J. Roháček leg. (SMOC, genit. prep.); Dubreuilville, along Magpie River, 48°21.12'N 84°34.04'W, sweeps, Equisetum ssuviatile, Carex, 10.vii.2010, 1 ♂; ~ 35 km WSW Dubreuilville, 2 km SE jct. Hwys.#17 & #519, 48°17.16'N 84°53.34'W, sweeps, roadside Calamagrostis / Scirpus, 23.vii.2003, 1 ♂, both K. N. Barber leg. (both CNCI); ~ 55 km NNW Elliot Lake, S of Rocky Island Lake, 46°49.32'N 82°59.54'W, 455 m, sweeping, vegetation with predominant Scirpus sp., 3.vii.2010, 2 ♂♂ 1 ♀ (1 ♂ 1 ♀ genit. prep.); ~ 61 km NNW Elliot Lake, Three Lakes, 46°49.94'N 83°06.31'W, 425 m, sweeping grasses nr. lakeshore, 3.vii.2010, 4 ♂♂ 1 ♀ (3 ♂♂ genit. prep.), all J. Roháček leg. (all SMOC); Finland, 17.vii.1960, 1 ♀, 21.vii.1960, 2 ♂♂, S. M. Clark leg. (CNCI); ~7.0 km E Foleyet, 48°14.34'N 82°20.75'W, hydro right-of-way, sweeps, mostly Carex utriculata, 13.vii.2013, 2 ♂♂ 1 ♀; Fraserdale, 49°50.71'N 81°36.99'W, sweeps, graminoids, herbs in disturbed area, 19.vii.2009, 2 ♀♀, all K. N. Barber leg. (all CNCI); Greenwater P. Pk., 49°10.91'N 81°16.28'W, sweeps, Carex, Calamagrostis, Phalaris, in creek noodplain, 21.vii.2009, 2 ♂♂ 1 ♀ (DEBU 01502126–28); Greenwater P. Pk., 49°11.05'N 81°16.04'W, sweeps, mostly lakeside Calamagrostis canadensis, 18.vii.2009, 1 ♀ (DEBU 01502104, -05); Greenwater P. Pk., 49°11.34'N 81°17.04'W, sweeps, grasses/herbs in hydro cut, 21.vii.2009, 2 ♂♂ 8 ♀♀ (DEBU 01502244–53); Greenwater P. Pk., Sandbar Lk. Trail, 49°13.10'N 81°17.35'W, sweeps, lakeshore Equisetum spp., graminoids, Caltha, 21.vii.2009, 2 ♀♀ (DEBU 01502104, -05), all K. N. Barber leg.; 6 km W Hagar, 46.47°N 80.48°W, sweep river edge at rest area, 29.vi.2007, 2 ♀♀, T. A. Wheeler leg. (LEMQ 0040285, -86); Hwy 17N & Trout Lake Road, 46°37.563'N 84°17.019'W, sweep, roadside, 23.vii.2011, 1 ♂ 1 ♀, J. E. Swann & D. R. Edwards (BDUC, 1 ♀ genit. prep.); Icewater Creek WS [Watershed], 13.5 km NNE Searchmont, mi.11.5 Whitman Dam Rd., sandy access road, 1.vii.1986, 1 ♀; ~ 25 km WNW Ignace, 49°29.52'N 92°00.83'W, sweeps, fen, mostly Carex utriculata with grasses, 4.vii.2012, 1 ♂ 1 ♀; 40 km S Ignace, Hwy#17, 49°15.6'N 91°07.9'W, sweeps/pooter, roadside Bromus inermis, 10.vii.1999, 1 ♀; ~ 11.9 km N Kejick, 49°43.89'N 95°04.14'W, sweeps, wet ditch, graminoids/ Equisetum, 30.vii.2008, 1 ♀, all K. N. Barber leg.; Kenora, 22.viii.1958, 1 ♀, J. G. Chillcott leg. (all CNCI); Manitoulin Is., 0.7 km N Michael’s Bay Pk., 45°36.5'N 82°06.2'W, sweeps, roadside grasses, 28.vii.1997, 1 ♂, sweeps/ pooter, graminoids in fen nat, 30.viii.2004, 1 ♀; Hwy#17, ~8.5 km NW Marathon, 48°47.69'N 86°26.07'W, sweeps, roadside graminoids, 31.vii.2008, 1 ♂, all K. N. Barber leg.; One-Sided [=Caliper] Lake, 13.vii.1960, 1 ♂, 19. vii.1960, 1 ♂, S. M. Clark leg., 26.vi.1960, 2 ♀♀, 29.vi.1960, 1 ♀, Kelton & Whitney leg. (all CNCI); Otter Rapids, 50°10.91'N 81°38.54'W, sweeps, mostly Calamagrostis canadensis in hydro cut, 19.vii.2009, 2 ♂♂ 3 ♀♀, 20.vii.2009, 2 ♂♂ 5 ♀♀; Otter Rapids, 50°10.96'N 81°37.88'W, sweeps, grasses,herbs on roadside slope, 20.vii.2009, 1 ♂ 1 ♀, sweeps, mostly Schedonorus arundinaceus, on roadside slope, 20.vii.2009, 10 ♂♂ 16 ♀♀, all K. N. Barber leg. (all CNCI); Thunder Bay Distr., mouth of Pic River, N side, 48°36'N 86°18'W, 20.vii.2001, 1 ♀, M. Buck leg.(DEBU 00169254); René Brunelle P. Pk., 49°25.89'N 82°08.44'W, sweeps, roadside Equisetum spp., 19.vii.2009, 1 ♂ 1 ♀ (DEBU 01502002, -03); ~ 2 km E Rossport, Hwy#17, picnic area, 48°50.3'N 87°29.4'W, sweeps of graminoids, 9.vii.1999, 2 ♂♂ 1 ♀ (CNCI), all K. N. Barber leg; S[ault] S[te.] Marie, S. of Algoma U[niversity] College, 46°29.9'N 84°17.2'W, sweeps, Calamagrostis canadensis & Carex aquatilis, 23.vii.1997, 3 ♂♂ 4 ♀♀ (CNCI 2 ♂♂ 3 ♀♀, SMOC 1 ♂ 1 ♀), sweeps, Calamagrostis canadensis / Carex aquatilis, 6.viii.2002, 1 ♀ (CNCI), sweeps/ pooter, Calamagrostis canadensis, 3.viii.1997, 4 ♂♂ 6 ♀♀ (CNCI 3 ♂♂ 5 ♀♀, 1 ♂ genit. prep.; SMOC 1 ♂ 1 ♀), 12.vii.2002, 1♂, sweeps, Calamagrostis canadensis, 29.vii.1999, 2 ♀♀, sweeps, mostly Calamagrostis canadensis, 28.vii.2001, 2 ♀♀ (1 ♀ genit. prep.), 29.vii.2001, 1 ♂ 1 ♀, 6.viii.2001, 3 ♂♂ 2 ♀♀ (1 ♀ genit. prep.), sweeps/ pooter, Carex aquatilis, 3.viii.1997, 1 ♀, sweeps, Carex aquatilis, 12.vii.2002, 1 ♀, 27.vii.2005, 1 ♀, sweeps, mostly Carex aquatilis, 15.vii.2001, 1 ♂, 17.vii.2001, 1 ♂, 18.vii.2001, 1 ♂ 3 ♀♀ (1 ♀ with wing glued to pin); same locality but 46°29.88'N 84°17.19'W, sweeps, Scirpus cyperinus, 21.viii.2004, 2 ♂♂ (all CNCI), all K. N. Barber leg.; S[ault] S[te.] Marie, Baseline Rd., 46°31.41'N 84°24.57'W, sweeps, sparse veg. on disturbed site, 26. vi.2005, 1 ♀; same locality but w. of creek, 46°31.61'N 84°24.68'W, sweeps, Carex edge of alder thicket, 22.vii.2005, 1 ♂; S[ault] S[te.] Marie, Bristol Pl. Pk., 46°30.8'N 84°16.6'W, sweeps/pooter, Calamagrostis canadensis, 1.viii.1997, 2 ♂♂ 1 ♀ (1 ♂ genit. prep.), sweeps/pooter, Phalaris arundinacea, 1.viii.1997, 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[te.] Marie, Finn Hill, 46°31.6'N 84°17.3'W, sweeps, graminoids/composites, 19.vii.2005, 1 ♂ 1 ♀; same locality but 46°31.48'N 84°17.36'W, sweeps, mostly Scirpus microcarpus, 8.vii.2006, 1 ♀; same locality but 46°31.57'N 84°17.37'W, sweeps, Calamagrostis, Rubus, ferns, 15.vii.2006, 1 ♀; same locality but 46°31.67'N 84°17.32'W, sweeps, Calamagrostis canadensis, 30.vii.2004, 1 ♂ 2 ♀♀; same locality but 46°31.63'N 84°17.33'W, sweeps, graminoids, herbs, composites, edge of Populus tremuloides, 8.viii.2008, 2 ♂♂ 2 ♀♀, 10.viii.2008, 1 ♂ 1 ♀, all K. N. Barber leg. (all CNCI); Sault Ste. Marie, Fish Hatchery Road, near Coldwater Creek, 46°34.29'N 84°17.21'W, sweeping graminoids, Impatiens, 9.vii.2010, 2 ♂♂, J. Roháček leg. (SMOC, both genit. prep.); S[ault] S[te.] Marie, Ft. Creek Cons[ervation] Area, 46°32.5'N 84°20.8'W, sweeps, mixed meadow veg., 9.viii.1997, 1 ♂; S[ault] S[te.] Marie, Landslide Rd., Coldwater Ck. noodplain, 46°33.8'N 84°16.6'W, sweeps/pooter, Calamagrostis canadensis, 7.viii.1997, 1 ♀, sweeps, mostly Carex sp., 7.viii.1997, 2 ♀♀; S[ault] S[te.] Marie, Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.2'W, sweeps, mixed meadow veg., 25.vii.1997, 2 ♀♀, sweeps, Phalaris arundinacea, 25.vii.1997, 1 ♀; S[ault] S[te.] Marie, hydro cut nr. Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.0'W, sweeps, mostly sedges, 10.vii.2003, 1 ♀; S[ault] S[te.] Marie, Whitensh Is./St. Mary’s Is., 46°30.68'N 84°21.20'W, riparian graminoids, sweeps, 8.viii.2004, 1 ♀ (genit.prep.); S[ault] S[te.] Marie, Hwy #17 city limits, 46°36.58'N 84°17.83'W, sweeps, mostly Carex / Calamagrostis in wet area, 23.viii.2004, 2 ♂♂ 2 ♀♀; same locality but 46°36.62'N 84°17.85'W, sweeps, Carex gynandra in alder thicket, 4.vii.2016, 1 ♂, all K. N. Barber leg. (all CNCI); ~ 10 km W S[ault] S[te.] Marie, Airport Rd., 46°29.9'N 84°28.9'W, natural gas r[igh]t-of-way, sweeps, graminoids, composites, Equisetum, Rubus, ferns, 25.viii.2008, 1 ♂; same locality but 46°29.72'N 84°28.96'W, sweeps, graminoids, composites, Equisetum, Rubus, ferns, 14.vii.2010, 5 ♂♂ 3 ♀♀, sweeps, graminoids, Equisetum, herbs, 5.viii.2009, 1 ♂ 2 ♀♀, sweeps, mostly Scirpus / Calamagrostis, 4.viii.2004, 1 ♂, all K. N. Barber leg. (all CNCI), sweeping graminoids, composites, Equisetum, Rubus, ferns, 12.vii.2010, 10 ♂♂ 14 ♀♀, J. Roháček leg. (SMOC, 5 ♂♂ 5 ♀♀ genit. prep.); 20 km W S[ault] S[te.] M[arie], Pt. de[s] Chenes Pk., 14.vii.1986, 1 ♂, K. N. Barber leg. (CNCI); MacIntyre Road, north of Sault Ste. Marie, 46°37.403'N 84°18.320'W, sweep, roadside, 23.vii.2011, 1 ♂ 2 ♀♀, J. E. Swann & D. R. Edwards (BDUC); Searchmont, N Hwy #552, 46°51.3'N 84°02.4'W, sweeps, graminoids around gravel pit, 24.vii.1998, 1 ♀; 12.4 km NNE Searchmont, mi.10 Whitman Dam Rd., herb/grass meadow by Goulais R., 23.vi.1986, 2 ♀♀; 18 km NNE Searchmont, mi.15 Whitman Dam Rd., grassy access road, 19.vi.1986, 1 ♂, 29.vi.1987, 1 ♂ 4 ♀♀; ~ 18.8 km NNE Searchmont, Goulais River WS [Watershed], ~mi.15 Whitman Dam Rd., 46°55.7'N 83°56.2'W, sweeps, jackpine plantation, 29.vii.1999, 1 ♀ (abdomen glued to pin), all K. N. Barber leg.; Sioux Narrows, 25.vi.1960, 1 ♂ (genit. prep.), Kelton & Whitney leg. (all CNCI); Thunder Bay Distr., Sleeping Giant Prov. Pk., Sawbill Lk. Trail, 48°21'N 88°50'W, 13.vii.2002, 1 ♂ 1 ♀, M. Buck leg. (DEBU 00199157, -170, 1 ♂ genit. prep.); ~ 21 km NNE Smooth Rock Falls, 49°20.91'N 81°32.01'W, sweeps, Equisetum ssuviatile in wet ditch, 19.vii.2009, 2 ♂♂ 1 ♀, sweeps, ditchside Equisetum spp. [including E. ssuviatile], grasses, herbs, 19.vii.2009, 1 ♂ 2 ♀♀, K. N. Barber leg. (CNCI); ~ 74 km NNE Thessalon, 46°53.94'N 83°16.23'W, shore of Mississagi R., sweeps, graminoids, herbs, Equisetum spp., 5.vii.2010, 9 ♂♂ 12 ♀♀ (CNCI, 1 ♀ genit. prep.), 17.vii.2010, 38 ♂♂ 54 ♀♀ (AMNH, CASC, DEBU, LACM, MCZC, USNM 4 ♂♂ 6 ♀♀ each; CNCI 14 ♂♂ 18 ♀♀ incl. pair in copula, 1 ♀ genit. prep.), sweeps, Equisetum spp. on mud bank, 17.vii.2010, 1 ♂ (CNCI, wing illustration), K. N. Barber leg., sweeping graminoids with Equisetum spp. on muddy shore, 5.vii.2010, 8 ♂♂ 8 ♀♀, J. Roháček leg. (SMOC, 4 ♂♂ 4 ♀♀ genit. prep.); ~ 92 km NNE Thessalon, nr. Mountain Ash Lake, 47°02.98'N 83°10.88'W, sweeping Carex aquatilis on edge of wetland, 4.vii.2010, 1 ♀, J. Roháček leg. (SMOC, genit. prep.); White River, 48°35.5'N 85°16.6'W, sweeps, mixed grasses edge of parking lot, 9.vii.1999, 4 ♂♂ 3 ♀♀, K. N. Barber leg. (CNCI). QUEBEC: Charlevoix, Ste-Aimé-des-Lacs, 47°41'N, 70°18.5'W, sweep vegetation, 5.viii.2001, 1 ♀, V. Dion leg. (LEMQ 0039190); Forillon N. P., Anse Saint-Georges, 48°46'N 64°12'W, sweep at roadside, 16.viii.2001, 1 ♂ 2 ♀♀ (LEMQ 0039172,-78, -79); Gaspé, Forillon N.P., Cap des Rosiers, 48°50'N 64°12'W, sweep grass, 6.viii.2000, 2 ♂♂ (LEMQ 0039262, -63); Gaspé, Forillon N. P., Haldiman St.-Jean R., 48°47'N 64°22'W, sweep salt marsh vegetation, 31. vii.2000, 1 ♂ (LEMQ 0039366); Gaspé, Forillon N. P., Ruisseau Petit Gaspé, 48°54'N 64°21'W, sweep muddy area, 6.viii.2000, 1 ♂ 1 ♀ (LEMQ 0039402, -04), all H. Varady-Szabo leg.; Ile Bonaventure, 3 km from Côte de Percé, 48°30'N 64°10'W, sweep grass, 28.vii.2000, 5 ♀♀, H. Varady-Szabo leg. (LEMQ 0039254, -257, -336, -338, -339), 2 ♀♀, A. Thibault leg. (LEMQ 0039207, -284, 1 ♀ genit. prep.); Îles de la Madeleine, Île de la Grande Entrée, Chemin du Bassin Ouest, 47°32.96'N 61°32.64'W, sweep neld at road, 9.viii.2004, 1 ♀, V. Dion leg. (LEMQ 0040481), sweep neld along road, 9.viii.2004, 1 ♀, S. Boucher leg. (LEMQ 0040505); Laniel, 1.vii.1944, 1 ♂ 4 ♀♀, A. R. Brooks leg.; La Verendrye Prov. Pk., mi.139 Rte.58, 29.vi.1965, 1 ♂, D. M. Wood leg. (all CNCI). UNITED STATES OF AMERICA: MAINE: Greenville, 1.viii.1930, 1 ♀; Pittston, 3.viii.1950, 1 ♂, both A. L. Melander leg. (both USNM). MICHIGAN: Delta Co., 3.vii.1955, 1 ♀, R. R. Dreisbach leg.; Keweenaw Co., Isle Royale, 11., 13.vii.1938, 2 ♂♂, 13., 15.vii.1938, 1 ♂ 1 ♀, G. Steyskal leg. (all USNM); Berrien Co., St. Joseph, 19.vi.1975, 1 ♀, D. D. Wilder leg. (CASC). MINNESOTA: Eaglesnest, 30.vi.1959, 2 ♂♂ 4 ♀♀ (INHS 40,087–089, -105–107, 1 ♀ genit. prep.), 16.vii.1959, 2 ♀♀ (INHS 40,108, -109), 6.vii.1959, 1 ♀ (INHS 40,094), 11.vii.1959, 1 ♂ 3 ♀♀ (INHS 40,095, -111–113), 11.vii.1952, 1 ♀ (INHS 40,098), 25.viii.1958, 1 ♂ (INHS 40,099), 12.viii.1958, 1 ♀ (INHS 40,100), 30.vi.1954, 1 ♂ (INHS 40,102), 6.vii.1954, 1 ♂ (INHS 40,103, genit. prep.), 13.vii.1957, 2 ♂♂ (INHS 40,119, -121), 28.vi.1957, 1 ♀ (INHS 40,126), 5.vii.1957, 1 ♀ (INHS 40,128), 23.vii.1958, 5 ♂♂ 1 ♀ (INHS 40,140, -142–144, -146, -147), all W. V. Balduf leg. (all part of mixed series [with A. vulgaris] with Sabrosky det. as Anthomyza gracilis); Aitkin Co., 14 mi W Willow River, 46.334°N 93.096°W, 27.vii.1997, 1 ♂, D. E. Hansen leg. (CNCI). NEW YORK: Franklin Co., Paul Smiths, 20.vii.1962, 2 ♂♂, J. R. Vockeroth leg. (CNCI). Other material examined (not included in type series). CANADA: ONTARIO: One-Sided [= Caliper] Lake, 1.viii.1960, 1 ♀, S. M. Clark leg. (CNCI, broken, abdomen, wing, 3 legs in gelatin capsule); ~ 10 km W Sault Ste. Marie,Airport Rd., 46°29.72'N 84°28.96'W, natural gas r[igh]t-of-way, sweeping graminoids, composites, Equisetum, Rubus, ferns, 2 ♂♂, 12.vii.2010, J. Roháček leg. (SMOC, used for molecular work); ~ 74 km NNE Thessalon, shore of Mississagi River, 46°53.94'N 83°16.23'W, sweeping graminoids with Equisetum spp.on muddy shore, 5.vii.2010, 1 ♀, J. Roháček leg. (SMOC, headless, genit. prep.). UNITED STATES OF AMERICA: MINNESOTA: Eaglesnest, 5.vii.1957, 2 ♀♀, W. V. Balduf leg. (INHS 40,129, -130, 1 ♀ missing wings & 2 legs, 1 ♀ missing tip of abdomen). Description. Male. Total body length 2.06–2.62 mm; general colour brown to blackish brown (abdomen usually lighter than thorax) brownish grey microtomentose, although less densely than in A. vulgaris. Head somewhat higher than long (Fig. 518), quadrangular and with face distinctly receding in pronle, dark brown and yellow. Occiput dorsomedially very slightly concave, shiny blackish brown due to very sparse microtomentum. Frons relatively narrow, medially largely dull brown, only frontal triangle grey microtomentose and anterior fourth to third of frons orange yellow; orbits (Fig. 519) anteriorly yellow and sparsely narrowly silvery white microtomentose, posteriorly (behind middle of distance between anterior and posterior ors) dark, brownish, with silvery grey microtomentum at eye margin. Frontal triangle narrow, reaching to anterior third of frons. Frontal lunule distinct, yellow, whitish microtomentose. Face narrow, medially less sclerotized and often with darker narrow stripe, otherwise yellow to orange-yellow and dull; parafacialia relatively broad and like gena yellow with sparse whitish microtomentum. Both parafacialia and gena with rather broad ochreous to pale brown marginal stripe; ventral angle of postgena yellow, whitish microtomentose; its dorsal part brown and greyish microtomentose, abruptly contrasting posteriorly with shiny occiput (Fig. 518); mouthparts (except for palpi) ochreous to pale brown, clypeus (small) and prementum darkest. Cephalic chaetotaxy: pvt small, convergent but usually not crossed; vti longest of cephalic setae; oc often as long as vti but nner; vte and posterior ors slightly shorter than vti; 3 ors but only 2 long, widely spaced, middle ors slightly to distinctly shorter than posterior, and anterior ors reduced to setula (sometimes enlarged); 1–2 (rarely 3) pairs of medial microsetulae in the anterior third of frons; 1 small setula behind vte (slightly longer than postoculars); postocular setulae (6–7) short, sparse, in single row; postgena with several setulae and 2 (1 longer) usual posteroventral setae; 1 long but thin vi (about as long as middle ors); subvibrissa small and weak, slightly longer than foremost peristomal setula; only 3–4 nne peristomal setulae. Palpus well developed, elongately clavate, yellow, with 1 ventral preapical seta and about 5 shorter subapical and ventral setulae. Eye moderately convex, large, of rounded quadrangular outline (Fig. 518), anteroventrally slightly widened, with longest diameter oblique and 1.3–1.4 times as long as the shortest. Smallest genal height about 0.14 times as long as shortest eye diameter. Antenna strongly geniculate, entirely yellow; 1st nagellomere nattened laterally, with short white pilosity on anteroventral margin. Arista brown to dark brown including basal segments, about 1.9 times as long as antenna, shortly ciliate. Thorax very slightly or not narrower than head, entirely blackish brown (mesonotum usually darker) and densely brownish grey microtomentose (Fig. 518), almost dull. Thoracic chaetotaxy (all macrosetae relatively weak): 1 hu (shorter than anterior npl); 2 npl (anterior longer); 1 small prs (shorter than hu); 1 sa (shorter than pa); 1 moderate pa; 2 postsutural dc (the shorter anterior about as long as anterior npl) and 4–6 dc microsetae in front of them; ac microsetae sparse, in 4 rows on suture, in 2 rows between anterior dc and reaching only slightly beyond them (no ac microsetae between posterior dc); 2 sc, laterobasal sc weak, about as long as prs, apical sc as long as posterior dc (longest thoracic setae); 1 hair-like ppl; 2 relatively long stpl (posterior longer than anterior npl, anterior usually shorter) and a few (1–2) setulae in dorsal half of sternopleuron; its ventral part with 3–5 longer setae. Scutellum rounded triangular, distinctly convex dorsally. Legs bright yellow (Fig. 518), only 2 terminal tarsal segments of all tarsi pale brown to brown; sometimes also coxae partly darkened, ochreous (never brown). f 1 with ctenidial spine rather variable, as long as to distinctly longer than maximum width of t 1 and with long thin setae in posteroventral (longer setae) and posterodorsal row; both f 2 and f 3 uniformly setulose; t 2 with relatively short and weak ventroapical seta; t 1, t 3 and all tarsi simply setulose. Wing (Fig. 524) relatively long and narrow but usually wider than that of A. vulgaris, with pale brown veins and hyaline unicolourous membrane (light ochreous brown). C with very small and sparse spinulae between apices of R 1 and R 2+3. R 2+3 long, slightly sinuous, parallel to C but with apex distinctly upcurving to it; R 4+5 almost straight to very slightly sinuous, parallel to or slightly divergent from M apically. Dis
Fungomyza buccata Rohacek & Barber 2004
Fungomyza buccata Roháček & Barber, 2004 (Figs 33–46) Fungomyza buccata Roháček & Barber, 2004: 135; ROHÁĆEK (2009a): 88 (key), 110–111 (phylogeny). Type material. HOLOTYPE: ♂, “ Stone Mt., DeKalb Co., Ga. XI-II-1953, Dodge”, “ ♂ ”, “ HOLOTYPUS ♂ Fungomyza buccata sp. n., J. Roháček & K. N. Barber det. 2004” (red), deposited in USNM (intact). PARATYPES: 5 ♂♂ 6 ♀♀ (DEBU, LEMQ, SMOC, USNM) (details in ROHÁĆEK & BARBER 2004). Other material examined (not included in type series). 1 spec. (USNM, only wing preserved, see ROHÁĆEK & BARBER 2004). Additional records. UNITED STATES OF AMERICA: ALABAMA: Baldwin Co., Silverhill, 30.53°N 87.74°W, (Site 9), Malaise trap, 15.xii.2004, 3 ♀♀, E. Benton leg. (DEBU, O. Lonsdale det., K. N. Barber rev.). Redescription. Male. Total body length 2.22–2.94 mm; general colour blackish brown, relatively shiny despite sparse greyish microtomentum, only extremities and head partly yellow or ochreous (Fig. 33). Head distinctly higher than long (Fig. 35); dorsal part of occiput slightly concave. Frons moderately broad, blackish brown posteriorly but becoming gradually paler anteriorly, orange ochreous in anterior third (lightest medially behind lunule), microtomentose and dull except for basal scars of ors and spots lateral to ocellar triangle. Orbits whitish microtomentose. Frontal triangle short, at most reaching half of frons. Frontal lunule orange, well developed. Occiput entirely blackish brown, shiny despite sparse greyish microtomentum, best developed in 2 bands behind ocellar triangle. Face (preafrons), parafacialia and gena whitish yellow to dark yellow, white microtomentose except for narrow and darker (pale brown) margin bordering parafacialia and gena; postgena orange ochreous ventrally, brown dorsally; mouthparts dark yellow. Cephalic chaetotaxy: pvt unusually long, strongly convergent or crossed; vte, vti and oc longest of cephalic setae, subequal or vte sometimes slightly longer than vti; 3 strong but relatively short ors, becoming shorter anteriorly, and 1 inclinate orbital microsetula in front of the foremost ors; 3–4 medial pairs of proclinate microsetulae in anterior half of frons; postocular setulae well developed but not longer than peristomals, in a single row; postgena with 1 longer ventral seta (as usual) and several shorter setulae; vi rather short (as long as anterior ors); subvibrissa reduced, unrecognizable from peristomal setulae that are all directed anterolaterally. Eye broadly suboval with longest diameter very slightly oblique and 1.2–1.3 times as long as shortest. Gena unusually high, with smallest height 0.20–0.23 times as long as shortest eye diameter. Palpus slender but clavate, with several black setulae ventrally, the subapical of which is distinctly longer. Antenna geniculate, orange, only 1st nagellomere brown in anterior half and shortly ciliate on anterior margin. Arista brown, about 1.7 times as long as antenna, very shortly ciliate. Thorax slightly narrower than head, dark to blackish brown, relatively shining despite sparse grey microtomentum; humeral and notopleural areas duller, with denser microtomentum; scutellum microtomentose on margins but bare and more shining on disc. Thoracic chaetotaxy: 1 minute but distinct ppl; 1 hu, 2 npl (anterior longer); 1 sa and 1 pa (longer) strong but not long; 1 prs, well developed, as long as hu or longer; 2 dc (both postsutural), anterior strong but markedly shorter than the very long posterior dc; a number (more than 5) of dc microsetae in front of anterior dc; ac microsetae nne, in 4 rows at suture, in 2 behind anterior dc, not reaching behind posterior dc; 2 sc, laterobasal as long as or slightly shorter than anterior dc, apical as long as posterior dc; 2 stpl (anterior markedly shorter) and 1 microseta in front of anterior stpl; 1–2 other microsetae in dorsal half of sternopleuron but more and longer setae ventrally. Scutellum relatively long, rounded triangular, with convex disc. Legs variegated, yellow and brown. Coxae ochreous yellow, trochanters paler. f 1 brown with yellow base and apex; t 1 dark brown except for yellow knee; fore tarsus with pale brown basitarsus, 2nd and 3rd segment yellowish ochreous, two apical segments brown. f 2 and f 3 pale ochreous basally and gradually darkened towards distal end but knee yellow; t 2 and t 3 brown but usually lighter than t 1 and with both ends yellow; mid and hind tarsi yellow, with 2 terminal segments brown (or apical segment dark brown). f 1 with ctenidial spine shorter than maximum width of t 1 and with posterodorsal and posteroventral setae thin and relatively short. t 2 with usual ventroapical seta. f 2 simply setulose. f 3 with posteroventral row of setae, 8–10 in apical two-thirds shortened and thickened. t 3 and hind basitarsus simply, nnely setulose. Wing (Fig. 34) long and relatively broad, with membrane and veins pale ochreous. C with minute spinulae not longer than other nne setulae between apices of R 1 and R 2+3. R 2+3 very long, bent parallel to C and ending usually nearer apex of R 4+5 than does M. R 4+5 very slightly bent; M long and almost straight. Discal cell (dm) medium-sized, strongly widened distally; r-m situated distally to middle of dm cell. CuA 1 slightly bent and ending near, A 1 ending far, from wing margin. Anal lobe large; alula also well developed, relatively broad. Wing measurements: length 2.38–2.82 mm, width 0.79–0.99 mm, Cs 3: Cs 4 = 0.76–1.00, r-m\ dm-cu: dm-cu = 1.67–2.09. Haltere yellow, with large pale yellow knob. Abdomen dark to blackish brown; terga more shining; sterna also dark brown but more densely greyish brown microtomentose and duller. Also T1 and T2 more microtomentose and less lustrous than other preabdominal terga. T3–T5 large, subequal in length and reaching far on ventral side of abdomen, all shortly setose. Preabdominal sterna narrow, becoming wider posteriorly, S5 largest and widest posteriorly. Postabdomen strongly sclerotized, shining blackish brown; T6 well developed, well sclerotized, forming brown, bare, transversely band-like sclerite. S6, S7 and S8 partly fused. S6 almost as long as S7, dark-margined anteriorly and usually both with 2 small setae; S8 long (as typical for Fungomyza), situated dorsally and more setose on posterior half. Genitalia. Epandrium (Figs 36, 38) dark brown, higher and wider than long, with more and stronger setae than in F. albimana, 3 pairs of laterodorsal setae longer and thicker than others. Anal nssure small, roundly triangular (Fig. 36); cerci below it also relatively small, each with a number of rather small setae including 2 longer ones and with microtomentum restricted only to posterior side (cercus laterally without microtomentum). Medandrium high, broad ventrally and strongly narrowed dorsally (Fig. 36) although less than in F. albimana. Gonostylus (Figs 36–38) elongate suboblong but with strongly projecting and incurved posteroventral corner bearing 2 small teeth on apex and with microtomentum restricted to a central area of outer side and setae mainly arising on inner side at posterior margin. Hypandrium (Fig. 38) relatively robust, well sclerotized but without anterior nat, dorsally projecting lobes. Pregonite (Fig. 38) large, nat, fused to hypandrium, ventrally thickened and dark but, in contrast to F. albimana, without posterior projection, and with 3 setae in anterior and 2 in posterior group. Postgonite (Fig. 38) simple, slender, pale, slightly bent, distinctly shorter than in F. albimana, with anterior setula in distal third, several grain-like sensillae on outer side and blunt apex. Transandrium well sclerotized, straight medially, with dorsally bent lateral parts and ventromedially carrying distinct (though pale-pigmented) forked caudal process (Fig. 40); basal membrane below caudal process with numerous dense short spines. Aedeagal complex (Fig. 41) with short phallophore having somewhat projecting anteroventral corner. Distiphallus bind and large, composed of largely membranous saccus and heavily sclerotized nlum. Saccus proximally sclerotized but these sclerites asymmetrical, enlarged distally on right side; membranous part of saccus relatively small and unarmed, without distinct spines or setulae. Filum formed by 2 dark band-like sclerites being closely attached in the middle and fused apically; apex of nlum (Fig. 39) slender, nattened and pointed but structurally similar to that of F. albimana. Aedeagal part of folding apparatus (Fig. 41) provided with elongate striae and grain-like tubercles in outer wall. Connecting sclerite (Fig. 41, cs) poorly delimited and weakly sclerotized. Phallapodeme (Figs 38, 41) medium-sized, similar to that of F. albimana, with shortly forked base, robust apical part and large ventral fulcrum. Ejacapodeme (Fig. 41) small, suboblong, with slender ventral projection lying in wall of ejaculatory duct. Female. Similar to male unless mentioned otherwise. Total body length 2.58–3.18 mm. Face, parafacialia and gena darker, orange yellow to orange ochreous; clypeus darker, brown. Antenna sometimes (North Carolina: Raleigh) more extensively darkened on the lateral surface, reaching ventral margin in apical half. Legs less distinctly variegated. Ctenidial spine on f 1 slightly shorter than in male. f 3 without posteroventral thickened setae. Wing measurements: length 2.58–3.30 mm, width 0.87–1.17 mm, Cs 3: Cs 4 = 0.81–0.97, r-m\dm-cu: dm-cu = 1.53–2.19. Preabdominal terga broader and more transverse, far reaching on ventral side of abdomen. T2–T4 shortly setose; T5 somewhat tapered posteriorly and with distinctly longer setae at posterior margin. Preabdominal sterna slightly narrower than in male, with short and nne setae. Postabdomen (Figs 42, 43) telescopically retractible and strongly tapered caudally. T6 broadly transverse but very tapered posteriorly, dark, shortly setose. S6 narrow, distinctly longer than broad, setose in posterior half, relatively dark. 6th spiracle situated at anterior corner of T6 (Fig. 43). T7 not fused with S7, markedly narrower than T6 and conically narrowed posteriorly, dark and heavily sclerotized, with longer setae only at posterior margin. S7 as long as S6 but distinctly narrower, with longer and denser microtomentum and long nne setae (more than in F. albimana) on posterior half, brown except for pale posterior fourth. 7th spiracle situated in front of anterior corner of T7. Intersegmental membrane (also dorsally) between 7th and 8th segment with long distinct pubescence. T8 (Fig. 42) markedly shorter than in F. albimana, distinctly transverse, lighter than T7, without microtomentum and nnely setose. S8 (Fig. 43) more or less divided medially, posteriorly curved dorsally and deeply invaginated into 8th segment (see also Fig. 44), strongly microtomentose apart from bare invaginated posterodorsal parts. Internal structures of the female genital chamber (uterus) better sclerotized than in F. albimana, with 2 pairs of partly fused posterior sclerites attached to invaginated parts of S8 (Figs 44, 45) and 1 poorly visible (seemingly incomplete), unpigmented, anterior looped structure; ducts of accessory glands slender, transversely ringed; accessory gland small; ventral receptacle (Fig. 44) large, hyaline membranous, elongate vesicular and similar to that of F. albimana except for terminal projection being short and blunt apically. Spermathecae (1+1) cup-like (not pyriform), with narrower base covered with nne spinulae terminated with usual stalked globuli; distal part broader and provided with broad, deep apical invagination (Fig. 46); spermathecal ducts short, without terminal cervix, hyaline near insertion into body of spermatheca. T10 (Fig. 42) narrowly pentagonal with somewhat bulging sides, and bare apart from usual pair of dorsal setae; apical portion depigmented leaving pigmented basal two-thirds medially and narrowly emarginate between bulging sides to near level of setal pair. S10 (Fig. 43) larger than T10, relatively nat with doubly emarginate anterior margin, also bare except for setae and sparse microtomentum at posterior margin. Cerci long and slender (more robust than in F. albimana), with rich setae that are longest apically. Discussion. Fungomyza buccata is the only Nearctic representative of Fungomyza. It closely resembles the W. Palaearctic F. albimana both as regards structures of the male genitalia (eg. form of epandrium, small cercus, dorsally tapered medandrium, robust pregonite, simple slender postgonite, forked caudal process of transandrium, relatively small saccus; similar nlum of distiphallus and ejacapodeme) and the female postabdomen (eg. T6, T7, S6, S7, S8, intersegmental pubescence, T10, ventral receptacle, terminal invagination of spermatheca). Most of these shared features are synapomorphic and four of them were used to demonstrate the sister-species relationships of these taxa, see ROHÁĆEK (2009a: Fig. 139). Fungomyza buccata differs distinctly from F. albimana in a number of external and genitalic features, however, including the following: anterior ors longer; frontal triangle short; eye less oval; gena unusually high (probably the highest in all described Anthomyzidae); legs differently variegated (e.g. f 1 brown except for both ends, 2 apical segments of all tarsi darkened, f 2 and f 3 darkened in distal half, etc.); gonostylus with projecting posteroventral corner and reduced microtomentum; pregonite without posterior projection; saccus of distiphallus without spinulae or setulae; nlum shorter, with narrower apex; female T8 short and transverse; internal sclerites of female genital chamber distinct and spermathecae more rounded. It is also more robust and has longer wings on the average than F. albimana. It is to be remarked that an additional species, Reliquantha variipes from Great Britain, is externally strikingly similar to F. buccata (including the variegated legs and cephalic chaetotaxy). It differs from F. buccata very distinctly in the male and female postabdominal structures (see ROHÁĆEK 2013c) and also in lacking the ctenidial spine on f 1 and in having a well-developed subvibrissa, short prs and only 2 setae on the ventral corner of the sternopleuron. Biology. A male specimen collected by Steve Marshall in Florida was caught on the sporocarp of a white fungus (Fig. 33) growing on a well-rotted, partially buried log in a sandy, partly wooded, habitat. It is therefore probable that F. buccata also develops in fungi like its European sister species. Adults were mainly obtained in autumn (in October – 5 specimens, November – 4 specimens, December – 4 specimens); only one female was captured in May. Together this suggests an association with sporocarps of fungi which mainly grow in autumn (ROHÁĆEK & BARBER 2004). Distribution. This uncommon species is widespread in the eastern United States of America: Alabama (new record), District of Columbia, Florida, Georgia, Maryland, New York, North Carolina, South Carolina, Virginia (ROHÁĆEK & BARBER 2004, and Table 2, Figs 600, 603).Published as part of Roháćek, Jindřich & Barber, Kevin N., 2016, Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera), pp. 1-412 in Acta Entomologica Musei Nationalis Pragae (suppl.) (suppl.) 56 on pages 33-36, DOI: 10.5281/zenodo.427282
Anthomyza pullinotum Roháćek & Barber 2016, sp. nov.
Anthomyza pullinotum sp. nov. (Figs 276, 296–312, 378) Type material. HOLOTYPE: ♂, “CAN:AB: ~22.7kmS Belle-vue, Hwy 774, 17.vii.2011, KNBarber, sweeps, road-side ditch, mostly Carex spp., Equisetum, grasses 49°22.62’N 114°22.58’W ” and “ Holotypus ♂ Anthomyza pullinotum sp. n., J. Roháček & K. N. Barber det. 2014” (red). The specimen is in good condition, with exposed genitalia (saccus and nlum visible on right side) and readily observable gonostyli (see Figs 296, 298) (DEBU, intact). PARATYPES: CANADA: ALBERTA: Banff N. P., 11 mi W Banff, 4500', 11.vii.1955, 1 ♂, G. E. Shewell leg. (CNCI); Banff Nat. Pk., 15 mi E Mt. Eisenhower Jct., 27.vii.1967, 1 ♂, S. P. Whitney leg. (USNM, genit. prep.); ~ 22.7 km S Bellevue, Hwy 774, 49°22.62'N 114°22.58'W, sweeps, roadside ditch, mostly Carex spp., Equisetum, grasses, 17.vii.2011, 10 ♂♂ 8 ♀♀, K. N. Barber leg. (CNCI 6 ♂♂ 4 ♀♀, SMOC 2 ♂♂ 2 ♀♀, USNM 2 ♂♂ 2 ♀♀); Calgary, Fish Creek Prov. Pk., 50°55.61'N 114°07.43'W, sweeps, mostly Carex utriculata & Equisetum ssuviatile, 12.vii.2011, 1 ♀, J. E. Swann & K. N. Barber leg.; same locality but 50°55.600'N 114°07.426'W, sweep, oxbow with sedges, 9.vii.2011, 1 ♂; same locality but 50°55.739'N 114°03.312'W, sweep near creek, 22.vii.2010, 1 ♂; same locality but pond near Shannon Terrace, 50°55.600'N 114°07.427'W, swept from sedges and Equisetum, 12.viii.2011, 1 ♂; same locality but Shannon Terrace, sweep around pond by 2 nd bridge, 9.vii.2010, 7 ♂♂ 1 ♀, all J. E. Swann leg. (all BDUC); Cypress Hills, Elkwater, trails, 49°39'24"N 110°17'32"W, 1250 m, 5.vii.2001, 2 ♀♀ (DEBU 00354151–52); Cypress Hills Prov. Pk., Elkwater, 5.vii.2001, 1 ♀ (DEBU 00362459), all S. A. Marshall leg.; w. border Elk Island N. P., Range Rd. 210, 0.5 km N Hwy #16, 53°34.52'N 112°57.09'W, sweeps, mixed sedges, 21.vii.2008, 2 ♀♀, sweeps, Calamagrostis canadensis, 21.vii.2008, 1 ♀, K. N. Barber leg.; Elkwater Lk., 19.vii.1956, 1 ♂, O. Peck leg. (all CNCI); ~ 22.5 km NW Highwood House, ~ 4 km W Mist Ck., 50°31.38'N 114°53.17'W, sweeps, Carex sp. (large), 25.vii.2008, 1 ♂, K. N. Barber leg. (DEBU); ~ 3.4 km SSW Hinton, Hwy #40, 53°21.27'N 117°37.32'W, sweeps, Equisetum ssuviatile, 22.vii.2008, 1 ♂; ~ 11 km WSW Hinton, nr. jct. Twp. Rd. 510A & Range Rd. 262, 53°22.72'N 117°44.13'W, sweeps, mostly Bromus inermis, 22.vii.2008, 1 ♂ 1 ♀, all K. N. Barber leg. (all CNCI); 2 mi S Jasper, 30.vii.1967, 1 ♂, S. P. Whitney leg. (USNM, genit. prep.); Kananaskis Country, Sibbald Area, Hwy 68, 3.1 km W Powderface Trail, 51°03.10'N 114°54.72'W, sweeps, mostly Carex utriculata, 15.vii.2011, 1 ♂ 1 ♀; same locality but Hwy 68 & Powderface Trail, 51°02.28'N 114°52.40'W, sweeps, mostly Carex utriculata ?, 15.vii.2011, 1 ♂, K. N. Barber (all CNCI); ~ 31.5 km S Kananaskis Village, 0.5 km W Little Highwood Pass, 50°38.55'N 115°02.94'W, sweeps, Bromus inermis, 25.vii.2008, 1 ♀, K. N. Barber leg. (genit. prep.); 50 mi N Nordegg, F[orestry] Trunk Rd., along beaver pond, 20.vii.1987, 1 ♂ 2 ♀♀ (incl. pair in copula), S. A. Marshall leg. (all DEBU); ~ 14.4 km E Obed, Range Rd. 213 @ RR crossing, 53°32.19'N 117°01.02'W, sweeps, mostly Carex utriculata, 25.vii.2011, 4 ♂♂ 2 ♀♀, K. N. Barber leg. (CNCI); Spray Valley P. Pk., 50°48.95'N 115°09.84'W, sweeps, fen, Carex utriculata ? and Poa sp., 13.vii.2011, 1 ♂, K. N. Barber leg. (DEBU 01502856). BRITISH COLUMBIA: Fernie,Annex Pk., 49°30.72'N 115°04.13'W, sweeps, wet ditch, Carex utriculata, 17.vii.2011, 1 ♂, K. N. Barber leg. (CNCI); Kaslo Cr., 18.vi. [-], 1 ♂, R. P.Currie leg. (USNM, genit. prep.); ~8.0 km SE Valemount, edge of Kinbasket Lake, 52°46.65'N 119°10.38'W, sweeps, mostly Carex utriculata, 23.vii.2011, 1 ♀, K. N. Barber leg.; Lac Le Jeune, 27.vi.1973, 1 ♂, H. J. Teskey leg.; Liard Hot Spring, mi. 496, 1500', Alaska Hwy, 9–10.vii.1959, 1 ♀, E. E. MacDougall leg. (all CNCI); Mt. Robson Prov. Pk., Hwy #16, small road towards Mt. Robson, 53°03'N 119°15'W, forest noor, swamp, (Universität Bielefeld, Ca1519), 6.viii.2002, 1 ♂, M. v. Tschirnhaus leg. (ZSMC, in ethanol); Revelstoke, 2.vii.1973, 1 ♀, H. J. Teskey leg.; Sawmill Lk., Telegraph Ck., 1100', 2.vii.1960, 2 ♂♂ 2 ♀♀, W. W. Moss leg., Carex, grass, Equisetum beside lake, 2.vii.1960, 1 ♂, 28.viii.1960, 1 ♂, R.Pilfrey leg.; ~ 27 km N Sparwood, Lower Elk Valley Rd., 49°50.24'N 114°53.29'W, sweeps, edge of creek, Carex utriculata ?, 20.vii.2011, 1 ♀, K. N. Barber leg.; Summit Lake, mi. 392 Alaska Hwy, 4500', 8.vii.1959, 1 ♀, E. E. MacDougall leg.; Spring Creek, Terrace, 220', 11.vi.1960, 1 ♂, R. Pilfrey leg. (all CNCI). LABRADOR: Goose Bay, 9.vii.1948, 1 ♂, 13.vii.1948, 1 ♀, W. E. Beckel leg., 11.vii.1950, 1 ♀, J. J. Tibbles leg. (CNCI). ONTARIO: Moosonee, 51.27717°N 80.64778°W, Repl. 3 wet, Malaise trap, 19–22.vi.2010, 1 ♀, NBP Field Party leg. (LEMQ). YUKON: Klondike Hwy, 8.8 km S Twin Lakes, Conglomerate Mt., 61°37.9'N 135°53.1'W, sweep along Klusha Creek, 15.vii.1998, 1 ♂; same locality but Conglomerate Mt., Klusha Creek, 61°38'N 135°53'W, sweep grass/sedges along creek, 8.vii.1997, 1 ♂ both (LEMQ); Alaska Highway at Yukon River crossing, 60°34'N 134°40'W, sweep grass/sedges along river margin, 2.vii.1997, 13 ♂♂ 6 ♀♀ (LEMQ 0039623 [only 1 ♂ with accession #], 1 ♂ wing illustration, 2 ♂♂ 1 ♀ genit. prep.); Yukon River at Alaska Hwy crossing, sweep grass/sedges along riverbank, 2.vii.1997, 1 ♀ (LEMQ 0039961), all T. A. Wheeler leg. UNITED STATES OF AMERICA: ALASKA: Knik Lake, NW of Wasilla, sweeping vegetation edge of lake, 18.vii.1978, 7 ♂♂ 1 ♀, P. H. Arnaud Jr. leg. (CASC, 1 ♂ genit. prep.). COLORADO: Electra Lake, F.4367E, ~ 37°33'N 107°48'W, ~8400', 28.vi.–1.vii.1919, 2 ♀♀, [no collector] (AMNH, 1 ♀ genit. prep.); Estes Park, 11.vii.1934, 1 ♂, A. L. Melander leg. (USNM, genit. prep.); Teller Co., 3.5 mi S Florissant, Sanborn Ranch, along tributary of Plum Creek, 14.vii.2004, 1 ♂ 1 ♀, The Nature Place, sweeping forbs, 10.vii.2004, 1 ♀, B. A. Foote leg. (CMNH); Summit Co., Frisco, 3.viii.2001, 2 ♂♂, I. S. Winkler leg. (BYUC); Jackson Co., Gould, 8.viii.1965, 1 ♀, F. C. Harmston leg. (LACM, genit. prep.); Boulder County, Middle Boulder Creek, 16 km W Boulder, Hwy 119, 2280 m, 8.viii.1973, 2 ♂♂ 2 ♀♀, P. H. Arnaud Jr. leg. (CASC, 1 ♂ genit. prep.); 3 mi N Nederland, 8500', marshy stream margin, 2.vii.1961, 1 ♀; 5 mi E Nederland, 7500', marshy lake & stream margin, 2.vii.1961, 1 ♂, both J. G. Chillcott leg.; State Bridge nr. Bond, 7000', 24–25.vi.1961, 1 ♂, C. H. Mann leg. (all CNCI). MICHIGAN: Keweenaw Co., Isle Royale, 15., 17.vii.1938, 1 ♂ 1 ♀, G. Steyskal leg. (USNM, 1 ♂ genit. prep.). NEW HAMPSHIRE: Pinkham Notch, 9.vii.1931, 1 ♀, J. M. Aldrich leg. (USNM). Description. Male. Total body length 2.48–2.90 mm; body distinctively bicolourous (Figs 296–298), with occiput (partly) and dorsal sides of thorax and abdomen largely brown to dark greyish brown (notum in particular), sharply contrasting with yellow lateral and ventral sides of thorax and abdomen, most of head, and all extremities (including antenna and palpus). Head about as long as high, anteriorly slightly angular in pronle, with face slightly receding, largely yellow except for brownish ocellar triangle and darker brown occipital pattern. Occiput very slightly concave, bicolourous, laterally with large dark brown subtriangular area extended from posterior eye margin to foramen and medially with small pale brown spot behind ocellar triangle; lateroventral parts of occiput and medial V-shaped area above foramen yellow (the latter dorsally transilient to yellow parts of frons and largely covered with 2 silvery microtomentose spots meeting above foramen). Frons relatively narrow, yellow and largely dull, only ocellar triangle brown; frontal triangle with sparse but distinct silvery golden glittering microtomentum as in A. mcalpinei. Also orbits as in latter species, with silvery whitish microtomentum reduced behind middle ors to form a narrow line. Frontal triangle very narrow and with attenuated and acutely pointed anterior corner reaching to anterior fourth to nfth of frons. Frontal lunule small but distinct, yellow. Face as in that of A. mcalpinei but separated from parafacialia by broader and usually darker, golden-orange microtomentose marginal stripe reaching onto ventral margin of gena; parafacialia, gena, postgena and mouthparts coloured and microtomentose as in A. mcalpinei. Cephalic chaetotaxy not particularly different from that of A. mcalpinei including dark colour and variability in length of setae; only postocular setulae (6–7) somewhat longer, particularly those situated more dorsally. Palpus as described for A. mcalpinei including chaetotaxy. Eye also very similar to that of A. mcalpinei, with longest diameter oblique and 1.3–1.4 times as long as shortest. Shortest genal height 0.16–0.18 times as long as shortest eye diameter, thus gena somewhat deeper (higher) than that of A. mcalpinei. Antenna geniculate, entirely yellow; 1st nagellomere with short white pilosity. Arista about 1.9 times as long as antenna; basal segments ochreous yellow, distal setiform part blackish brown and with (dark) cilia shorter than those on 1st nagellomere as in A. mcalpinei. Thorax slightly narrower than head. Scutum dorsally invariably dark greyish brown with dense pale grey microtomentum with distinct bluish tinge (Fig. 297). Humeral callus and notopleural area yellow (but the latter usually with ochreous to pale brown darkening around anterior npl), often with yellow or ochreous colour extended along suture up to level of prs, and sometimes also with short yellow stripe between sa and pa. Scutellum also brown and similarly microtomentose to scutum but sometimes (particularly medially) paler brown. Pleural part of thorax sparsely whitish grey microtomentose, more shining than scutum, usually entirely bright yellow (Fig. 297), rarely with very narrow brownish marginal darkening dorsally on mesopleuron and propleuron. Postscutellum and postnotum brown to dark brown. Thoracic chaetotaxy: 1 hu (usually longer than posterior npl) plus 1 (rarely 2) hu setula on humeral callus; 2 npl (anterior distinctly longer than hu); 1 distinct prs (as long as or longer than hu); 1 sa (as long as prs) and 1 pa (usually longer than sa); 2 long postsutural dc (anterior about as long as anterior npl, posterior longest of thoracic setae) and 6–8 dc microsetae in front of them (the hindmost distinctly enlarged); ac microsetae more sparse than in A. mcalpinei, arranged in only 2 medial rows but with 1–2 lateral ac microsetae behind suture in addition; hindmost (medial) ac pair usually situated somewhat beyond level of posterior dc; 2 sc (shorter laterobasal about as long as sa, apical almost as long as posterior dc); 1 small hair-like ppl (exceptionally 2, observed in single specimen); 2 long stpl (anterior shorter) and 4–5 upcurved setulae in dorsal half of sternopleuron, its ventral part with 5–6 longer setae. Scutellum rounded triangular, slightly convex dorsally. Legs pale yellow, only distal half to three-fourths of last tarsal segment of all tarsi dark brown. Pedal chaetotaxy very similar to that in A. mcalpinei: f 1 with ctenidial spine only slightly longer than maximum width of t 1; f 3 in distal two-nfths with 5–7 shortened and thickened setae in posteroventral row; t 2 with short ventroapical seta; fore and hind basitarsus with 2–3 enlarged (also thickened on hind basitarsus) ventrobasal setulae, also mid basitarsus with 1 or 2 somewhat longer and thicker ventrobasal setulae. Wing (Fig. 276) less elongate (somewhat wider) than that of A. mcalpinei, having pale yellowish ochreous veins and membrane. C with distinct sparse spinulae (not well visible on Fig. 276 because erect and perpendicular to alar plane) among hair-like setosity between apices of R 1 and R 2+3. R 2+3 long, bent parallel to C with apex slightly upcurved to it; R 4+5 very slightly bent, subparallel with almost straight or indistinctly bent M. Discal cell (dm) moderately long and narrow; r-m situated slightly in front of the middle of cell dm. Apical portion of CuA 1 longer than dm-cu and ending near wing margin; A 1 short, ending far from it. Alula distinct, moderately narrow. Wing measurements: length 2.56–2.90 mm, width 0.81–0.99 mm, Cs 3: Cs 4 = 0.86–1.25, rm\dm-cu: dm-cu = 2.63–3.17. Haltere yellowish white with stem more yellow. Abdomen dorsally largely brown, ventrally pale to whitish yellow (Fig. 298). Preabdominal terga T1–T5 mostly brown but their side areas variously yellow; T2–T3 usually darkest with only margins yellow, T1 and T4–T5 (particularly) usually with lateral yellow regions larger, covering up to one third on each side. T1–T5 relatively shortly and sparsely setose, subshining, with greyish microtomentum distinct but sparser than that on thorax. T1 and T2 separate, only narrowly fused laterally. T1 shortest and most transverse, T2 slightly shorter than T3–T5, the latter subequal in size, all reaching onto lateroventral sides of abdomen. Preabdominal sterna pale to whitish yellow, relatively broad (only S1 and S2 narrower) and becoming wider posteriorly; S1 short and transverse, S2 slightly, S3–S5 distinctly transverse, all suboblong to slightly trapezoidal (wider posteriorly), S5 widest. S2–S5 nnely but not very densely setose, only S1 bare and with darker posterior marginal stripe. T6 submembranous as in A. mcalpinei, very short, transversely strip-like, bare and almost unpigmented. S6 and S7 pale brown (usually) to brown, often with central part lighter (up to ochreous yellow), both with dark brown anterior marginal ledge; both S6 and S7 with 2–3 (usually 2) setae; S8 somewhat longer than epandrium, brown (usually darker than S6 and S7 but paler than epandrium), setose in posterior two-thirds. Genitalia. Epandrium (Figs 299, 300) darker than S8 (Fig. 298), moderately long and relatively broad as in A. mcalpinei, but slightly less setose, with 2 or 3 pairs of longer and thicker setae dorsolaterally; anal nssure relatively small but more acutely triangular than that of A. mcalpinei. Cercus relatively short and pale, with nne setae, apical and preapical longest. Cerci (Fig. 299) distinctly closer to each other than in A. mcalpinei. Medandrium (Fig. 299) simple, slightly smaller (lower) than in A. mcalpinei, dorsally slightly narrowed and with dorsolateral corners simple, obtuse-angled, ventrally with shallow broad emargination, bare. Gonostylus (Figs 299, 300, 303) elongate and slender but shorter than epandrial height, bent anteriorly (with distinct anterior concavity in lateral view – Figs 298, 300), distally gradually tapered and apically somewhat pointed, with 1 short apical and 1 small subapical tooth (Fig. 303), thus more resembling that of the Palaearctic A. dissors but distinctly different from that of A. mcalpinei although similarly micropubescent on most of outer side and with longer setae only on inner side. Hypandrium (Fig. 302) resembling that of A. mcalpinei but somewhat more robust. Transandrium (Fig. 301) also as in the latter species although more arched dorsally, without caudal process except for a pair of short medial sclerotizations transilient to nnely spinose parts of basal membrane. Pregonite (Fig. 302) most similar to that of A. mcalpinei but with anterior tooth more slender and acutely pointed, ventrally with 5 (2 middle usually longest) setae. Postgonite (Fig. 302) clearly different from that of A. mcalpinei, short and unusually broad, widest preapically (in lateral view), with wide rounded apex, dark basally, pale apically, with 1 setula in basal third to fourth of anterior margin and several sensillae on outer side. Dorsal internal sclerite at base of postgonite distinct, narrower than, and almost as long as postgonite. Basal membrane (Figs 301, 302) with the same nne armature as in A. mcalpinei, but spinulae on posterolateral areas smaller. Aedeagal part of folding apparatus with different structure from those of A. mcalpinei and A. dissors, laterally with elongate group of nne dark tubercles in addition to elongate hyaline striae. Connecting sclerite strong, dark, proximally fused to phallophore, distally slightly wider and with a few (in contrast to that of A. mcalpinei) spines on apex (Fig. 304); membrane posterior to connecting sclerite with only nne unpigmented spinulae and nat warts. Phallapodeme very similar to that of A. mcalpinei including position and shape of fulcrum and laterally projecting apex (Fig. 304). Aedeagus (Fig. 304) with phallophore short and compact; distiphallus large and long, bind from near its base. Saccus more voluminous and longer than in A. mcalpinei and A. dissors, often more dilated distally, membranous, basally with usual slender sclerites plus one short lateral sclerite and armed with 8 robust dark-pigmented spines (thus more than in A. mcalpinei). Filum most resembling that of A. mcalpinei but its curved slender terminal part distal to small subterminal membranous lobe with fewer spinulae (Fig. 305). Ejacapodeme small, pale, with small pointed terminal projection (Fig. 304). Female. Similar to male unless mentioned otherwise. Total body length 3.09–3.73 mm. Head with occiput sometimes yellower, with lateral brown areas interrupted by a yellow stripe connecting orbit with lateroventral area of occiput. Frons with up to 4 pairs of medial microsetulae in front of frontal triangle. Outer side of 1st nagellomere with large darker (ochreous to pale brown) anterodorsal spot below insertion of arista, also dorsal margin of inner side sometimes faintly darkened. Thorax often paler than in male because yellow lateral areas extend dorsally or even onto posterior nfth of scutum to form small yellow spots in prescutellar area; scutellum also sometimes partly (usually laterally) yellow to (rarely) completely ochreous yellow. Largest female specimens with more ac microsetae (forming 4 rows also presuturally) and with more setose sternopleuron (with up to 6 upcurved setulae and 7 ventral setae). f 3 without row of shortened and thickened posteroventral setae. Wing generally more elongate. Wing measurements: length 3.07–3.67 mm, width 0.99–1.19 mm, Cs 3: Cs 4 = 0.87–1.11, rm\ dm-cu: dm-cu = 2.29–3.20. Abdomen with T1–T6 distinctly lighter, brownish only medially but these darker areas often reduced to small medial spots or sometimes disappear to leave terga completely yellow. T2–T5 shorter and more transverse than in male, T1 distinctly, T2 slightly narrower than T3. T3–T5 broad and subequal in size, all wider than T6. Preabdominal sterna pale yellow, somewhat more densely setose and slightly narrower than in male. S2 as long as wide, S3 slightly, S4 and S5 distinctly transversely suboblong. S5 largest and widest abdominal sternum, being slightly wider than S6. Postabdomen (Figs 306, 308) relatively long (more elongate than in A. mcalpinei), telescopic, yellow with brown markings. T6 simple, large, narrower than T5 and much longer than that of A. mcalpinei, slightly tapered posteriorly and with broadly rounded posterior corners, dark yellow or with medial brownish area of various size, with relatively short and dense setae in posterior two-thirds (dorsomedially in only posterior half), marginal setae longest. S6 also relatively narrow (less transverse than in A. mcalpinei), slightly trapezoidal with anterior corners rounded, less broad than S5, pale yellow and nnely densely setose. Tergosternum T7+S7 relatively long, subcylindrical, narrowing slightly posteriorly, dorsomedially (Fig. 306) shortened due to anteromedial incision (shallower than that seen in A. mcalpinei), ventrally longer but without anteroventral pouch-like structures (Fig. 308). T7+S7 with variable yellow and brown pattern, dorsally with brown posteromedial area that is variable in size, ventrolateral region often with brown area that becomes darker anteroventrally; ventral part of T7+S7 with original S7 well discernible but fused anteriorly with dilated anteroventral lobes of T7, relatively dark but with paler elongate area medially, with nne setae and sparse micropubescence (Fig. 308); dorsal and lateral parts of T7+S7 with rather short setae, those dorsally distinctly shorter and thicker (Figs 306, 378); 7th spiracle embedded in expanded lateral part of original T7 (Figs 308, 378). 8th segment nnely micropubescent laterally. T8 (Fig. 306) about as long as wide, with sides bent ventrally, brownish, with deep emargination anteromedially and with unpigmented crescent-shaped marginal area posteromedially (hence different from that of A. mcalpinei), with sparse micropubescence centrally and nne exclinate setae laterodorsally; S8 (Fig. 308) shorter than T8, anteriorly strongly tapered, medially divided into 2 posteriorly convex (dorsally bent), nnely hirsute and micropubescent sclerites that have a rather cordate shape. Genital chamber (uterus) posteriorly with distinctive darkpigmented internal sclerotization (Figs 309, 311, 312) formed by 1 compact ventral sclerite (different from that of A. mcalpinei because convex ventrally), a complex (doubled) pair of weak, elongate and pale-pigmented dorsal sclerites and 1 subcircular, curved (in pronle) annular sclerite situated in front of the former. Membranous pa
Anthomyza oblonga Roháćek & Barber 2016, sp. nov.
Anthomyza oblonga sp. nov. (Figs 206, 223, 225–238) Type material. HOLOTYPE: ♂, “CAN:ON: SSMarie, Bristol Pl.Pk., 04.vii.2008, KNBar-ber, sweeps, Impatiens, Clematis, Equisetum, Rub-us, ferns, Phalaris 46°30.77’N 84°16.66’W ” and “ Holotypus ♂ Anthomyza oblonga sp. n., J. Roháček & K. N. Barber det. 2013” (red). The specimen is in perfect condition, with highly visible, partly exposed genitalia (see Fig. 206) (CNCI, intact). PARATYPES: CANADA: MANITOBA: Ninette, 13.vi.1958, 1 ♀, C. D. F. Miller leg.(CNCI); Ninette, ex. Rudbeckia laciniota, 3.vi.1958, 1 ♂, maple/elm noodplain community, 12.vi.1958, 2 ♂♂ 2 ♀♀, J. F. McAlpine leg. (CNCI). NEWFOUNDLAND: Port aux Basques, 6.viii.1961, 1 ♀, C. P.Alexander leg. (USNM, 1 ♀ genit. prep.). NOVA SCOTIA: C[ape] B[reton] H[ighlands] N. P., Beulach Ban Falls, PG812870, swept along fast rocky stream, 8.vii.1983, 2 ♀♀; Truro, 14.vii.1983, 1 ♀, all J. R. Vockeroth leg. (all CNCI). ONTARIO: Algonquin, mixed wood, 1.vi.1991, 1 ♂, M. Barták leg.(MBPC, genit. prep.); Cootes Paradise nr. Dundas, sweeping undergrowth of deciduous forest, 20.viii.1994, 17 ♂♂ 9 ♀♀, J. Roháček leg. (SMOC 14 ♂♂ 8 ♀♀, 3 ♂♂ 3 ♀♀ genit prep., NMPC 3 ♂♂ 1 ♀); Dubreuilville, 48°21.05'N 84°33.84'W, sweeping Diervilla, ferns, Clintonia, Cornus, Aralia, Eurybia, Vaccinium under Populus / Pinus, 10.vii.2010, 1 ♀, J. Roháček leg. (SMOC, genit. prep.); Fergus, Malaise trap, 21.vii.1990, 1 ♂, S. A. Marshall leg. (DEBU); Fort Frances, 10 mi E on Hwy. 11, 8–9.vii. 1978, 1 ♀, H. J. Teskey leg. (CNCI); Greenwater P. Pk., Green Trail, 49°11.73'N 81°16.76'W, sweeps, Eurybia, Cornus, Clintonia, Diervilla, Aralia under Populus, 21.vii.2009, 1 ♂, K. N. Barber leg. (DEBU 01502212); 7 mi E Grifnth, 22.vi.1985, 1 ♂, B. E. Cooper leg. (CNCI); Icewater Creek WS [watershed], ~ 12.7 km NNE Searchmont, mi. 10.5 Whitman Dam Rd., alder thicket, 1.viii.1986, 1 ♀ (CNCI); Icewater Creek WS [watershed], 46°53.7'N 84°03.4'W, sweeps, Thalictrum, Eupatorium [Eutrochium], sedge, fern in mixed forest, 7.vii.1998, 6 ♂♂ 4 ♀♀ (CNCI 4 ♂♂ 2 ♀♀, 1 ♀ genit. prep., USNM 2 ♂♂ 2 ♀♀), sweeps, Thalictrum, sedge, fern, riparian mixed forest, 7.vii.1998, 3 ♂♂ 4 ♀♀ (AMNH 2 ♂♂ 2 ♀♀, CNCI 1 ♂ 2 ♀♀, 1 ♂ 1 ♀ genit. prep.), sweeps, Thalictrum, Eupatorium [Eutrochium], fern in mixed forest, 10.vii.1998, 2 ♀♀ (1 ♀ genit. prep.), sweeps, trailside sedges, ferns, grasses, 10.vii.1998, 2 ♀♀, sweeps, trailside veg. incl. sedges, ferns, grasses, 17.vii.1998, 1 ♀ (genit. prep.) (CNCI); King Mt., 26 km N S[ault] S[te.] Marie, riparian sweeps, 16.vi.1987, 1 ♀ (CNCI), all K. N. Barber leg.; Moosonee, 51.24622°N 80.67281°W, Repl. 1 mesic, Malaise trap, 18–21.vi.2010, 2 ♀♀; Moosonee, 51.24466°N 80.67767°W, Repl. 2 mesic, Malaise trap, 21–24.vi.2010, 1 ♂, NBP Field Party leg. (LEMQ); Moosonee, 51°14.75'N 80°40.33'W, sweeps, mostly Rubus, Ribes, under Populus, 9.vii.2014, 1 ♀; Moosonee, 51°14.79'N 80°40.35'W, 9.vii.2014, sweeps, mostly Solidago, Calamagrostis, 1 ♀; Moosonee, 51°16.33'N 80°39.11'W, sweeps, mostly Rubus, Impatiens, under Salix, Alnus, 10.vii.2014, 2 ♂♂ 5 ♀♀; Moosonee, 51°16.36'N 80°39.11'W, sweeps, railside ditch, mostly Equisetum ssuviatile, Carex spp., 11.vii.2014, 1 ♀, all K. N. Barber leg. (all CNCI); Ottawa, 15.vii.1957, 1 ♀ (genit. prep.), J. E. H. Martin leg., 30.vi.1963, 1 ♀, J. R. Vockeroth leg. (CNCI); S[ault] S[te.] Marie, S of Algoma U[niversity] College, 46°29.9'N 84°17.2'W, sweeps, Impatiens under Populus / Betula, 13.vii.1998, 1 ♂ 1 ♀ (♀ genit. prep.); S[ault] S[te.] Marie, Baseline Rd., 46°31.40'N 84°24.45'W, sweeps, edge of forest, Solidago, Rubus, Equisetum, grasses, 25.vi.2005, 1 ♂, all K. N. Barber leg. (all CNCI); Sault Ste. Marie, Baseline Road, 46°31.40'N 84°24.40'W, sweeping Aster [Doellingeria], Rubus, Equisetum, Carex, Clematis, ferns under aspen (Populus), 7.vii.2010, 2 ♂♂ 2 ♀♀ (2 ♂♂ 1 ♀ genit. prep.), 12.vii.2010, 1 ♂ (genit. prep.), J. Roháček leg. (SMOC); S[ault] S[te.] Marie, Baseline Rd., 46°31.40'N 84°24.40'W, Malaise #1, Aster [Doellingeria], Rubus, Equisetum, Carex, Solidago, in aspen clearing, 8–18.vii.2005, 1 ♀, sweeps, Aster [Doellingeria], Rubus, Equisetum, Carex, ferns under aspen, 25.vi.2005, 2 ♂♂, 26.vi.2005, 1 ♂ 2 ♀♀, 22.vii.2010, 1 ♀ (CNCI), 19.vi.2011, 1 ♂ 2 ♀♀ (INHS), 29.vii.2012, 2 ♀♀, sweeps, mostly ferns under aspen, 19.vi.2011, 3 ♂♂ 2 ♀♀ (LACM), 27.vii.2012, 1 ♂ 3 ♀♀ (1 ♀ genit. prep.), 28.vii.2012, 2 ♂♂ 1 ♀ (1 ♂ genit. prep.), 29.vii.2012, 2 ♀♀ (CNCI), K. N. Barber leg.; S[ault] S[te.] Marie, Birchwood Pk., mixed forest, 15.vi.1986, 1 ♂ (MCZC), 28.vi.1986, 1 ♂, 6.vii.1986, 1 ♂ (CNCI); same locality but 46°30.7'N 84°15.6'W, sweeps, mostly Impatiens, under Betula / Acer, 19.vi.1998, 3 ♀♀ (1 ♀ genit.prep.), 23.vi.1998, 1 ♀, sweeps, Impatiens, under Betula / Acer, 27.vi.1998, 1 ♀ (genit. prep.), sweeps, graminoids, Impatiens, ferns under Betula / Acer, 17.ix.2004, 1 ♀; same locality but 46°30.67'N 84°15.63'W, sweeps, Rubus, Aralia, graminoids, ferns, under Betula / Acer, 29.vi.2008, 1 ♂, sweeps, mostly Impatiens, fern, Aster [Eurybia], 23.viii.2009, 1 ♂ (all CNCI), all K. N. Barber leg.; S[ault] S[te.] Marie, Bristol Place, 28.vi.1987, 1 ♀, 1.vii.1987, 1 ♂, K. N. Barber leg. (CNCI); S[ault] S[te.] Marie, Bristol Pl[ace] Pk., 46°30.8'N 84°16.6'W, sweeps, Impatiens under Betula / Acer, 9.vii.1998, 1 ♂ 1 ♀, sweeps, Clematis, Rubus, Impatiens, grasses, 28.v.1999, 1 ♂, sweeps, mostly Impatiens, Clematis, Rubus, grasses, 11.vi.1999, 2 ♂♂ (CNCI); same locality but 46°30.77'N 84°16.66'W, sweeps, Impatiens, Clematis, Equisetum, Rubus, ferns, Phalaris, 29.vi.2008, 2 ♂♂ 3 ♀♀ (CASC), 30.vi.2008, 1 ♂ 1 ♀ (MCZC), 1. vii.2008, 2 ♀♀, 4.vii.2008, 6 ♂♂ 2 ♀♀ (1 ♂ wing illustration, one pair in copula), 5.vii.2008, 1 ♂ 2 ♀♀ (CNCI), 9.vii.2008, 2 ♂♂ 3 ♀♀ (DEBU, 1 ♀ genit. prep.), 11.vii.2008, 7 ♂♂ 3 ♀♀ (1 ♀ genit. prep.), 13.vii.2008, 1 ♀, 27.vii.2009, 1 ♂ 5 ♀♀ (CNCI), 21.vii.2014, 1 ♂ 1 ♀ (SMOC), all K. N. Barber leg.; S[ault] S[te.] Marie, Finn Hill, 46°31.6'N 84°17.4'W, sweeps, vegetation under Populus, 4.vii.2002, 1 ♀; S[ault] S[te.] Marie, Fish Hatchery Road, along Coldwater Ck., 46°34.33'N 84°17.23'W, sweeps, trailside Clematis, Eutrochium, 17.vi.2010, 1 ♂; S[ault] S[te.] Marie, Florwin Dr. greenbelt, 46°30.3'N 84°17.2'W, sweeps, Impatiens under Acer / Betula, 23.vi.1999, 1 ♀; S[ault] S[te.] Marie,Sault Coll[ege] Woodlot, 46°32.08'N 84°18.25'W, sweeps, Clintonia, ferns, under Acer, 31.v.2010, 1 ♂, all K. N. Barber leg. (all CNCI). QUEBEC: Abbotsford, 4.vi.1937, 1 ♂, G. E. Shewell leg.; Beechgrove, 7. vi.1955, 1 ♀, J. F. McAlpine leg.; Beechgrove, 45°39'N 76°08'W, 29.vi.1962, 1 ♂, J. R. Vockeroth leg.; Bolton Pass, Knowlton, 800', 5.vi.1963, 1 ♂, J. G. Chillcott leg. (all CNCI); 1.2 km E Bristol, Silver Creek, 45°31'N 76°27'W, noodplain meadow, 5.vi.1996, 1 ♀, L. Dumouchel leg. (LEMQ 0040277); Gatineau Pk., Harrington Lk., 3.vii.1963, 1 ♂, J. R. Vockeroth leg.; Lac Roddic, 16 km S Maniwaki, 23.vi.1991, 1 ♂ 1 ♀, M. Barták leg. (MBPC, both genit. prep.); Magog, 1.vi.1965, 1 ♀, D. M. Wood leg. (genit. prep.); Old Chelsea, 18.vi.1963, 1 ♂, J. R. Vockeroth leg. (genit. prep.); Old Chelsea, King Mt., 26.v.1963, 1 ♂, J. G. Chillcott leg.; Wakeneld, 26.vi.1946, 1 ♀, 9.vii.1946, 1 ♀, G. E. Shewell leg. (all CNCI). SASKATCHEWAN: Kenosee, 7.vi.1958, 2 ♂♂ 7 ♀♀; Saskatoon, 9.vii.1951, 1 ♀, all A. R. Brooks leg. (all CNCI). UNITED STATES OF AMERICA: CONNECTICUT: Canaan, 17.viii.1952, 1 ♀, A. Stone leg.; Redding, 8.vi.1928, 1 ♀ (genit. prep), A. L. Melander leg. (both USNM). ILLINOIS: Savanna, 13.vi.1917, 1 ♂, [no collector] (INHS 40,188). INDIANA: Lafayette, 24.v.1917, 1 ♂, J. M. Aldrich leg. (USNM). MARYLAND: Bethesda, 14.viii.1981, 1 ♀, G. C. Steyskal leg. (USNM); Montgomery Co., Bethesda, 5.v.1968, 3 ♂♂, L. V. Knutson leg., 26.v.1968, 1 ♂, G. Steyskal leg.; Cabin John, 21.vii.1921, 1 ♂, J. R. Malloch leg. (Malloch det. as A. tenuis); Montgomery Co., Dickerson, 14.vii.1974, 1 ♂ (genit. prep.), G. A. Foster leg.; Glen Echo, 23.vii.1922, 1 ♂, J. R. Malloch leg.; Lavale, 9.v.1970, 1 ♂ (genit. prep., left wing and apical half of right wing missing), G. Steyskal leg. (all USNM); Plummers Is., 12.v.1907, 1 ♂, W. L. McAtee leg. (with det. as A. tenuis), 9.viii.1909, 1 ♀ (genit. prep.), Barber & Schwarz leg., 28.vii.1912, 1 ♀, H. L. Viereck leg., 5.v.1914, 1 ♂, 10.v.1914, 1 ♂, at light, 13.vi.1914, 1 ♀ (genit. prep.), R. C. Shannon leg., Malaise trap, 8–20.vii.1968, 1 ♂ (genit. prep.), Paul Spangler leg. (all USNM). MASSACHUSETTS: Franklin Co., ~0.5 km E Farley, 42°36.16'N 72°25.94'W, sweeps, asters, ferns, Impatiens, Rubus, under canopy, 26.vii.2006, 2 ♂♂ 3 ♀♀, K. N. Barber leg. (CNCI, 1 ♀ genit. prep.); Middlesex Co., Lincoln, Malaise trap, 4–8.vi.1952, 1 ♀, E. T. Armstrong leg. (USNM, genit. prep.). MICHIGAN: Hillsdale Co., 21.v.1960, 1 ♂, R. & K. Dreisbach leg. (USNM). MINNESOTA: Itasca Pk., 12.vii.1952, 1 ♂, [no collector] (AMNH). NEW HAMPSHIRE: Mt. Washington, 18.vii. [-], 1 ♀, C. W. Johnson leg. (MCZC). NEW YORK: Bemus Pt., Chautauqua Lk., swampy woods, 31.v.1963, 1 ♀, W. W. Wirth leg. (USNM); Ithaca, 24.v.1900 [?], 1 ♂, [no collector] (AMNH, genit. prep., wings in genit. vial); New York, 31.v.1923, 1 ♂, A. H. Sturtevant leg.; Tompkins Co., Ringwood Res., swamp, 16–17.vi.1963, 1 ♀, W. W. Wirth leg.; Rome, 24.vi.1935, 1 ♀, H. K. Townes leg. (all USNM); Whiteface Mt., 4600–4872', 19.vii.1962, 1 ♂, J. R. Vockeroth leg. (CNCI). NORTH CAROLINA: Buncombe Co., Blue Ridge Pkwy. at Craggy Gardens, 35.70°N 82.39877°W, 5400', sweep forest path and clearing, 17.viii.2007, 1 ♀, T. A. Wheeler leg. (LEMQ); Swain Co., Rte 441, 3 mi N Cherokee, Great Smoky Mountains National Park, 35°31.3'N 83°18.5'W, 2000', 27.v.1999, 1 ♀, S. D. Gaimari leg. (99-5, Nat’l Pk. Svc. Permit #GRSM-99-074) (USNM). ENNSYLVANIA: Dauphin Co., Grantville, 24.v.1962, 1 ♀, J. R. Vockeroth leg. (CNCI, genit.prep.); Roxborough, 23.v.1909, 1 ♀, [no collector] (USNM, genit. prep.). VIRGINIA: Bland Co., Brushy Mt. at Rd. 623, 37.05229°N 81.30209°W, sweep forest edge, 15.viii.2007, 6 ♂♂ 5 ♀♀, J. Mlynarek leg., 4 ♂♂ 1 ♀ (LEMQ 0040768–72, 1 ♂ genit. prep.), T.A. Wheeler leg., 1 ♂ (LEMQ 0040310, genit. prep.), A. MacLeod leg.; Chain Bridge, 7.v.1922, 1 ♀ (genit. prep.), 20.viii.1922, 1 ♀, 14.v.1924, 1 ♂, J. R. Malloch leg.; Fairfax Co., Dead Run, 28.vii.1915, 1 ♂ 1 ♀, R. C. Shannon leg. (♀ genit. prep.); Falls Church, Holmes Run, light trap, 24.viii.1960, 1 ♀, W. W. Wirth leg. (all USNM); Page Co., G.Washington N. F., Gap Creek Trail, 38.70764°N 78.56077°W, 1662', sweep forest path at creek, 19.viii.2007, 1 ♀, J. Mlynarek leg. (LEMQ 0040534); Great Falls, “ 12-vi ”, 1 ♀, N. Banks leg. (MCZC); Giles Co., Mountain Lake Biol. Stn., 37°22'31"N 80°31'18"W, sweep nr. station, 20.v.2005, 1 ♂, S.A. Marshall leg. (DEBU 00252870); near Plummers Island, MD [Maryland], at light, 20.v.1914, 1 ♀ (genit. prep.), R. C. Shannon leg. (USNM); Shenandoah N. P., mi. 65–100, sweeps, 29.v.1979, 2 ♂♂, M. J. Sharkey leg. (DEBU); Hawksbill Gap, Shenandoah N. P., 1600 m, 17.viii.1981, 1 ♂ 1 ♀, J. R. Vockeroth leg. (CNCI); Shenandoah Co., Mt. Jackson, 25.v.1962, 1 ♂ 1 ♀, J. G. Chillcott leg., 2 ♂♂, J. R. Vockeroth leg. (CNCI); Tazewell, Burkes Garden, Sta[tion] Spr[in]g, 37°05.9'N 81°22.4', 960 m, 18.v.2005, 4 ♂♂, W. N. & D. Mathis leg. (USNM). WEST VIRGINIA: Grant Co., Dahle Soda, 13. vi.1986, 1 ♂, A. L. Norrbom leg. (USNM). Other material examined (not included in type series). Provenance unknown, “Loew Coll.”, “Type 13427” (red label), 1 ♀, (MCZC, genit.prep., erroneously labelled a syntype of Anthophilina tenuis Loew). CANADA: ONTARIO: Moosonee, 51°16.33'N 80°39.11'W, sweeps, mostly Rubus, Impatiens, under Salix, Alnus, 10.vii.2014, 1 ♀, K. N. Barber leg. (CNCI, deformed postabdominal tergites); S[ault] S[te.] Marie, Bristol Pl[ace] Pk., 46°30.77'N 84°16.66'W, sweeps, Impatiens, Clematis, Equisetum, Rubus, ferns, Phalaris, 21.vii.2014, 3 ♂♂, K. N. Barber leg. (SMOC, used for molecular work). SASKATCHEWAN: Kenosee, 7.vi.1958, 1 ♀ (CNCI, headless, genit. prep.). UNITED STATES OF AMERICA: NEW HAMPSHIRE: “N.H.”, “205”, “Loew Coll.”,“ Type 14558” (red label), 1 ♀ (MCZC, headless, double mount with a ♀ of Arganthomyza duplex on the same pinned bricket, erroneously labelled as type specimens of Anthophilina terminalis Loew). NEW YORK: Delaware Co., Franklin Mtn. nr. Oneonta, 3.vii.1980, 1 ♀, D. J. Bickel leg. (MCZC, head glued to pin, genit. prep.). Other A. macra group material of questionable identity (Anthomyza sp. cf. oblonga). CANADA: ONTARIO: S[ault] S[te.] Marie, Baseline Rd., 46°31.40'N 84°24.40'W, sweeps, Aster [Eurybia], Rubus, Equisetum, Carex, ferns, under aspen, 8.viii.2005, 1 ♀ (genit. prep.); S[ault] S[te.] Marie, Birchwood Park, mixed forest, 28.vi.1986, 1 ♀ (genit. prep.), both K. N. Barber leg. (both CNCI). UNITED STATES OF AMERICA: PENNSYLVANIA: Dauphin Co., Grantville, 24.v.1962, 1 ♀, J. R. Vockeroth leg. (CNCI, genit. prep.). Description. Male. Total body length 2.02–2.66 mm. Externally extremely similar to A. tenuis including colouration and chaetotaxy, thus body bicolourous and sparsely pale grey microtomentose as in the latter species. Head shape as in A. tenuis, about as long as high. Colouration of head practically identical with that of the latter species including microtomentose pattern of all its parts. Also mouthparts very similarly coloured. Cephalic chaetotaxy as in A. tenuis, only pvt somewhat longer (usually slightly longer than half length of vti), 3–4 (usually 3) pairs of medial microsetulae in anterior third of frons, 9–10 postocular setulae; vi long as in A. tenuis; subvibrissa weak but usually distinctly longer than anterior peristomal setula. No differences in colour and chaetotaxy of palpus. Eye large and oval as in A. tenuis, with longest (oblique) diameter 1.3–1.4 times as long as shortest. Gena higher than that of A. tenuis, about 0.12 times as long as shortest eye diameter. Also antenna (including arista) as in the latter species although whitish marginal cilia on 1st nagellomere slightly longer. Thorax dark brown dorsally and yellow laterally and ventrally as in A. tenuis but the pattern is yet more variable. In lightest specimens the yellow is extended not only on humeral and notopleural areas but also on lateral sides of mesonotum (up to supralar seta); even disc of scutellum is broadly yellow leaving only its sides brownish. In darkest specimens, on the contrary, humeral callus and a band across dorsal margin of mesopleuron and pteropleuron are also brownish, in addition to darkened laterotergite, mediotergite, subscutellum and scutellum. Intermediates between these extremes are frequent. Thoracic chaetotaxy: hu, npl(s), prs, sa and pa as in A. tenuis; also both (postsutural) dc similar; 5–6 dc microsetae in front of anterior dc, the hindmost only slightly longer than others; ac microsetae less numerous than in A. tenuis, normally in only 2 rows on suture (plus a few single microsetae laterally to them), usually ending slightly behind anterior dc, only rarely reaching up to posterior dc; 2 sc and 2 stpl as in A. tenuis but upcurved setulae on sternopleuron more numerous (4–5) and differently arranged, in more or less perpendicular row with most dorsal setulae longer and situated almost between stpl macrosetae; no setula in front of anterior stpl. Scutellum formed as in A. tenuis but usually paler on disc medially, sometimes largely yellow (see above). Legs yellow as in A. tenuis, with only distal third to half of apical segment of all tarsi brownish. f 1 with ctenidial spine about as long as maximum width of t 1; other pedal chaetotaxies as in A. tenuis. Wing (Fig. 223) closely resembling that of A. tenuis, with hyaline pale ochreous brown membrane and ochreous veins. Venation as in A. tenuis, only with r-m sometimes situated in middle of cell dm, and terminal section of CuA 1 slightly to distinctly longer than dm-cu. Wing measurements: length 2.30–2.74 mm, width 0.79–0.97 mm; Cs 3: Cs 4 = 1.48–1.84, rm\ dm-cu: dm-cu = 1.95–2.79. Haltere yellowish white, knob often lighter. Abdomen colouration and structures very similar to those of A. tenuis. Preabdominal terga brown to pale brown (sometimes slightly paler dorsomedially), distinctly lighter than mesonotum or epandrium. T1 concolourous or slightly paler than T2. T2–T5 as in A. tenuis. Preabdominal sterna somewhat wider than in A. tenuis, pale yellow and with denser setosity. S1 as in A. tenuis, S2–S5 slightly to distinctly wider than long, becoming wider and more transverse posteriorly; S5 the largest and widest, posteriorly distinctly widened. T6 as in A. tenuis but somewhat longer, seemingly bipartite, with dorsomedial unpigmented part narrower. S6–S8 dorsally fused and more or less asymmetrical as usual. S6 and S7 ochreous to yellow in pale specimens, brown and concolourous with T 5 in dark ones, with anterior ledge-like margins always darker brown. Both S6 and S7 usually with more setulae (S6 with 3–5, S7 with 2–4). S8 long, brown to pale brown as T5 or (in pale specimens) with lightened marginal areas. Genitalia. Epandrium (Figs 225, 226) very broad as in A. tenuis but smaller, with similar chaetotaxy, usually with setae denser laterally; anal nssure higher (longer) and cercus slightly larger (compared to epandrium) than in A. tenuis. Medandrium distinctly higher, with dorsolateral corners small (Fig. 225). Gonostylus (Figs 225, 226, 231) distinctive, dissimilar to those of all relatives, roughly suboblong, relatively narrow but with outer side convex, with both distal corners more or less angular, setose only on inner side and externally largely micropubescent, with only anterior marginal band bare (Fig. 226). Hypandrium (Fig. 227) as in A. tenuis but more robust anteriorly. Transandrium (Fig. 228) generally constructed as in relatives, somewhat wider and with terminal arms of caudal process thicker than those of A. tenuis. Pregonite (Fig. 227) similar to that of A. tenuis, 3 anterior and only 4 posterior setae (latter on short lobe-like process). Postgonite more slender (only when viewed in pronle) than that of A. tenuis and with 1–2 setiform microsensilla below 1 usual seta inserted more proximally at anterior margin (Fig. 227). Aedeagal part of folding apparatus with the same armature as in A. tenuis; connecting sclerite also similar although somewhat paler and less sclerotized; basal membrane with dense short excrescences more spine-like (Fig. 227). Phallapodeme more robust, particularly the fulcrum (Fig. 230). Phallophore as in A. tenuis. Saccus (Fig. 230) voluminous but somewhat smaller and its membranous distal part with markedly fewer spines than that of A. tenuis hence resembling that of A. silvatica. Filum (Fig. 230) compact ribbon-shaped, largely dark, fading only towards apex which is (in contrast to A. tenuis) parallel-sided with blunt tip (Fig. 229). Ejacapodeme larger than in relatives (Fig. 230). Female. Similar to male unless mentioned otherwise. Total body length 2.46–3.21 mm. Cephalic and thoracic setae longer and thicker. Orbit rarely (2 specimens seen) with 1–3 additional microsetulae among ors. Antenna with 1st nagellomere only sometimes darker yellow around base of arista. Mesonotum with ac microsetae sometimes more numerous forming up to 4 rows on suture. Ctenidial spine on f 1 distinctly longer and thicker, longer than maximum width of t 1. Wing measurements: length 2.73–3.25 mm, width 0.93–1.13 mm; Cs 3: Cs 4 = 1.33–1.68, rm\dm-cu: dm-cu = 1.85–2.82. Preabdomen with terga more transverse, bicolourous, typically dorsally largely yellow and laterally brown (cf. Fig. 232). T1 more or less ochreous to pale brown; T2 not only laterally but also anteriorly brownish, otherwise yellow, or (more rarely, in dark specimens) completely pale brown. T3–T5 largely yellow, only laterally (on bent sides) brown, with dark parts sharply contrasting with yellow dorsum of sclerites. More rarely T3–T5 (or only some of them) with dorsal pale areas smaller and ochreous and/or with dorsomedial pale brown spot. Preabdominal sterna pale yellow, narrower than in male but also densely (more than in A. tenuis) setose; S2 and S3 about as long as wide; S4 and S5 wider t
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