128,886 research outputs found
Oxyina kashmira Baba & Usmani 2021, sp. n.
Oxyina kashmira sp. n. (Figure 1 M–N, Figure 9, Plate 7) HOLOTYPE. ♀. INDIA: Kashmir: Baramulla: Eco Park, 34.1925°N, 74.2987°E, 28.viii.2020, Tajamul Hassan & Mehdi Tahir, Sweeping Net, (Zoology Museum, Aligarh Muslim University) [1]. Identification: Larger in size; tegmen well developed and exceeding beyond the tip of abdomen; antennae filliform with 22 segments; dorsum of pronotum moderately flattened, median carina diminutive, lateral carinae absent; apical part of hind femur slightly reddish. ♀ genitalia: Subgenital plate long and broad, ventral surface of subgenital plate with a weak median apical concavity, posterior ventral margin of subgenital plate triangular bearing a comparatively short process; cerci conical with subacute apex, exceeding beyond supra-anal plate; supra-anal plate triangular; the dorsal and ventral valves of ovipositor bear diminutive spines, apodeme of ovipositor comparatively shorter, posterior ventral basivalvular plate without spines; apical diverticulum of spermatheca short and straight, not at all coiled, with an apical bifurcation of apex, apices of apical diverticulum turgid; preapical diverticulum much broader, question mark shaped. ♂ Unknown. Geographical distribution: India: Kashmir Morphometry of Oxyina kashmira s p. n.: Etymology: The species name implies to the distribution of this new species in the Kashmir Valley, a region of Jammu & Kashmir state, India.Published as part of Baba, Tajamul Hassan & Usmani, Hd. Kamil, 2021, A revision and first record of the genus Oxyina Hollis, 1975 (Acrididae: Oxyinae) from India, with description of a new species, pp. 151-175 in Zootaxa 4985 (2) on pages 164-168, DOI: 10.11646/zootaxa.4985.2.1, http://zenodo.org/record/494279
Sadayoshia adaro Macpherson & Baba, 2012, n. sp.
Sadayoshia adaro n. sp. (Figs. 3, 7 C) Sadayoshia inermis Macpherson & Baba 2010: 428 (in part, specimens from the Solomon Islands). Sadayoshia tenuirostris Macpherson & Baba 2010: 444 (in part, only one specimen from the Solomon Islands). Material examined. Holotype: Solomon Islands. SALOMON 1, Stn DW1822, 9° 51.8 'S, 160 ° 51.8 'E, 51–54 m, 0 3 October 2001: 1 Ƥ 3.6 mm (MNHN-IU- 2010-5374). Paratype: Solomon Islands. SALOMON 1, Stn DW1744, 10°32.0'S, 159 ° 38.9 'E, 49–55 m, 23 September 2001: 1 ov. Ƥ 3.9 mm (MNHN-IU- 2010-5376).—Stn DW1822, 9° 51.8 'S, 160 ° 51.8 'E, 51–54 m, 0 3 October 2001: 1 ov. Ƥ broken (MNHN-IU- 2010-5375). Etymology. The name adaro refers to a sea-spirit in the Salomon Islands myths. The name is considered as a substantive in apposition. Description. Carapace: As long as broad; dorsal surface nearly horizontal from anterior to posterior; cervical groove distinct. Dorsal surface with 4 pairs of epigastric spines, 1 parahepatic and 1 or 2 anterior branchial spines on each side; ridges with numerous short uniramous setae. Gastric region with 5 transverse ridges behind epigastric spines, first and third to fifth uninterrupted, second interrupted (scale-like median ridge flanked by longer ridge on each side). Mid-transverse ridge of carapace uninterrupted, preceded by cervical groove. Posterior branchial region laterally with 6 ridges (exclusive of mid-transverse ridge and posteriormost transverse ridges directly anterior to posterior margin of carapace). Lateral margins slightly or moderately convex, with 7 spines. Rostrum 1.8 times as long as broad, length 0.4 that of carapace, lateral margin with 2 spines; rostral spine 1.2 times longer than wide (measured at sinus between rostral and anterior lateral spines), dorsal surface flat, with short setiferous striae, nearly horizontal in lateral view, one minute distal tooth on each side. Pterygostomian flap rugose with sparse short ridges, ending in blunt point. Sternum: Sternite 3 1.6 times as broad as long, anterior margin strongly convex, with small median notch. Sternite 4 1.6 times as long as and 2.7 times broader than preceding sternite, 2.7 times broader than long; surface with 2 transverse ridges, anterior medially interrupted. Sternites 4–6 with 0–2 short ridges. Lateral parts of sternite 7 without granules. Abdomen: Somite 2 unarmed; somites 2–4 each with 4 transverse ridges, anterior first and third uninterrupted, second and fourth interrupted; ridges with numerous short setae and sparse long iridescent setae. Eyes: Peduncles 1.4 times longer than broad; cornea moderately dilated, maximum corneal diameter about 0.4 times distance between bases of anterolateral spines; eyelashes short, not reaching midlength of cornea. Antennule: Article 1 with 4 distal spines: mesial and medioventral smaller than others, lateral largest; lateral margin unarmed; ventral surface smooth. Antenna: Article 1 with distomesial spine slightly exceeding distal margin of article 2. Article 2 with distolateral spine longer than distomesial, terminating in midlength of article 3. Articles 3 and 4 unarmed. Mxp 3: Ischium with very small distal spine on flexor margin; extensor margin bluntly produced distally; crista dentata with 23–26 denticles. Merus as long as ischium, flexor margin with 2 subequal spines, distal one terminal, proximal one at midlength; extensor margin unarmed. Carpus unarmed. P 1: 1.8 times carapace length, relatively stout, with long uniramous iridescent setae along lateral and mesial margins of merus, palm and fingers; dorsal and ventral surfaces of palm and fingers with short setae. Merus 0.6 times as long as carapace, 2.2 times as long as carpus, with strong distal spines on dorsal and mesial margins, and a few additional small spines on dorsal surface. Carpus 0.7 times length of palm, and 0.9 times as long as broad; dorsal surface with spines; mesial surface with well-developed spines. Palm 0.9 times longer than broad; with dorsolateral and mesial rows of spines continued on to entire margins of fixed and movable fingers, respectively; dorsal surface with a few spines. Fingers 1.3 times as long as palm, distally spooned. P 2–4: Somewhat compressed mesio-laterally, sparsely with long uniramous iridescent setae, and short setae. Meri successively shorter posteriorly; P 2 merus 0.6 times carapace length, 3.1 times as long as broad, 1.2 times longer than P 2 propodus; P 3 merus 2.7 times as long as broad, 1.2 times length of P 3 propodus; P 4 merus 3.0 times as long as broad, 1.1 times length of P 4 propodus; dorsal margins with row of proximally diminishing spines on P 2 and P 3, nearly unarmed on P 4; ventrolateral margins with strong terminal spine. Carpi with 3 extensor marginal spines distinct on P 2 and P 3, obsolescent on P 4, distalmost larger; flexor margins with distal spine; lateral surface with 0 or 1 small spine. Propodi 3.2–3.6 times as long as broad; extensor margins with 2 proximal spines, lateral side unarmed; flexor margin with 6–8 slender movable spines. Dactyli subequal in length, 0.8–0.9 times length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 or 6 seta-like movable spines, each arising from obsolescent tooth. Remarks. The new species is closely related to S. inermis Macpherson & Baba, 2010, sharing the thick rostrum and the absence of postcervical spines on the carapace. However, these species can be differentiated by the shape of the rostral spine. This spine is very short and thick, being less than 2 two times longer than wide (measured at the sinus between the rostral and anterior lateral spines) in S. adaro, whereas it is 2.5 –3.0 times longer in S. inermis. Furthermore, the flexor margin of the Mxp 3 usually has 2 spines in S. adaro and usually 3 – or 4 spines in S. inermis. Distribution. Solomon Islands, at 51– 54 m.Published as part of Macpherson, Enrique & Baba, Keiji, 2012, The squat lobsters of the genus Sadayoshia Baba, 1969 (Crustacea: Decapoda: Anomura: Munididae): new records including six new species from the Pacific Ocean, pp. 30-48 in Zootaxa 3589 on pages 36-38, DOI: 10.5281/zenodo.21464
Uroptychus imparilis Baba 2018, n. sp.
Uroptychus imparilis n. sp. Figures 105, 106 Uroptychus scandens Baba 1981: 132 (not Uroptychus scandens Benedict, 1902). TYPE MATERIAL — Holotype: Philippines. MUSORSTOM 1 Stn CP 40, 13°57.4’N, 120°27.8’E, 287- 265 m, 24.III.1976, ov. ♀ 3.8 mm (MNHN-IU-2013-8565). Paratypes: Philippines. MUSORSTOM 1, collected with holotype 1 ♂ 3.2 mm, 1 ov. ♀ 3.6 mm (MNHN-IU-2014-10143 & MNHN-IU-2014-10144), 1 ov. ♀ 4.0 mm (MNHN-IU-2014-16579). MUSORS- TOM 2 Stn CP75, 13°51’N, 120°30’E, 300-330 m, 1.XII.1980, 1 ov. ♀ 3.2 mm (MNHN-IU-2014-16580). Indonesia, Kai Islands. KARUBAR Stn CP06, 5°49’S, 132°21’E, 298- 287 m, 22.X.1991, 1 ♂ 3.5 mm (MNHN-IU-2014-16582). Vanuatu. MUSORSTOM 8 Stn CP1025, 17°49.01’S, 168°39.37’E, 385-410 m, 28.IX.1994, 1 ov. ♀ 3.0 mm (MNHN-IU-2014-16583). OTHER MATERIAL EXAMINED — Japan, Tosa Bay. On pennatulacean, 24.XII.1959, K. Sakai coll., 1 ov. ♀ (carapace broken), 1 ♀ 3.6 mm (ZLKU 7466); 150 m, 31.I.1959,K.Kurohara coll., 1 ♂ 3.6 mm (ZLKU 5871); IV.1960,K.Sakai coll., 1 ov. ♀ 3.2 mm (ZLKU 14393). Southern Kyushu,SOYO-MA- RU Stn 72d, 31°13.6’N, 129°58.5’E, 310 m, 12.II.1959, 2 ♂ 2.7, 3.1 mm, 1 ♀ 3.2 mm (NSMT). Hachijo-jima, SOYO-MARU Stn B3, 33°06.5’N, 140°04.8’E, 490-495 m, 12.XII.1963, 1 ♂ 3.7 mm (NSMT). ETYMOLOGY„ From the Latin imparilis (different), suggesting that the species is different from the closest congener U. parisus n. sp. (see below). DISTRIBUTION„ Philippines, Indonesia, Vanuatu and Japan; 150- 495 m. SIZE„ Males, 2.7-3.7 mm; females, 3.0- 3.8 mm; ovigerous females from 3.0 mm. DESCRIPTION„ Small species. Carapace: Broader than long (0.8 × as long as broad); greatest breadth 1.8 × distance between anterolateral spines. Dorsal surface setose laterally, with very small spines on epigastric, hepatic and anterior lateral branchial regions, those on hepatic region somewhat larger. Lateral margins convexly divergent posteriorly, occasionally slightly constricted between hepatic and branchial regions, bearing row of small spines; anterolateral spine much larger than others, directed straight forward, overreaching much smaller lateral orbital spine. Rostrum narrow triangular, with interior angle of 25°; lateral margins feebly concave, with 8-9 denticles; dorsal surface somewhat concave; length slightly smaller than (0.9 ×) breadth, 0.4 × that of remaining carapace, breadth less than half carapace breadth measured at posterior carapace margin; lateral orbital spine slightly anterior to level of anterolateral spine. Pterygostomian flap anteriorly angular, produced to strong sharp spine; surface with small spines on anterior half, including row of spines along anterior part of dorsal margin. Sternum: Excavated sternum with slightly convex anterior margin; surface smooth, without ridge and central spine, with setae along anterior margin; sternal plastron 0.8 × as long as broad (1.3 × broader than long); anterior margin of sternite 3 broadly and deeply excavated nearly in semicircular shape, with pair of small median spines basally contiguous (one of the spines occasionally missing), anterolateral margin with several small or obsolescent spines. Sternite 4 having anterolateral margin feebly convex, anteriorly produced to distinct spine and followed by a few smaller spines; posterolateral margin 0.7 × as long as anterolateral margin. Anterolateral margin of sternite 5 as long as posterior margin of sternite 4. Abdomen. Somite 1 gently convex from anterior to posterior. Somite 2 tergite 2.5 × broader than long, pleura similar in males and females, slightly concavely divergent posteriorly, ending in rounded margin. Pleura of somites 3-5 laterally rounded. Telson slightly less than half as long as broad; posterior plate slightly emarginate or feebly convex, 1.7 × longer than anterior plate. Eyes: Elongate, 1.9-2.1 × longer than broad, reaching or very slightly falling short of rostral tip, lateral margin convex, mesial margin concave. Cornea slightly inflated, very slightly broader than and half as long as remaining eyestalk. Antennule and antenna: Ultimate article of antennular peduncle 2.7-2.9 × longer than high. Antennal peduncle slightly overreaching rostral tip. Article 2 with strong distolateral spine. Antennal scale articulated with article 2, ending in midlength of article 5, laterally with 1 or 2 spines; breadth 1.3-1.4 × that of article 5. Article 3 distomesially rounded or produced to small spine. Article 4 with distinct distomesial spine. Article 5 with small distomesial spine, length 1.5 × that of article 4, breadth 0.4-0.5 × height of ultimate article of antennule. Flagellum of 8-10 segments ending in midlength of P 1 merus. Mxp: Mxp1 with bases broadly separated. Mxp3 with long setae. Basis with 1 distal denticle on mesial ridge. Ischium with small spine lateral to distal end of flexor margin, crista dentata with a few to several obsolescent denticles on proximal half. Merus 2.5 × length of ischium, ridged along flexor margin, not well compressed; with 3 spines on distal third of flexor margin and 2 distolateral spines. Carpus unarmed. P 1: Slender, subcylindrical, 5.7-6.6 × (males), 4.4-6.8 × (females) longer than carapace. Ischium dorsally with basally broad, depressed, bifurcate short spine, ventromesially unarmed. Merus 1.5 × longer than carapace, dorsally bearing 2 rows of spines: 1 row along mesial margin and another row of smaller (some bifurcated) spines directly dorsolateral to it). Carpus 1.3-1.4 × longer than merus, unarmed. Palm 4.1-5.0 × (males), 6.4-6.6 (females) as long as broad, slightly (females) or distinctly (males) broadened distally, length 0.9 × that of carpus. Fingers relatively narrow distally, strongly gaping in proximal two-thirds in males, weakly so in proximal third in females; opposable margins fitting to each other in distal two-thirds (females) or in distal third (males) when closed, with row of denticles; movable finger 0.4 × length of palm, with obtuse process at midpoint of gaping portion. P 2-4: Thickly setose like P 1. Meri successively shorter posteriorly (P 3 merus 0.9 × length of P 2 merus, P 4 merus 0.8 × length of P 3 merus), slightly broader on P 3 and P 4 than on P 2, or slightly broader on P 3 than on P 2 and P 4; lengthbreadth ratio, 4.1-4.2 on P 2, 3.4 on P 3, 2.9-3.0 on P 4; dorsal margin with 7-9 small spines on P 2, 6-7 on P 3, unarmed on P 4; P 2 merus subequal to length of carapace, 1.3-1.4 × length of P 2 propodus; P 3 merus 1.3 × length of P 3 propodus; P 4 merus 1.2-1.3 × length of P 4 propodus. Carpi successively shorter posteriorly; carpus-propodus length ratio, 0.6 on P 2, 0.5 on P 3 and P 4. Propodi successively shorter posteriorly; flexor margin straight in lateral view, with pair of small terminal spines only. Dactyli subequal on P 2-4; dactylus-carpus length ratio, 0.5-0.6 on P 2, 0.6-0.7 on P 3, 0.6-0.8 on P 4; dactylus-propodus length ratio, 0.3 on P 2, 0.3-0.4 on P 4; truncate, bearing 7 slender spines obscured by setae on P 2 and P 3, 6 spines on P 4, 3 or 4 of these located on terminal margin, remainder perpendicular to flexor margin, terminal spine smaller. Eggs. Number of eggs carried, about 40; size, 0.63 × 0.68 mm - 0.61 × 0.71 mm. REMARKS — The material registered under ZLKU 5871 contains three specimens. One is identified as U. imparilis and the other two, with the label “associated with a pennatulacean,” are Uroptychus scandens. The materials reported under U. scandens by Baba (1981) are referable to U. imparilis. This species shares truncate dactyli of P 2-4 with U. articulatus n. sp., U. parisus n. sp. and U. scandens Benedict, 1902. Their relationships are discussed under U. scandens (see below).Published as part of Baba, Keiji, 2018, Chirostylidae of the Western and Central Pacific: Uroptychus and a new genus (Crustacea: Decapoda: Anomura), pp. 1-612 in Mémoires du Muséum national d'Histoire naturelle 212 on pages 232-236, DOI: 10.5281/zenodo.376097
Heteroptychus galapagos Baba & Wicksten 2019, n. sp.
Heteroptychus galapagos n. sp. (Figs. 5, 6) Type material. CDF NA064-031-01-01-A, holotype, ovigerous female (CL 6.1 mm), ROV Dive H1436, East Darwin Seamount, 1°40.5'N, 91°41.2'W, 1012 m, 28 June 2015. Description. Carapace: 0.8 × as long as broad, greatest breadth 1.9 × distance between anterolateral spines. Dorsal surface polished, somewhat convex from anterior to posterior, without depression between gastric and cardiac regions, ridged along posterior half of lateral margin, greatest breadth measured at posterior third. Lateral margins convexly divergent posteriorly; anterolateral spine well developed, directed anterodorsally and straight forward; smooth except for 2 small, distinct protuberances, one at anterior end of anterior branchial margin, another at anterior end of posterior branchial margin. Rostrum slightly overreaching apex of eye, 0.7 × as long as broad, distinctly concave on lateral margin; distally blunt and setose, with interior angle of 22°, length one-quarter postorbital carapace length, breadth less than half (0.4) that of carapace measured at posterior margin; dorsal surface flattish but feebly concave basally, horizontally directed straight forward. Lateral limit of orbit rounded, without spine. Pterygostomian flap with posterior height 0.4 × anterior height, anteriorly produced to sharp spine; anterior surface well inflated (convex from dorsal to ventral), with 2 low ridges in midline. Thoracic sternum: Excavated sternum with triangular anterior end followed by rounded longitudinal ridge in midline. Sternal plastron 0.6 × as long as broad, lateral extremities divergent posteriorly, sternite 6 as broad as sternite 7. Posterior margin strongly concave. Anterior margin of sternite 3 deeply excavated in semicircular shape, with small, shallow median notch, lacking submedian spines. Abdomen: Smooth, polished. Somite 1 tergite gently convex from anterior to posterior. Somite 2 tergite 2.0 × broader than long; pleural lateral margins moderately concave, strongly divergent posteriorly, with blunt terminus. Pleura of somites 3–4 ending in rounded margin. Telson 0.4 × as long as broad; posterior plate slightly concave on posterior margin, 1.1 × longer than, and 0.8 × as broad as anterior plate. Eyes: Ovate, 1.7 × as long as broad. Cornea 0.8 × as long as remaining eyestalk. Antennule and antenna: Ultimate article of antennular peduncle 4.1 × longer than high. Antennal peduncle overreaching apex of rostrum by half length of article 5. Antennal scale fused with article 2, 1.2 × broader than article 5, terminating in distal quarter of article 4. Distal 2 articles unarmed; article 5 1.4 × longer than article 4, breadth 0.5 × height of ultimate antennular article. Flagellum of 18 segments. Mxp: Mxps 1 with bases widely separated. Mxp 3 basis with distal denticle on mesial ridge; ischium crista dentata with 5 widely spaced denticles; merus 2.6 × longer than ischium, relatively thick mesio-laterally, unarmed, with rounded ridge along flexor margin; extensor and flexor margins subparallel; carpus unarmed. P1: Left P1 missing, right P1 with distal part missing, presumably taken for tissue sample. Ischium with basally broad, distally blunt distodorsal process and proximal dorsal lobe-like process. Merus with short distodorsal and smaller distoventral spine mesially, length 1.7 × that of carapace. Carpus 0.8 × as high as broad, 1.4 × longer than merus, with row of 8 small spines along mesial margin, and 4 spines on distal margin at juncture with palm. P2–4: Long setae on propodi, thickly setose along prehensile margins of propodus and dactylus. P2 and P3 subequal. P4 merus 0.4 × as long as P3 merus; length-breadth ratio, 7.5 on P2 and P3, 3.6 on P4; P2 merus 1.1 × carapace length, 1.4 × length of propodus. P3 merus 1.4 × length of P3 propodus. P4 merus 0.7 × length of P4 propodus. P2 and P3 carpi 3.1 × longer than P4 carpus, 1.2 × longer than P2 and P3 propodi; P4 carpus 0.4 × length of P4 propodus. Propodi 1.6 (P2 and P3), 1.4 × (P4) longer than dactyli, flexor margins concavely curving in lateral view, unarmed. Dactyli 0.5 × (P2 and P3), 1.7 × (P4) length of carpi; flexor margin with row of 15 or 16 sharp spines proximally diminishing and nearly perpendicular to margin, ultimate and penultimate spines subequal. Eggs: Number of eggs carried, 36; size, 1.32 × 1.40 mm to 1.45 × 1.14 mm. Coloration: Information not available. Etymology. Named for the type locality of the species; used as a noun in apposition. Remarks. Only two specimens referable to this genus are identified as new species, H. galapagos and H. nautilus (see below). These species share a non-articulated antennal scale, which links them to H. lemaitrei Baba, 2018 and H. claudeae Baba, 2018, both from the western Pacific. Heteroptychus galapagos n. sp. differs from H. lemaitrei Baba, 2018 in having two instead of only one protuberance on the carapace lateral margin, in having the rostrum horizontally directed straight forward rather than dorsally directed, in having the anterolateral spine of the carapace directed straight forward and dorsally instead of anteromesially and horizontally, in having the lateral orbital angle rounded instead of bearing a distinct spine, and in having the pterygostomian flap surface with two distinct ridges instead of being smooth without ridges. Heteroptychus claudeae has no protuberance on the carapace lateral margin, the anterolateral spine of the carapace closer to the lateral limit of the orbit, the rostrum more or less depressed distally rather than spiniform, the pterygostomian flap much lower in the posterior half (posterior height/anterior height, 0.1 versus 0.4). Morphological differences between H. galapagos and H. nautilus are outlined under the remarks of H. nautilus (see below). Distribution. East Darwin Seamount, Galapagos Islands at 1012 m depth.Published as part of Baba, Keiji & Wicksten, Mary K., 2019, Chirostyloidean squat lobsters (Crustacea: Decapoda: Anomura) from the Galapagos Islands, pp. 391-421 in Zootaxa 4564 (2) on pages 397-400, DOI: 10.11646/zootaxa.4564.2.5, http://zenodo.org/record/258923
Uroptychus septimus Baba 2018, n. sp.
Uroptychus septimus n. sp. Figures 235, 236 TYPE MATERIAL — Holotype: Wallis and Futuna Islands. MUSORSTOM 7 Stn DW 523, 13°12’S, 176°16’W, 455-515 m, with corals of Chrysogorgiidae (Calcaxonia), 13.V.1992, ov. ♀ 7.1 mm (MNHN-IU-2014-16947). ETYMOLOGY„ From the Latin septimus (seventh), alluding to the seventh cruise of MUSORSTOM, by which the species was taken. DISTRIBUTION„ Wallis and Futuna Islands (SW Pacific); 455- 515 m. DESCRIPTION„ Medium-sized species. Carapace: Nearly as long as broad; greatest breadth 1.5 × distance between anterolateral spines. Dorsal surface smooth and glabrous, nearly horizontal in profile, smoothly sloping down anteriorly to rostrum. Lateral margins slightly convex; anterolateral spine small, located somewhat posterior to level of lateral orbital angle, and not overreaching that spine; tubercle-like small spine at point one-fifth from anterior end (anterior end of branchial margin), followed by granulation; no ridge along posterior portion. Rostrum narrow triangular, with interior angle of 23°, slightly overreaching eyes; dorsal surface somewhat concave; ventral surface straight horizontal; lateral margins somewhat concave; length less than half that of remaining carapace, breadth half carapace breadth at posterior carapace margin. Lateral limit of orbit acuminate, lacking distinct spine. Pterygostomian flap with somewhat roundish anterior margin lacking distinct spine; surface smooth. Sternum: Excavated sternum with anterior margin with small median spine between bases of Mxp1; surface with small spine in center. Sternal plastron 1.1 × broader than long, lateral extremities straight divergent posteriorly. Sternite 3 depressed well, anterior margin strongly excavated in broad V-shape, with pair of sharp submedian spines separated by V-shaped notch, anterolateral angle rounded. Sternite 4 having convex anterolateral margin anteriorly ending in spine nearly as large as submedian spines of sternite 3, followed by posteriorly diminishing denticles; posterolateral margin about half as long as anterolateral margin. Sternite 5 having anterolateral margin strongly convex, 1.3 × longer than posterolateral margin of sternite 4. Abdomen: Smooth and glabrous, with relatively long somites. Somite 1 feebly convex from anterior to posterior. Somite 2 tergite 2.1 × broader than long; pleuron posterolaterally blunt; lateral margins concavely strongly divergent posteriorly. Pleuron of somite 3 with blunt lateral terminus. Telson two-thirds as long as broad; posterior plate distinctly emarginate, length 1.8 × that of anterior plate. Eye: Slightly falling short of apex of rostrum, about twice as long as broad, lateral and mesial margins somewhat concave. Cornea somewhat inflated, length much more than half that of remaining eyestalk. Antennule and antenna: Ultimate article of antennular peduncle 2.5 × longer than high. Antennal peduncle relatively slender, reaching distal end of cornea. Article 2 distolaterally acuminate, lacking distinct spine. Antennal scale 1.5 × broader than article 5, tapering, slightly falling short of distal end of article 5 and distal end of eye. Article 4 unarmed. Article 5 with small distomesial spine; length 2.7 × that of article 4, breadth half height of ultimate article of antennule. Flagellum consisting of 17 or 20 segments, nearly reaching distal margin of P 1 merus. Mxp: Mxp1 with bases very close to each other. Mxp3 scarcely setose laterally. Basis with 1 distal denticle proximally followed by 1 or 2 obsolescent denticles. Ischium with 25-26 denticles on crista dentata, flexor margin distally not rounded. Merus 1.9 × longer than ischium, unarmed, not well compressed mesio-laterally, ridged along smoothly convex flexor margin. Carpus with obsolescent distolateral spine. P 1: Moderately massive, 5.2 × longer than carapace; sparsely setose except for fingers, setae relatively short. Ischium dorsally with broad short spine, ventromesially feebly denticulate, without subterminal spine. Merus 1.3 × longer than carapace, unarmed except for very small distomesial and distolateral spines on ventral surface, ventromesially feebly granular. Carpus 1.2 × longer than merus. Palm smooth dorsally, rounded on mesial margin, 2.8 × longer than broad, 0.9 × length of carpus. Fingers slightly gaping, distally crossing when closed; fixed finger with low median eminence on opposable margin; movable finger about half as long as palm, opposable margin with disto-proximally broad, bluntly bidentate proximal process. P 2-4: Slender, with sparse long setae. Meri moderately compressed mesio-laterally, dorsally unarmed, ventrolaterally ending in small spine; successively diminishing posteriorly (P 3 merus 0.9 × length of P 2 merus, P 4 merus 0.9 × length of P 3 merus), subequally broad on P 2 and P 3, somewhat narrower on P 4 (P 4 merus 0.9 × breadth of P 3 merus); lengthbreadth ratio, 6.2 on P 2, 5.6 on P 3 and P 4; P 2 merus as long as carapace, 1.5 × longer than P 2 propodus; P 3 merus 1.2 × length of P 3 propodus; P 4 merus as long as P 4 propodus. Carpi subequal in length on P 2 and P 3, shortest on P 4; much longer than dactyli (carpus-dactylus length ratio, 2.0 on P 2, 1.7 on P 3, 1.5 on P 4); carpus-propodus length ratio, 0.7 on P 2, 0.6 on P 3, 0.5 on P 4; extensor margin with small distal spine at most on P 2. Propodi longest on P 3, shortest on P 2; flexor margin nearly straight, with pair of distal spines slightly distant from juncture with dactylus and preceded by row of spines (8 or 11 on P 2, 8 on P 3, 6 or 7 on P 4) nearly along entire length on P 2, slightly more distally on P 3 and P 4, distalmost of these unpaired spines equidistant between distal second and terminal pair. Dactyli relatively slender, strongly curving at proximal third, ending in prominent spine preceded by 7-8 smaller flexor marginal spines moderately inclined, loosely arranged and diminishing toward base of article; dactylus-carpus length ratio, 0.5 on P 2, 0.6 on P 3, 0.7 on P 4; dactyluspropodus length ratio, 0.35-0.36 on P 2, 0.35 on P 3, 0.36 on P 4. Eggs. Number of eggs carried, 7; diameter, 1.4 mm. REMARKS„ The new species resembles U. comptus Baba, 1988, U. empheres Ahyong & Poore, 2004, U. pollostadelphus n. sp. and U. sagamiae Baba, 2005 in the carapace shape, in having the pterygostomian flap roundish on the anterior margin, and in having the P 2-4 dactyli with the ultimate of the flexor spines strongest. Uroptychus comptus and U. empheres can be separated from the other species including U. septimus by having no epigastric spines. They also differ from U. septimus in having no spine on the antennal article 5 and in having a ventrally granulose P 1 merus. Uroptychus septimus is differentiated from U. pollostadelphus by the P 2-4 meri that are successively shorter posteriorly, with the P 4 merus 0.9 times as long as the P 3 merus (in U. pollostadelphus, the P 2 merus is as long as the P 3 merus and the P 4 merus is much shorter, 0.6 times the length of the P 3 merus); the antennal article 2 is distolaterally acuminate instead of bearing a distinct spine, and the article 5 bears a distinct distomesial spine instead of being unarmed. Uroptychus septimus differs from U. sagamiae Baba, 2005 in having the antennal article 2 distolaterally acuminate instead of bearing a distolateral spine; the antennal scale is barely reaching instead of overreaching the distal end of the article 5; the P 1 palm is smooth instead of denticulate on the ventral surface; the P 2 merus is as long as instead of much shorter than the carapace and 1.5 times longer than instead of as long as the P 2 propodus; and the P 2 carpus is longer, being 0.7 instead of 0.5 times the length of the P 2 propodus.Published as part of Baba, Keiji, 2018, Chirostylidae of the Western and Central Pacific: Uroptychus and a new genus (Crustacea: Decapoda: Anomura), pp. 1-612 in Mémoires du Muséum national d'Histoire naturelle 212 on pages 470-473, DOI: 10.5281/zenodo.376097
Sadayoshia actaea Macpherson & Baba, 2012, n. sp.
Sadayoshia actaea n. sp. (Figs. 2, 7 B) Sadayoshia latisternata Macpherson & Baba 2010: 433 (in part, only specimens from New Caledonia). Material examined. Holotype: New Caledonia, Lifou Island. LIFOU, Stn 1467, 20º 46.6 'S, 167 º05.7'E, 90 m, 20 November 2000: Ƥ 3.2 mm (MNHN-IU- 2010-5372). Paratypes: New Caledonia, Lifou Island. LIFOU, Stn DW 14 Stn 1467, 20º 46.6 'S, 167 º05.7'E, 90 m, 20 November 2000: 1 Ƥ 3.5 mm (MNHN-IU- 2010-5373). Etymology. The name actaea refers to one of the Nereids in the Greek mythology. The name is considered as a substantive in apposition. Description. Carapace: As long as broad; dorsal surface nearly horizontal from anterior to posterior; cervical groove distinct. Four pairs of epigastric spines and 1 parahepatic spine on each side; 4 uninterrupted ridges on gastric region behind epigastric spines; mid-transverse ridge of carapace uninterrupted, preceded by cervical groove. Posterior branchial region laterally with 4 ridges (exclusive of mid-transverse ridge and posteriormost transverse ridges directly anterior to posterior margin of carapace). Dorsal surface with short uniramous and a scattered long setae arising from transverse ridges. Lateral margins medially convex, with 7 spines. Rostrum moderately broad, 1.1 times as long as broad, length 0.4 times that of carapace; dorsal surface nearly horizontal in lateral view, slightly convex from side to side, without setiferous striae; rostral spine 2.1 times longer than wide (measured at sinus between rostral and anterior lateral spines). Pterygostomian flap rugose with sparse setae, anterior margin ending in blunt point. Sternum: Sternal plastron as long as broad, lateral limits divergent posteriorly. Sternite 3 3.2 times as broad as long, anterior margin moderately produced, with small V-shaped median notch. Sternite 4 2.4 times longer and twice as broad as preceding sternite, 3.9 times broader than long; surface with 1 anterior stria flanking midline. Following sternites smooth. Lateral parts of sternite 7 smooth. Abdomen: Somite 2 unarmed; somites 2–4 without transverse ridges other than anterior one bearing both short and scattered long uniramous setae. Eyes: Peduncle as long as broad; cornea slightly dilated; maximum corneal diameter about 0.4 times distance between bases of anterolateral spines; eyelashes short, not reaching midlength of cornea. Antennule: Article 1 with 4 distal spines: mesial and medioventral subequal and clearly smaller than others; lateral margin unarmed; ventral surface with a few short scales. Antenna: Article 1 with distomesial spine nearly reaching distal margin of article 2. Article 2 with distolateral spine larger than distomesial, reaching midlength of article 3. Articles 3 and 4 unarmed. Mxp 3: Ischium with small distal spine on flexor margin; extensor margin unarmed; crista dentata with 29 denticles. Merus subequal in length to ischium, flexor margin with 2 subequal small spines: distal terminal, proximal at midlength; extensor margin unarmed. Carpus unarmed. P 1: 1.2 times carapace length, stout, with long uniramous iridescent setae along lateral and mesial margins of merus, palm and fingers; dorsal and ventral surfaces of palm and fingers with scattered short setae. Merus about 0.4 times length of carapace, 2.5 times as long as carpus, with dorsal and lateral spines, and 1 strong distomesial spine. Carpus 0.5 times length of palm, 0.5 times as long as broad; with small spines on dorsal surface, 2 well-developed spines on mesial margin and a few spines along distal margin. Palm 0.8 times as long as broad; mesial and lateral margins with strong spines continued on to fingers, dorsal surface with sparse spines. Fingers distally spooned; movable finger as long as palm, dorsal surface with 2 proximal spines. P 2–4: Somewhat compressed mesio-laterally, sparsely with long uniramous iridescent setae and some short plumose setae. Meri successively shorter posteriorly; P 2 merus 0.6 times carapace length, 3.0 times as long as broad, 1.4 times longer than P 2 propodus; P 3 merus 2.9 times as long as broad, 1.2 times length of P 3 propodus; P 4 merus 3.1 times as long as broad, 1.1 times length of P 4 propodus; dorsal margins of meri with row of proximally diminishing spines on P 2 and P 3, minute spines on P 4; dorsolateral surface unarmed on P 2 and P 3, with 2 spines on P 4; ventrolateral margins with well-developed terminal spine. Carpi with 3 spines on extensor margin of P 2 and P 3, unarmed on P 4, distalmost largest; lateral surface unarmed or with 1 spine; flexor margins with distal spine. Propodi subequal in length on P 2 and P 3. slightly shorter on P 4, each about 3 times as long as broad; extensor margin with 1 or 2 proximal spines, lateral side unarmed; flexor margin with 5–7 slender movable spines on P 2–4. Dactyli subequal in length, 0.7–0.8 times length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 or 6 obsolescent teeth, each with seta-like movable spine. Remarks. The new species strongly resembles S. latisternata Macpherson & Baba 2010 in the relatively smooth abdominal somites 2–4 bearing only an anterior ridge and the shape of sternite 3 short relative to breadth. Sadayoshia actaea can be distinguished from S. latisternata by lacking instead of possessing the anterior branchial spine on the carapace. Furthermore, the rostral spine is about 2.0 times longer than wide (measured at sinus between rostral and anterior lateral spines) in S. actaea, whereas it is about 3 times longer in S. latisternata. Distribution. New Caledonia, Lifou Island, at 90 m.Published as part of Macpherson, Enrique & Baba, Keiji, 2012, The squat lobsters of the genus Sadayoshia Baba, 1969 (Crustacea: Decapoda: Anomura: Munididae): new records including six new species from the Pacific Ocean, pp. 30-48 in Zootaxa 3589 on pages 34-36, DOI: 10.5281/zenodo.21464
Uroptychus adiastaltus Baba 2018, n. sp.
Uroptychus adiastaltus n. sp. Figures 5, 6 TYPE MATERIAL — Holotype: New Caledonia, Isle of Pines. BIOCAL Stn DW 33, 23°11’S, 167°10’E, 675-680 m, 29.VIII.1985, 1 ov. ♀ 4.3 mm (MNHN-IU-2012-687). Paratypes: New Caledonia, Isle of Pines. BIOCAL Stn DW33, 23°11’S, 167°10’E, 675-680 m, 29.VIII.1985, 3 ♂ 3.3-4.9 mm, 7 ov. ♀ 3.7-4.6 mm (MNHN-IU-2014-17157). – Stn DW36, 23°09’S, 167°11’E, 650-680 m, 29.VIII.1985, 1 ♂ 2.8 mm, 2 ov. ♀ 3.5, 3.7 mm (MNHN-IU-2012-685). – Stn DW51, 23°05’S, 167°45’E, 680- 700 m, 31.VIII.1985, 13 ♂ 2.3-3.1 mm, 12 ov. ♀ 2.7-3.6 mm, 5 ♀ 2.2-3.9 mm (MNHN-IU-2012-684). MUSORSTOM 4 Stn DW220, 22°58.5’S, 167°38.3’E, 505-550 m, 29.IX.1985, 5 ♂ 3.3-3.7 mm, 2 ov. ♀ 3.5, 3.6 mm, 1 ♀ 3.1 mm (MNHN- IU-2012-686). New Caledonia, Norfolk Ridge. NORFOLK 1 Stn DW1666, 23°42’S, 167°43’E, 469-860 m, 20.VI.2001, 1 ov. ♀ 4.9 mm (MNHN-IU-2012-693). NORFOLK 2 Stn DW2027, 23°26.34’S, 167°51.38’E, 465-650 m, 21.X.2003, 2 ♂ 4.9, 5.0 (MNHN-IU-2014-17158). – Stn DW2091, 24°45,36’S, 168°06.24’E, 600-896 m, 29.X.2003, 1 ♂ 3.7 mm (MNHN- IU-2012-690). – Stn DW2101, 23°55.09’S, 167°44.10’S, 700-730 m, 30.X.2003, 1 ♀ 4.0 mm (MNHN-IU-2012-688). – Stn DW2106, 23°53.72’S, 167°41.76’E, 685-757 m, 30.X.2003, 1 ov. ♀ 3.7 mm (MNHN-IU-2012-689). ETYMOLOGY„ From the Greek adiastaltos (not clearly defined), alluding to sternite 3 medially bearing a poorly defined sinus, by which the species is readily distinguished from U. kareenae n. sp., U. depressus n. sp. and U. levicrustus Baba, 1988. DISTRIBUTION„ Isle of Pines and Norfolk Ridge, in 465- 896 m. SIZE„ Males, 2.3-5.0 mm; females, 2.2-4.9 mm; ovigerous females from 2.7 mm. DESCRIPTION — Small species. Carapace: Slightly broader than long (0.9 × as long as broad); greatest breadth 1.7-1.8 × distance between anterolateral spines. Dorsal surface feebly convex from anterior to posterior, without any distinct border; surface nearly smooth, sparingly with short fine setae. Lateral margins divergent posteriorly to point at anterior end of branchial region, then subparallel or slightly divergent posteriorly; ridged along branchial margin; anterolateral spine well developed, overreaching lateral orbital spine; branchial margin with well-developed spine at anterior end, followed by a few to several granulate ridges visible in lateral view, occasionally 1 or 2 very small spines. Rostrum narrow triangular, with interior angle of 22-24°, length usually slightly less than half, rarely about half that of carapace, breadth about one-third to two-fifths carapace breadth measured at posterior carapace margin; dorsal surface slightly concave. Pterygostomian flap anteriorly moderately angular, ending in distinct spine, smooth on surface. Sternum: Excavated sternum broad triangular on anterior margin, with weak ridge in midline; sternal plastron slightly broader than long, lateral extremities divergent posteriorly. Sternite 3 well depressed, anterior margin of broad V-shape medially sharply notched, laterally angular. Sternite 4 having anterolateral margin relatively short, three-quarters as long as posterolateral margin, convex, anteriorly ending in blunt corner; anterolateral margin of sternite 5 convex, about half as long as posterolateral margin of sternite 4. Abdomen: Smooth and nearly glabrous; somite 1 with antero-posteriorly convex transverse ridge. Somite 2 tergite 2.4-2.8 × longer than broad; pleural lateral margins feebly concave and weakly divergent posteriorly. Pleuron of somite 3 posterolaterally blunt angular. Telson half as long as broad or slightly less than so, feebly emarginate or slightly concave on posterior margin, posterior plate 1.2-1.4 × longer than anterior plate. Eye: Relatively short (1.4 × longer than broad), reaching or slightly overreaching midlength of rostrum, medially somewhat swollen; cornea not inflated, more than half length of remaining eyestalk. Antennule and antenna: Ultimate article of antennular peduncle 3.4 × longer than high. Antennal article 2 with small lateral spine; antennal scale reaching distal end of antennal article 5; article 4 usually unarmed, occasionally with tuberclelike mesial distoventral spine; article 5 unarmed, 1.4 × length of article 4, breadth about half height of ultimate article of antennule; flagellum of 9-12 segments far falling short of distal end of P 1 merus. Mxp: Mxp1 with bases separated. Mxp3 basis without denticle on mesial ridge. Ischium with distally rounded flexor margin, no distinct denticle on crista dentata. Merus 2.1 × longer than ischium, flattish on mesial face, weakly ridged along flexor margin, distolateral spine obsolescent; flexor margin unarmed or with obsolescent spine somewhat distal to midlength; carpus with tiny distolateral spine. Pereopods sparsely with soft fine setae. P 1: Slender, 5.5-6.0 × longer than carapace. Ischium dorsally with dorso-ventrally flattened, short spine, ventromesially with strong subterminal spine proximally followed by a few denticles. Merus 1.2-1.3 × longer than carapace, ventromesially with strong or moderate-sized spines: 1 at distal end, 1 at proximal end (these spines not prominent in small specimens), and occasionally additional few small spines between. Carpus 1.4-1.5 × longer than merus. Palm 3.5-4.3 × (males), 5.5- 5.7 × (females) longer than broad, subequal to or slightly shorter than carpus. Fingers slightly incurved distally, gaping in large males, not gaping in females and small males, movable finger 0.4-0.5 × length of palm; opposable margin of movable finger with obtuse process of triangular shape having proximal margin oblique; that of fixed finger with prominence or lower process distal to position of opposite process of movable finger. P 2-4: Meri flattish on mesial face, moderately inflated on lateral face, dorsal margin rounded, successively shorter posteriorly (P 3 merus 0.9 × length of P 2 merus, P 4 merus 0.8-0.9 × length of P 3 merus), subequally broad on 3 and P 4, slightly narrower on P 2, length-breadth ratio, 3.5-3.6 on P 2, 2.9-3.1 on P 3, 2.4-2.6 on P 4; P 2 merus 0.7-0.8 × length of carapace,1.0-1.2 × longer than P 2 propodus; P 3 merus about as long as P 3 propodus; P 4 merus 0.7-0.8 × length of P 4 propodus. Carpi subequal, 0.4 × length of propodus on P 2, 0.3 × on P 3 and P 4. Propodi usually subequal on P 3 and P 4, shorter on P 2, rarely successively longer posteriorly; flexor margin straight, ending in pair of long movable spines preceded by 4, 3-4, 2-3 single spines at most on distal half of length on P 2, P 3, P 4, respectively. Dactyli much longer than carpi (1.3-1.5 ×, 1.6-1.9 ×, 1.8-2.0 × longer on P 2, P 3, P 4, respectively), distinctly more than half (0.6 × on P 2 and P 3, 0.6- 0.7 × on P 4) as long as propodi, well compressed, relatively broad distally; flexor margin straight, with 16-20 (P 2), 18-22 (P 3-4) obliquely directed spines nearly contiguous to one another and obscured by setae, ultimate more slender than antepenultimate, penultimate more than 2 × broader than antepenultimate, remaining spines distinctly broader than ultimate spine except for proximal spines; antepenultimate spine 2.0-2.7 × longer than broad. Eggs. Number of eggs carried, up to 12; size, 0.83 × 0.92 mm - 1.10 × 1.30 mm. PARASITES„ One of the females from BIOCAL Stn DW51 (MNHN-IU-2012-684) bears a rhizocephalan externa on the abdomen. REMARKS„ Uroptychus adiastaltus keys out in a couplet together with U. alius Baba, 2005 (see above under the key to species), having the anteriormost branchial marginal spine not followed by distinct spines. However, the very characteristic sternite 4, having the anterolateral margin much shorter than the posterolateral margin, links U. adiastaltus much more closely to U. depressus n. sp., U. kareenae n. sp. and U. levicrustus Baba, 1988. Their relationships are discussed under the remarks of U. kareenae.Published as part of Baba, Keiji, 2018, Chirostylidae of the Western and Central Pacific: Uroptychus and a new genus (Crustacea: Decapoda: Anomura), pp. 1-612 in Mémoires du Muséum national d'Histoire naturelle 212 on pages 42-45, DOI: 10.5281/zenodo.376097
Heteroptychus claudeae Baba 2018, n. sp.
Heteroptychus claudeae n. sp. Figures 292-294 Uroptychus scambus — Baba 1981: 120; 1988: 43. — Baba et al. 2009: 59 (part), figs 49-51. TYPE MATERIAL — Holotype: Wallis and Futuna Islands. MUSORSTOM 7 Stn CP564, 11°46’S, 178°27’W, 1015-1020 m, with Chrysogorgiidae gen. sp. (Calcaxonia), 20.V.1992, ♂ 5.0 mm (MNHN-IU-2013-8571). Paratypes: New Caledonia, Chesterfield Islands. MUSORSTOM 5 Stn CP387, 20°53.41’S, 160°52.14’E, 650-660 m, with Chrysogorgiidae gen. sp. (Calcaxonia), 22.X.1986, 1 ov. ♀ 4.5 mm (MNHN-IU-2013-8573). Solomon Islands. SALOMON 2 Stn CP2189, 8°19.6’S, 160°01.9’E, 660-854 m, 23.X.2004, 1 ov. ♀ 3.9 mm (MNHN-IU-2014-17105); 2 ♂ 3.8, 4.0 mm, 2 ov. ♀ 3.9, 4.1 mm, 1 ♀ 3.7 mm (MNHN-IU-2014-17106). – Stn DW2190, 08°24’S, 159°27’E, 140-263 m, 24.X.2004, 1 ♀ 4.0 mm (MNHN- IU-2014-17107). – Stn CP2250, 7°29.2’S, 156°16.7’E, 845-970 m, 2.XI.2004, 1 ov. ♀ 4.5 mm (MNHN-IU-2014-17108). – Stn CP2253, 7°26.5’S, 156°15.0’E, 1200-1218 m, 2.XI.2004, 1 ov. ♀ 4.9 mm (MNHN-IU-2014-17109). Wallis and Futuna Islands. MUSORSTOM 7 Stn CP564, collected with holotype, 7 ♂ 4.2-5.5 mm, 8 ov. ♀ 4.2-5.6 mm (MNHN- IU-2014-17110). – Stn CP567, 11°47’S, 178°27’W, 1010-1020 m, 20.V.1992, 4 ♂ 4.6-4.9 mm, 6 ov. ♀ 4.2-5.5 mm (MNHN- IU-2013-12286, MNHN-IU-2013-12284, MNHN-IU-2013-11272), 1 ♂ 5.8 mm, (MNHN-IU-2013-12285). – Stn DW589, 12°16’S, 174°41’W, 400 m, 23.V.1992, 1 ov. ♀ 4.1 mm (MNHN-IU-2014-17111). – Stn CP551, 12°15’S, 177°28’W, 791-795 m, 18.V.1992, 2 ♂ 4.2, 4.3 mm, 1 ov. ♀ 4.3 mm (MNHN-IU-2013-8572). – Stn CP550, 12°15’S, 177°28’W, 800- 810 m, 18.V.1992, 1 ♂ 4.1 mm, 1 ov. ♀ 4.5 mm (MNHN-IU-2014-17112). New Caledonia, Loyalty Ridge. BIOGEOCAL Stn CP297, 20°37’S, 167°11’E, 1230-1240 m, 28.IV.1987, 1 ♂ 3.8 mm (MNHN-IU-2014-17113). MUSORSTOM 6 Stn CP438, 20°23’S, 166°20’E, 800 m, 18.II.1989, 1 ♂ 4.4 mm, 1 ov. ♀ 3.6 mm (MNHN-IU-2014-17114). New Caledonia. HALIPRO 2 Stn BT96, 23°59’S, 161°55’E, 1034-1056 m, 25.XI.1996, 1 ♂ 4.2 mm (MNHN-IU-2014-17115). – Stn BT101, 24°19’S, 161°43’E, 970-1063 m, 26.XI.1996, 1 ♂ 4.8 mm (MNHN-IU-2014-17116). New Caledonia, Norfolk Ridge. BIO- CAL Stn CP31, 23°08’S, 166°51’E, 850 m, 29.VIII.1985, 1 ♂ 3.5 mm (MNHN-IU-2014-17117). – Stn CP32, 23°07’S, 166°51’E, 825 m, 29.VIII.1985, 1 ov. ♀ 3.7 mm (MNHN-IU-2014-17118). – Stn CP54, 23°10’S, 167°43’E, 950-1000 m, 1.IX.1985, 1 ♂ 4.5 mm, 2 ov. ♀ 4.7, 4.8 mm, 1 ♀ 3.8 mm (MNHN-IU-2014-17119). – Stn CP61, 24°11’S, 167°32’E, 1070 m, 2.IX.1985, 1 ♂ 3.4 mm, 3 ov. ♀ 3.7-4.2 mm, 1 sp., sex indet., 2.4 mm (MNHN-IU-2013-12287, MNHN-IU-2013-12288, MNHN-IU-2013-11271). BATHUS 3 Stn CP823, 23°23’S, 167°52’E, 980-1000 m, 29.XI.1993, 1 ♂ 5.0 mm (MNHN- IU-2014-17120). Vanuatu. MUSORSTOM 8 Stn CP992, 18°52.34’S, 168°55.16’E, 775- 748 m, 24.IX.1994, 1 ov. ♀ 4.7 mm (MNHN-IU-2014-17121). – Stn CP1125, 15°57.63’S, 166°38.43’E, 1160-1220 m, 10.X.1994, 1 ov. ♀ 4.4 mm (MNHN- IU-2014-17122). – Stn CP1036, 18°01.00’S, 168°48.20’E, 920-950 m, 29.IX.1994, 1 ♂ 4.6 mm (MNHN-IU-2014-17123). Tonga. BORDAU 2 Stn CP1613, 23°03’S, 175°47’W, 331-352 m, 17.VI.2000, 1 ♂ 4.1 mm (MNHN-IU-2014-17124). – Stn CP1625, 23°28’S, 176°22’W, 824 m, 19.VI.2000, 1 ♀ 4.1 mm (MNHN-IU-2014-17125). ETYMOLOGY„ Named for Claude Crosnier for her hospitality during my stays in Paris. DISTRIBUTION„ Japan, Taiwan, Indonesia, in 1175-1184 m, and now Wallis and Futuna Islands, Chesterfield Islands, Solomon Islands, New Caledonia, Vanuatu, Loyalty Ridge, Norfolk Ridge, and Tonga, in 331-1240 m. SIZE„ Males, 3.4-5.5 mm; females, 3.7-5.6 mm; ovigerous females from 3.7 mm. DESCRIPTION„ Carapace: Broader than long (0.7 × as long as broad), greatest breadth 2.1-2.3 × distance between anterolateral spines. Dorsal surface polished, moderately convex in profile on anterior half, feebly so on posterior half, ridged along posterior half of lateral margin. Lateral margins strongly convex around posterior third; anterolateral spine prominent, horizontally inclined anteromesially, occasionally directed straight forward, not reaching apex of rostrum. Rostrum distally narrow triangular, laterally concave, with interior angle of 20-30°, nearly horizontal or directed somewhat or moderately dorsally, reaching or slightly overreaching tip of eye; length 0.9 × breadth, 0.3 × length of remaining carapace; dorsal surface flattish but feebly concave basally. Lateral limit of orbit unarmed, acuminate or with small spine. Pterygostomian flap anteriorly produced to sharp strong spine; anterior surface well inflated, smooth or with feeble or distinct ridge in lateral midline on anterior portion; posterior height 0.1 × anterior height. Sternum: Excavated sternum with roundly produced anterior margin, surface bearing sharp ridge in midline on anterior half. Sternal plastron in males 0.5-0.6 × as long as broad; in females, left and right parts of sternites 5-7 discontinuous, interrupted by loss of median parts. Sternite 3 having anterior margin broadly and deeply excavated, with small (rarely obsolescent) median notch, with or without small submedian processes. Sternite 4 with rounded anterolateral margin. Sternites 5 and 7 subequal in breadth, sternite 6 broadest. Abdomen: Smooth and polished. Dorsal surface of somite 1 gently convex from anterior to posterior. Somite 2 tergite 2.4- 2.8 × (males), 2.1-2.6 × (females) broader than long; pleural lateral margin slightly concave, strongly divergent posteriorly, ending in blunt tip. Pleura of somites 3-4 somewhat more narrowed in females than in males, ending in rounded margin. Telson 0.4 × as long as broad; posterior plate feebly concave on posterior margin, length 1.4-1.8 × that of anterior plate. Eyes: 1.3-1.4 × as long as broad, subovate. Cornea 0.5-0.7 × length of remaining eyestalk. Antennule and antenna: Ultimate article of antennular peduncle 2.7-2.8 × longer than high. Article 2 of antennal peduncle fused with antennal scale. Antennal scale narrow triangular, reaching or barely reaching midlength of article 4, laterally with or without small spine. Distal 2 articles unarmed; article 5 1.2-1.3 × as long as article 4, breadth less than half height of ultimate antennular article. Flagellum of 9-11 segments. Mxp: Mxp1 with bases broadly separated. Mxp3 basis unarmed on mesial ridge. Ischium relatively short,‰.4 × as long as merus, crista dentata without distinct denticles, flexor margin not rounded distally. Merus relatively thick mesiolaterally, with rounded ridge along flexor margin, unarmed. No spine on carpus. P 1: Smooth and nearly glabrous except for fingers, length 6.3-8.7 × (males), 6.3-6.9 × (females) that of carapace. Ischium with basally broad, blunt distodorsal process (elongate in large specimens) and lobe-like proximal process overhanging basis, unarmed elsewhere. Merus mesially with 1 well-developed distodorsal and 1 small or obsolescent distoventral spine, length 1.6-1.9 × (males), 1.6-1.8 × (females) that of carapace. Carpus 1.4-1.5 × longer than merus, with 2 terminal spines dorsomesially and several obsolescent spines ventromesially. Palm 5.1-5.4 × (males), 3.8-5.9 × (females) longer than broad, 0.8-0.9 × length of carpus, subequally broad medially and distally, proximally narrowed; mesial margin variably ridged from roundly to sharply, more sharply in large specimens, usually with no spine or serration (with obsolescent blunt spines in large specimens). Fingers directed somewhat anterolaterally, gaping in proximal half, straightly fitting to each other in distal half when closed, relatively slender, distally spooned; movable finger‰.5-0.6 × length of palm, opposable margin with prominent blunt process at midlength of gaping portion, distal to position of opposite truncate process on fixed finger. P 2-4: Setose along prehensile margins of propodi and dactyli. Meri subequal on P 2 and P 3, P 4 merus 0.3-0.4 × length of P 3 merus; length-breadth ratio, 5.1-5.8 on P 2, 5.1-5.5 on P 3, 2.5-2.8 on P 4; P 2 merus 0.9 × (1.1 × in largest male MNHN- IU-2013-12285) length of carapace, 1.1-1.2 × length of P 2 propodus, P 3 merus 1.1-1.3 × length of P 3 propodus, P 4 merus 0.6-0.7 × length of P 4 propodus. Carpi subequal in length on P 2 and P 3, P 4 carpus 0.3 × length of P 3 carpus; carpusdactylus length ratio, 1.1-1.2 on P 2 and P 3, 0.4 on P 4. Propodi subequal on P 2 and P 3, P 4 propodus 0.8 × length of P 3 propodus; propodus-dactylus length ratio, 1.4 on P 2, 1.3-1.4 on P 3, 1.2 on P 4; flexor margin concavely curving in lateral view, unarmed. Dactyli subequal on P 2 and P 3, slightly shorter on P 4; dactylus-carpus length ratio, 0.9 on P 2 and P 3, 2.2- 2.6 on P 4; flexor margin with row of 13-16 sharp slender spines proximally diminishing and perpendicular to margin, ultimate somewhat larger than penultimate. Eggs. Number of eggs carried, 20; size, 1.33 × 1.04 mm - 1.33 × 1.08 mm (MNHN-IU-2013-8573); 25 eggs; size, 1.23 × 1.49 mm - 1.41 × 1.67 mm (MNHN-IU-2013-12285). Color. Two specimens in different color of Uroptychus scambus from Taiwan (now referred to H. claudeae; see below) were illustrated by Baba et al. (2009: Figures 49, 50). REMARKS„ The antennal scale is fused with article 2 in H. claudeae, H. lemaitrei n. sp. and the western Atlantic H. brevis (Benedict, 1902), whereas in all the other species it is articulated. Heteroptychus brevis has both the rostrum and the anterolateral spines distally blunt and the P 1 ischium bearing an obsolescent posterior dorsal lobe (see Figure 295), the features differentiating it from H. claudeae and H. lemaitrei. Heteroptychus claudeae differs from H. lemaitrei in having the carapace lateral margin smooth instead of bearing a small but distinct protuberance at the anterior end of branchial margin (see below under the remarks of H. lemaitrei). Unlike all the other species, the presence or absence of lateral orbital spine and the shape of the anterior margin of sternite 3 are unstable in H. claudeae (Figure 294). The specimens reported under U. scambus in my earlier paper (Baba, 1981) from Japan are referable to H. claudeae, as also are the specimens from Taiwan (Baba et al. 2009) except for the male from TAIWAN Stn CD141 (= H. scambus; see below) and the specimens from Station PCP342 (not examined). One of the specimens, the largest of the type series (1 ♂ 5.8 mm, MNHN-IU-2013-12285) collected together with the holotype from Wallis and Futuna, resembles H. lemaitrei in having a spiniform rostrum and in having the P 1 ischium with a spiniform distodorsal spine. Unfortunately, the molecular analyses of this lot were unsuccessful (L. Corbari, personal comm.). Also, one of the Japanese specimens collected at Soyo-Maru Stn B4, 1700 m (Baba, 1981) has a spiniform rostrum but the distodorsal process of the P 1 ischium is distally obtuse, not spiniform. These two specimens are provisionally placed in U. claudeae. The variability noted above, as well as the color difference in Taiwan specimens (Baba et al. 2009: figs 49, 50) suggest that the species may likely contain cryptic species, for which further study is required.Published as part of Baba, Keiji, 2018, Chirostylidae of the Western and Central Pacific: Uroptychus and a new genus (Crustacea: Decapoda: Anomura), pp. 1-612 in Mémoires du Muséum national d'Histoire naturelle 212 on pages 580-586, DOI: 10.5281/zenodo.376097
Munidopsis gibbosa Baba 1978
Munidopsis gibbosa Baba, 1978 Munidopsis gibbosa Baba, 1978: 31, fig. 1, 2 (S of Hong Kong, 19°20.0´N, 114°13.2´E, 952–1025 m). — Komai, 2000: 359 (list). — Baba, 2005: 288 (key, synonymies). Type data: holotype, male, NSMT-Cr. 5655. Type locality: S of Hong Kong, 19°20.0´N, 114°13.2´E, 952–1025 m.Published as part of Baba, Keiji, Macpherson, Enrique, Poore, Gary C. B., Ahyong, Shane T., Bermudez, Adriana, Cabezas, Patricia, Lin, Chia-Wei, Nizinski, Martha, Rodrigues, Celso & Schnabel, Kareen E., 2008, Catalogue of squat lobsters of the world (Crustacea: Decapoda: Anomura-families Chirostylidae, Galatheidae and Kiwaidae), pp. 1-220 in Zootaxa 1905 (1) on page 142, DOI: 10.11646/zootaxa.1905.1.1, http://zenodo.org/record/513458
Sadayoshia acamar Macpherson & Baba, 2012, n. sp.
Sadayoshia acamar n. sp. (Figs. 1, 7 A) Sadayoshia lipkei Macpherson & Baba 2010: 436 (in part, specimens from Chesterfield Islands and some from New Caledonia). Material examined. Holotype: Chesterfield Islands. CHALCAL 1984, Stn DC45, 20º 48.93 'S, 158 º 30.21 'S, 50 m, 23 July 1984: ov. Ƥ 5.1 mm (MNHN-IU- 2010-5377). Paratypes: New Caledonia. Grotte Merlet, 20–30 m, 19–21 January 1993: 2 3 4.7 –5.0 mm, 1 Ƥ 3.1 mm (MNHN-IU- 2010-5383). Chesterfield Islands. CHALCAL 1984, Stn DC45, 20º 48.93 'S, 158 º 30.21 'S, 50 m, 23 July 1984: 2 ov. Ƥ 5.0– 5.1 mm (MNHN-IU- 2010-5378, MNHN-IU- 2010-5379).—Stn DC51, 21º 13.21 'S, 158 º 42.50 'E, 55 m, 24 July 1984: 8 3 3.5–5.4 mm, 1 ov. Ƥ 4.7 mm, 1 Ƥ 4.0 mm (MNHN-IU- 2010-5380, MNHN-IU- 2010-5381). Etymology. The name acamar refers to one of the stars of the southern hemisphere (constellation Eridanus). The name is considered as a substantive in apposition. Description. Carapace: Slightly longer than broad; dorsal surface nearly horizontal from anterior to posterior; anterior cervical groove distinct. Dorsal surface with 4 pairs of epigastric spines, 1 parahepatic, 1 anterior branchial and 1 postcervical spine on each side; ridges with numerous short uniramous and some scattered long iridescent setae. Gastric region with 6 transverse ridges: first uninterrupted, second interrupted into 3 or 4 more or less scalelike ridges, third uninterrupted, fourth medially interrupted, fifth and sixth uninterrupted but short. Mid-transverse ridge uninterrupted, preceded by cervical groove. Posterior branchial region laterally with 5 ridges (excluding midtransverse ridge and posteriormost transverse ridges directly anterior to posterior margin of carapace). Lateral margins slightly or moderately convex, with 7 spines: first anterolateral and well developed; second smaller than first, situated at midlength between anterolateral spine and anterior cervical groove; third to fifth on anterior branchial margin, and sixth and seventh on posterior branchial margin. Rostrum 1.3 times as long as broad, length 0.6 that of carapace, rostral spine about 5.0 times longer than wide (measured at sinus between rostral and anterior lateral spines); dorsal surface convex from side to side, horizontal in profile; lateral margin with 2 spines. Pterygostomian flap rugose with sparse small ridges, ending in acute point. Sternum: Sternal plastron as long as broad, lateral limits divergent posteriorly. Sternite 3 2.3 times as broad as long, anterior margin strongly convex, with small median notch. Sternite 4 1.9 times as long as and 2.7 times broader than preceding sternite, 3.3 times broader than long; surface with 2 medially interrupted, setiferous transverse ridges. Following sternites with 1 or 2 minute ridges. Lateral parts of sternite 7 without granules. Abdomen: Somite 2 unarmed; somites 2–4 each with 2 uninterrupted transverse ridges and 1 additional interrupted transverse ridge between; ridges with numerous short setae and some scatteredlong iridescent setae. Eyes: Ocular peduncles 1.2 times longer than broad; cornea moderately dilated, maximum corneal diameter about 0.4 times distance between bases of anterolateral spines; eyelashes short, not reaching midlength of cornea. Antennule: Article 1 with 4 distal spines: mesial and medioventral spines clearly smaller than others, lateral largest; lateral margin with accesory small spine proximal to distolateral spine; ventral surface with a few short scales. Antenna: Article 1 with distomesial spine reaching distal margin of article 2. Article 2 with distolateral spine terminating in midlength of article 3 and longer than distomesial spine. Article 3 with distomesial spine usually small, sometimes absent. Article 4 unarmed. Mxp 3: Ischium with small distal spine on flexor margin, extensor margin bluntly produced distally; crista dentata with 26–30 denticles. Merus subequal in length to ischium; flexor margin with 2 well-developed spines of subequal size, terminally unarmed or with small spine (in a few specimens); extensor margin unarmed. Carpus unarmed. P 1: 1.9 times carapace length, relatively stout, with sparse long uniramous iridiscent setae along lateral and mesial margins of merus, palm and fingers; dorsal and ventral surfaces of palm and fingers with scattered short setae. Merus 0.7 times length of carapace, 2.5 times length of carpus, with strong distal spines on dorsal and mesial margins, and a few small spines on dorsal surface. Carpus 0.6 times as long as palm, and 0.7 times as long as broad; dorsal surface with a few small spines; mesial margin with 2–3 well-developed spines. Palm 1.1 times longer than broad; with dorsolateral and mesial rows of spines continued on to fingers; dorsal surface with scattered small spines. Fingers as long as palm, distally spooned; 1 or 2 small spines on dorsal proximal part of movable finger. P 2–4: Somewhat compressed mesiolaterally, sparsely with long uniramous and iridescent setae, and short plumose setae. Meri successively shorter posteriorly; P 2 merus 0.8 times carapace length, more than 3.5 times as long as broad, 1.6 times longer than P 2 propodus; P 3 merus more than 3.5 times as long as broad, 1.4 times length of P 3 propodus; P 4 merus 3.5 times as long as broad, 1.2 times length of P 4 propodus; dorsal margins with row of proximally diminishing spines on P 2 and P 3, unarmed on P 4; ventrolateral margins with strong terminal spine followed by proximally diminshing prominences; dorsolateral surfaces unarmed on P 2–4, with 2 spines on P 4. Carpi extensor margins with 3–4 spines on P 2 and P3, 2 spines on P 4, distalmost largest; flexor margins with distal spine; 0–2 small spines on lateral surface. Propodi 3.5 –4.0 times as long as broad; extensor margins with 2 or 3 proximal spines, lateral surface with 1 or 2 proximal spines; flexor margins with 7–10 slender movable spines. Dactyli subequal in length, 0.6–0.8 times length of propodi, ending in incurved, strong, sharp spine; flexor margins with 6 seta-like movable spines, each arising from very low tooth. Remarks. Sadayoshia acamar belongs to a group of species having the antennular article 1 with a lateral marginal spine (= accessory spine) proximal to the distolateral spine and no spine mesial to the anterolateral spine of the carapace. The group contains three species: S. miyakei Baba, 1969, S. lipkei Mapherson & Baba, 2010 and the present new species. Sadayoshia miyakei can be easily distinguished from the other two species by lacking instead of possessing the postcervical spine on each side of the carapace. Sadayoshia acamar and S. lipkei can be distinguished by the number of spines along the flexor margin of Mxp 3 merus: 3 three spines in S. lipkei and 2 two in S. acamar. However, the median spine can be very small in some specimens of S. lipkei, and a third small distal spine can be observed in a few specimens of S. acamar. Furthermore, P 2–4 are slightly more slender in S. acamar than in S. lipkei. For instance, the P 2–3 meri are 3.2–3.3 times as long as broad in S. lipkei, instead of being more than 3.5 times longer as in the new species. Distribution. New Caledonia, and Chesterfield Islands, at 20– 55 m.Published as part of Macpherson, Enrique & Baba, Keiji, 2012, The squat lobsters of the genus Sadayoshia Baba, 1969 (Crustacea: Decapoda: Anomura: Munididae): new records including six new species from the Pacific Ocean, pp. 30-48 in Zootaxa 3589 on pages 31-34, DOI: 10.5281/zenodo.21464
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