11 research outputs found
Sciocoris (Parasciocoris) angusticollis Puton 1895
Sciocoris (Parasciocoris) angusticollis Puton, 1895 Sciocoris angusticollis Puton, 1895: 83. Distribution in Iran. Khorasan (Nateq Golestan & Modarres Awal 2012). General distribution. North Africa, Near East (Israel, Syria). Host plants. Under Artemisia sp. (Asteraceae) (Nateq Golestan & Modarres Awal 2012). Comments. Cited from France and Syria by Rider (2006), but not by Derjanschi & Péricart (2005), neither author cites this species from Iran. Dusoulier & Lupoli (2006) exclude this taxon from the French fauna.Published as part of Ghahari, Hassan, Moulet, Pierre & Rider, David A., 2014, An annotated catalog of the Iranian Pentatomoidea (Hemiptera: Heteroptera: Pentatomomorpha), pp. 1-95 in Zootaxa 3837 (1) on page 52, DOI: 10.11646/zootaxa.3837.1.1, http://zenodo.org/record/28672
Homoeocerus albiguttulus Stal 1873
Homoeocerus albiguttulus Stål, 1873 Homoeocerus albiguttulus Stål, 1873: 61 (as new species). Material examined. Sistan & Baluchestan: Iranshahr (538 m), 2 Ƥ, 13, April 2002. New record for Iran. General distribution. Bangladesh, Far East, India, Malaysia, Pakistan, Sumatra (Perveen 1991). Comments. Moreover, one damaged specimen of Homoeocerus sp. was found from Saravan (Sistan & Baluchestan province). On the basis of preliminary identification by the first author and also I.M. Kerzhner, it is probably H. lacertosus Distant 1889 with distribution in India and Pakistan. Because Sistan & Baluchestan province is a vast region in southeastern Iran and especially near of Pakistan, such record cannot be excluded.Published as part of Ghahari, Hassan, Moulet, Pierre, Linnavuori, Rauno E. & Ostovan, Hadi, 2012, An Annotated Catalog of the Iranian Coreidae, Rhopalidae, and Stenocephalidae (Hemiptera: Heteroptera: Pentatomomorpha: Coreoidea), pp. 1-31 in Zootaxa 3519 on pages 9-10, DOI: 10.5281/zenodo.21229
Carabid beetles (Coleoptera: Carabidae) collected from different ecosystems in Iran with new records
The Carabidae (Coleoptera) of apple orchards and alfalfa fields in seventeen agricultural sites of ten provinces of Iran have been investigated. 43 species/subspecies collected from these regions have been listed. Seven of the species are new records for Iran and two species are second find for the Palaearctic Region. The geographical distribution and chorotype data are given for all species.Shahre Rey Islamic Azad UniversityThe authors are indebted to Dr. Borislav Gueorguiev (Bulgaria) for his valuable help and advice. They also thank M. Tabari (Iran Rice Research Institute), H. Sakenin (Ghaemshahr Islamic Azad University), H. Rakhshani (Isfahan University of Technology) and many agricultural students of first author for collecting some specimens. The research was supported by Shahre Rey Islamic Azad University
Chalcophorella Kerremans 1903
Genus CHALCOPHORELLA Kerremans, 1903 Chalcophorella bagdadensis (Laporte & Gory) 1836: 125 (Buprestis); Théry 1925: 79; Kubáň 2006: 345; Bellamy 2008 a: 551; Abaei 2014: 691 (author: Bily [sic!]). synonym: amarensis Obenberger 1942: 8 (subspecies of bagdadensis). synonym: aureoscripta Mandl & Pochon 1957: 326. synonym: berhauti Mandl & Pochon 1957: 325. [subgenus Stigmatophorella]. Distribution within Iran. "Chaldée persane: Kouh-Hadjoudj, Neima; Poucht-é-Kouh: Tagh-i-Mowla" (Théry 1925). Fars (Saghaei et al. 2010), Khuzestan (Théry 1925), Zagros area (Abaei 2014), Iran (no locality cited) (Kubáň 2006; Bellamy 2008 a; Ali 2010). Distribution outside Iran. Armenia, Iraq, Syria, Turkey (Kubáň 2006; Bellamy 2008 a; Ali 2010). Note. A robber fly species, Stenopogon junceus (Wiedemann in Meigen, 1820) (Diptera: Asilidiae) is the predator of C. bagdadensis (Saghaei et al. 2010). Chalcophorella bagdadensis freyi Obenberger 1942: 8 (subspecies of bagdadensis); Radjabi 1968: 71, 1976: 128; Bílý 1983 a: 53; Modarres Awal 1997: 134 (as bagdanensis frey); Borumand 2002: 29; Kubáň 2006: 345; Bellamy 2008 a: 551. Host plant(s) within Iran. Wild and cultivated almond (Amygdalus spp.; Rosaceae) (Modarres Awal 1997). Distribution within Iran. Bushehr, Kuhgiloyeh & Boyerahmad (Borumand 2002), Fars (Radjabi 1976; Bílý 1983 a; Modarres Awal 1997; Borumand 2002), Kerman, Kermanshah, Khuzestan, Ilam (Radjabi 1968; Modarres Awal 1997; Borumand 2002), Tehran (Modarres Awal 1997), Iran (no locality cited) (Kubáň 2006; Bellamy 2008 a). Distribution outside Iran. Iraq (Bellamy 2008 a). Chalcophorella escalerae (Abeille de Perrin) 1904: 212 (Buprestis); Richter 1952: 197 (abberation of bagdadensis); Radjabi 1968: 71, 1976: 128; Modarres Awal 1997: 134; Borumand 2002: 29; Kubáň 2006: 345; Bellamy 2008 a: 551; Ghobari et al. 2012: 236. [subgenus Stigmatophorella]. Host plant(s) within Iran. Wild almond, white-thorn (Amygdalus sp., Crataegus sp., Rosaceae) (Modarres Awal 1997), Crataegus sp. (Rosaceae) (Borumand 2002). Distribution within Iran. Fars (Radjabi 1968, 1976; Modarres Awal 1997; Borumand 2002), Lorestan (Modarres Awal 1997; Borumand 2002), Kuhgiloyeh & Boyerahmad ("Valee du Chindaar, dans le Haut Karoum") (Abeille de Perrin 1904; Radjabi 1976; Borumand 2002), Yazd (Radjabi 1976), Kordestan (Ghobari et al. 2012), Iran (no locality cited) (Richter 1952; Kubáň 2006; Bellamy 2008 a). Distribution outside Iran. None recorded. * Chalcophorella fabricii (Rossi) 1792: 100 (Buprestis); Théry 1925: 79; Kubáň 2006: 345; Bellamy 2008 a: 551. [subgenus Rossiella]. Immatures: Gory & Laporte 1840: 2; Bertoloni 1842: 87; Pecchioli 1843: 2; Perris 1877: 131. Biology/Bionomy: Gory & Laporte 1839: 2; Bertoloni 1842: 89; Pecchioli 1843: 2; Bertoloni 1872: 351; Xambeu 1892: 225; Berlese 1924: 184. Distribution within Iran. "Poucht-é-Kouh: Mollah-Giavan" (Théry 1925). Iran (no locality cited) (Bellamy 2008 a). Distribution outside Iran. Albania, Bosnia-Herzegovina, Bulgaria, Croatia, Greece, Iraq, Lebanon, Italy, Macedonia, Syria, Turkey, former Yugoslavia (Kubáň 2006; Bellamy 2008 a). Note. The occurrence of this species in Iran needs confirmation. Chalcophorella morgani Théry 1925: 79 (Chalcophorella), 171 (Chalcophora); Kubáň 2006: 345; Bellamy 2008 a: 552. [subgenus Chalcophorella]. Distribution within Iran. Lorestan (Théry 1925), Iran (no locality cited) (Kubáň 2006; Bellamy 2008 a). Distribution outside Iran. Iraq, Turkey (Kubáň 2006; Bellamy 2008 a). Chalcophorella orientalis (Obenberger) 1924 c: 14 (Chalcophorella); Kubáň 2006: 345, Bellamy 2008 a: 552; Ghobari et al. 2012: 236. [subgenus Stigmatophorella]. Distribution within Iran. Kordestan (Ghobari et al. 2012), Sistan & Baluchestan (Obenberger 1924 c), Iran (no locality cited) (Kubáň 2006; Bellamy 2008 a). Distribution outside Iran. Afghanistan, Pakistan (Kubáň 2006; Bellamy 2008 a). Chalcophorella quadrioculata (Redtenbacher) 1843: 993 (Chalcophora); Kubáň 2006: 345; synonym: akbesiana Cobos 1957: 200 (subspecies of quadrioculata); Kubáň 2006: 46 (synonym of quadrioculata), 345; Bellamy 2008 a: 552. [subgenus Stigmatophorella]. Distribution within Iran. Iran (no locality cited) (Kubáň 2006; Bellamy 2008 a). Distribution outside Iran. Georgia, Iraq, Syria, Turkey (Kubáň 2006; Bellamy 2008 a). Chalcophorella stigmatica (Dalman in Schönherr) 1817: 119 (Buprestis); Théry 1925: 79; Richter 1952: 195; Davatchi et al. 1959: 245; Farahbakhsh 1960: 73; Radjabi 1974: 38; Abai & Adeli 1984: 19; Modarres Awal 1997: 134; Borumand 2002: 30; Kubáň 2006: 345; Bellamy 2008 a: 552; Ghobari et al. 2012: 236. synonym: quadrinotata (Klug) 1829: No. 20 (Buprestis). synonym: lefebvrei (Laporte & Gory) 1836: 17 (Buprestis). synonym: quadrimaculata (Redtenbacher) 1850: 47 (Chalcophora). synonym: marseuli (Garbiglietti) 1867: lxvii (Buprestis); Modarres Awal 1997: 134 (subspecies of stigmatica, author Obenberger). synonym: balcanica Obenberger 1936 c: 175 (subspecies of stigmatica). [subgenus Chalcophorella]. Host plant(s) within Iran. Amygdalus spp. (Rosaceae) (Davatchi et al. 1959; Abai & Adeli 1984; Modarres Awal 1997), wild almond (Amygdalus sp.) (Farahbakhsh 1960), Quercus sp. (Fagaceae) (Borumand 2002). Distribution within Iran. East Azarbayjan, Markazi (Modarres Awal 1997),? Fars (Modarres Awal 1997, as stigmatica marseuli Obenberger), Isfahan and central provinces (Abai & Adeli 1984; Modarres Awal 1997), Kermanshah (Radjabi 1974; Modarres Awal 1997; Borumand 2002), Khuzestan (Théry 1925), Kordestan (Ghobari et al. 2012), Iran (no locality cited) (Richter 1952; Davatchi et al. 1959; Kubáň 2006; Bellamy 2008 a). Distribution outside Iran. Albania, Bosnia-Herzogovina, Bulgaria, Cyprus, Croatia, Egypt, Greece, Iraq, Israel, Italy (Sardinia), Jordan, Lebanon, Macedonia, Syria, Turkey (Kubáň 2006; Bellamy 2008 a).Published as part of Ghahari, Hassan, Volkovitsh, Mark G. & Bellamy, Charles L., 2015, An annotated catalogue of the Buprestidae of Iran (Coleoptera: Buprestoidea), pp. 1-141 in Zootaxa 3984 (1) on pages 34-35, DOI: 10.11646/zootaxa.3984.1.1, http://zenodo.org/record/24515
Agrilus Curtis 1825
Genus AGRILUS Curtis, 1825 synonym: Paradomorphus Waterhouse 1887. synonym: Samboides Kerremans 1900. synonym: Sarawakita Obenberger 1924. synonym: Callichitones Obenberger 1931. synonym: Therysambus Descarpentries & Villiers 1967. synonym: Wallaceilus Holynski 2003. Agrilus albogularis Gory 1841: 235; Jendek 2006: 395; Bellamy 2008 d: 1956; Jendek & Grebennikov 2011: 235. synonym: sieversi Abeille de Perrin 1897: 28 (variety of zigzag Marseul 1866); Jendek 2006: 396 (synonym of zigzag); Bellamy 2008 d: 2367 (synonym of zigzag); Jendek & Grebennikov 2011: 235 (synonym of albogularis). [subgenus Xeragrilus; albogularis species group]. Immatures: Alexeev 1960: 1509, 1961: 9, 1981 b: 79; Bílý 1999: 37. Biology/Bionomy: Bílý 2003: 10. Distribution within Iran. Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Albania, Armenia, Austria, Azerbaijan, Belarus, Bosnia-Herzegovina, Bulgaria, Czech Republic, France, Georgia, Germany, Greece, Hungary,? Iraq, Italy, Kazakhstan,? Kyrgyzstan, Macedonia, Moldavia, Portugal, Romania, Russia, Slovakia, Slovenia, Spain, Turkey, Turkmenistan, Ukraine (Jendek 2006; Bellamy 2008 d). Agrilus angustulus (Illiger) 1803: 240 (Buprestis); Jendek 2006: 391; Bellamy 2008 d: 1965. synonym: olivaceus (Gyllenhal) 1808: 454 (Buprestis). synonym: laetefrons Mannerheim 1837: 114. synonym: rugicollis (Ratzeburg) 1837: 55 (Buprestis). synonym: laeticeps Semenov-Tian-Shankij 1891: 342. synonym: semenovi Kerremans 1892: 271 (nomen novum for laeticeps Semenov-Tian-Shankij 1890, superfluous name). [subgenus Quercuagrilus; sulcicollis species group]. Immatures: Ratzeburg 1837: 54; Perris 1875: 385, 1877: 127; Alexeev 1964 b: 20, 1981 b: 73; Molino-Olmedo 1998 b: 29; Bílý 1999: 35. Biology/Bionomy: Bílý 2003: 11. Distribution within Iran. Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Albania, Algeria, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bosnia- Herzegovina, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Georgia, Germany, Great Britain, Georgia, Greece, Hungary, Italy, Kazakhstan, Latvia, Lichtenstein, Lithuania, Luxembourg Macedonia, Moldavia, Montenegro, Morocco, the Netherlands, Norway, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Syria, Tunisia, Turkey, Ukraine (Jendek 2006; Bellamy 2008 d). Agrilus biguttatus (Fabricius) 1777: 237 (Buprestis); Davatchi et al. 1959: 247; Farahbakhsh 1960: 71; Adeli 1972: 10; Abai & Adeli 1984: 89; Modarres Awal 1997: 132; Jendek 2006: 390; Bellamy 2008 d: 1997; Barimani et al. 2009: 66, 2010: 131. synonym: pannonicus (Piller & Mitterpacher) 1783: 37 (Buprestis). synonym: subfasciatus (Ménétriés) 1832: 153 (Buprestis). synonym: morosus Gory & Laporte 1837: 35. [subgenus Anambus; biguttatus species group]. Immatures: Ratzeburg 1837: 57; Goureau 1843: 23; Schiødte 1870: 360, 374; Alexeev 1960: 1506, 1961: 5, 1981 b: 68; Bílý 1999: 33. Biology/Bionomy: Bílý 2003: 13. Host plant(s) within Iran. Oak (Quercus sp., Fagaceae) (Davatchi et al. 1959; Farahbakhsh 1960; Adeli 1972; Abai & Adeli 1984, Modarres Awal 1997, dry wood). Distribution within Iran. Mazandaran (Barimani et al. 2009, 2010), northern provinces (Abai & Adeli 1984; Modarres Awal 1997), Iran (no locality cited) (Davatchi et al. 1959; Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Albania, Algeria, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bosnia- Herzegovina, Bulgaria, Czech Republic, Croatia,? Denmark, Estonia, France, Great Britain, Germany, Greece, Hungary, Italy, Kazakhstan, Latvia, Lithuania, Luxembourg, Macedonia, Moldavia, Morocco, the Netherlands,? Norway, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Slovenia, Spain, Sweden, Syria, Switerland Turkey, Ukraine (Jendek 2006; Bellamy 2008 d). Agrilus chlorophyllus Abeille de Perrin 1904: 223; Jendek 2006: 397; Bellamy 2008 d: 2025; Jendek & Grebennikov 2011: 59. synonym: esfandiarinus Davatchi 1958: 68 (subspecies of viridicaerulans); Davatchi et al. 1959: 248, Salavatian 1950: 11 (subspecies of viridicaerulans; author Obenberger nomen nudum); Farahbakhsh 1960: 71 (subspecies of viridicaerulans); Radjabi 1968: 70, 1976: 128; Adeli 1972: 10 (subspecies of viridicaerulans); Abai & Adeli 1984: 66 (subspecies of viridicerulans); Modarres Awal 1997: 132 (subspecies of viridicaerulans); Abivardi 2001: 675, 678; Borumand 2002: 50 (subspecies of viridicaerulans). synonym: pistaciophagus Alexeev & Kulinitsh 1962: 92; Jendek 2006: 394; Bellamy 2008 d: 2237; Jendek & Grebennikov 2011: 59 (synonym of chlorophyllus). [subgenus Rosagrilus; roscidus species group]. Immatures: Davatchi 1958: 68, Davatchi et al. 1959: 248 (as viridicaerulans esfandiarinus); Alexeev & Kulinitsh 1962: 94 (as pistaciophagus). Biology/Bionomy: Davatchi 1958: 68; Balachowsky et al. 1962: 298 (as viridicaerulans esfandiarinus). Host plant(s) within Iran. Pistacia sp. (Anacardiaceae) (as viridicaerulans esfandiarinus Obenberger) (Salavatian 1950; Davatchi 1958; Davatchi et al. 1959, Salavatian 1950; Adeli 1972; Abai & Adeli 1984; Modarres Awal 1997); Pistacia mutica (as viridicaerulans esfandiarinus) (Davatchi 1958; Farahbakhsh 1960; Balachowsky et al. 1962); P. vera (Farahbakhsh 1960, as viridicaerulans esfandiarinus). Distribution within Iran. Fars (as viridicaerulans esfandiarinus) (Farahbakhsh 1960; Radjabi 1968, 1976; Adeli 1972; Abai & Adeli 1984; Modarres Awal 1997; Borumand 2002; Jendek & Grebennikov 2011, as chlorophyllus), Kerman (as viridicaerulans esfandiarinus) (Davatchi et al. 1959; Farahbakhsh 1960; Radjabi 1968; Adeli 1972; Abai & Adeli 1984; Modarres Awal 1997), Kordestan, Zanjan and most other pistachio planting areas (as viridicaerulans esfandiarinus) (Modarres Awal 1997), Qazvin (as viridicaerulans esfandiarinus) (Salavatian 1950; Davatchi et al. 1959; Farahbakhsh 1960; Radjabi 1968; Borumand 2002), central Iran (as viridicaerulans esfandiarinus) (Adeli 1972; Abai & Adeli 1984), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Kyrgyzstan, Israel, Syria, Tajikistan, Turkey, Turkmenistan, Uzbekistan (Jendek & Grebennikov 2011). Agrilus cuprescens (Ménétriés) 1832: 154 (Buprestis); Jendek 2006: 388; Bellamy 2008 d: 2046; Jendek & Grebennikov 2011: 65. synonym: aurichalceus Redtenbacher 1849: 286. synonym: proximus Rey 1891 b: 19. synonym: egenus Abeille de Perrin 1895 b: cxxii. synonym: lacrymans Abeille de Perrin 1895 d: cclxxiv. synonym: epistomalis Abeille de Perrin 1897: 13. synonym: foveolatus Abeille de Perrin 1897: 15. synonym: obtusus Abeille de Perrin 1897: 15. synonym: rubicola Abeille de Perrin 1897: 15. synonym: calcicola Obenberger 1916: 271. synonym: krasai Obenberger 1924 c: 40. synonym: communis Obenberger 1924 c: 41. synonym: mokrzeckii Obenberger 1927g: 204. synonym: altaicola Obenberger 1935 c: 168. synonym: kuznecovi Obenberger 1935 c: 169. synonym: kuznecovinus Obenberger 1936 a: 1038. synonym: obtusifera Schaefer 1949: 402. [subgenus Agrilus; viridis species group]. Immatures: Alexeev 1960: 1508 (as chrysoderes), 1961: 7, 1981 b: 76; Bílý 1999: 34 (as aurichalceus). Biology/Bionomy: Bílý 2003: 15. Distribution within Iran. Guilan (Jendek & Grebennikov 2011). Distribution outside Iran. Albania, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, China, Croatia, Czech Republic, Estonia, Finland, France, Georgia, Germany, Great Britain, Greece, Hungary, Italy, Japan, Kazakhstan, Kyrgyzstan, Latvia, Lichtenstein, Lithuania, Luxembourg, Macedonia, Moldavia, Mongolia, the Netherlands, North America (introduced), Poland, Portugal, Romania, Russia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Syria, Turkey, Turkmenistan, Ukraine, former Yugoslavia (Jendek & Grebennikov 2011). Agrilus derasofasciatus Lacordaire in Boisduval & Lacordaire 1835: 613; Davatchi 1958: 69; Jendek 2006: 391; Bellamy 2008 d: 2064; Barimani et al. 2009: 67, 2010: 131. synonym: hungerbuehleri Pochon 1963: 69. [subgenus Quercuagrilus; sulcicollis species group]. Immatures: Perris 1851 teste Perris 1877: 132; Alexeev 1964 b: 22, 1981 b: 72; Bílý 1999: 35. Biology/Bionomy: Bílý 2003: 17. Distribution within Iran. Mazandaran (Barimani et al. 2009, 2010), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Albania, Algeria, Armenia, Austria, Azerbaijan, Belarus, Bosnia-Herzegovina, Bulgaria, Croatia,? Cyprus, Czech Republic, Egypt, France, Georgia, Germany, Greece, Hungary, Israel, Italy, Lebanon, Macedonia, Madeira, Malta, Moldavia, Montenegro, Morocco, Poland, Portugal, Romania, Russia, Slovakia, Slovenia, Spain, Syria, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine; introduced: U.S.A. (Jendek 2006; Bellamy 2008 d). Agrilus desertus (Klug) 1829: No. 14 (Buprestis); Bílý 1983 a: 84; Jendek 2006: 391; Bellamy 2008 d: 2066; Bílý et al. 2011: 211. [subgenus Diplolophotus]. Host plant(s) within Iran. Acacia flava (Fabaceae) (larva) (Bílý 1983 a). Distribution within Iran. Hormozgan (Bílý 1983 a), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d, Bílý et al. 2011). Distribution outside Iran. Algeria, Chad, Djibouti, Egypt, Israel, Jordan, Lybia, Morocco, Saudi Arabia, Tunisia, UAE, Yemen (Jendek 2006; Bellamy 2008 d; Bílý et al. 2011). Agrilus graminis Kiesenwetter 1857: 137; Jendek 2006: 392; Bellamy 2008 d: 2112; Barimani et al. 2009: 67, 2010: 131. synonym: verticalis Abeille de Perrin 1895 b: cxxvi. synonym: reitteri Abeille de Perrin 1897: 16. synonym: barbatus Abeille de Perrin 1897: 34; Jendek 2006: 392 (synonym of hastulifer); 2007: 90 (synonym of graminis, lectotype). synonym: cervicornis Abeille de Perrin 1897: 35; Jendek 2006: 391; 2007: 90 (synonym of graminis). synonym: talyschensis Obenberger 1917 c: 215; Jendek 2006: 402, 2007: 90 (synonym of graminis, lectotype). synonym: disparicornis Bedel 1921: 216 (nomen novum for graminis Gory & Laporte 1837). synonym: akbesinus Obenberger 1924 c: 50 (variety of disparicornis). synonym: mamorensis Théry 1930 b: 446 (subspecies of disparicornis); Jendek 2006: 392 (subspecies of graminis); 2007: 90 (synonym of graminis, lectotype). synonym: armenus Obenberger 1936 b: 106; Jendek 2006: 396, 2007: 90 (synonym of graminis, lectotype). [subgenus Quercuagrilus; sulcicollis species group]. Immatures: Xambeu 1903: 79; Alexeev 1960: 1509, 1961: 8, 1981 b: 74; Bílý 1999: 35. Biology/Bionomy: Bílý 2003: 17. Distribution within Iran. Mazandaran (Barimani et al. 2009, 2010), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Albania, Algeria, Armenia, Austria, Azerbaijan, Belarus, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Republic, France, Georgia, Germany, Greece, Hungary, Italy, Macedonia, Moldavia, Montenegro, Morocco, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Slovenia, Spain, Switzerland, Syria, Tunisia, Turkey, Ukraine (Jendek 2006; Bellamy 2008 d). Agrilus hastulifer (Ratzeburg) 1837: 54 (Buprestis), 1839 a: 9; Davatchi et al. 1959: 247; Farahbakhsh 1960: 71; Adeli 1972: 10; Modarres Awal 1997: 132; Jendek 2006: 392; Bellamy 2008 d: 2122; Barimani et al. 2009: 67; Abaei 2014: 691 (as A. lustifer Rhet. [sic!]). synonym: aladaghensis Obenberger 1918 b: 24; Bílý 1983 a: 83 (subspecies of hastulifer); Borumand 2002: 50; Jendek 2006: 392 (subspecies of hastulifer); 2007: 90 (synonym of hastulifer, lectotype). [subgenus Quercuagrilus; sulcicollis species group]. Immatures: Perris 1877: 147; Zverezomb-Zubovskii 1914: 19; Alexeev 1964 b: 22, 1981 b: 74; Bílý 1999: 35. Biology/Bionomy: Bílý 2003: 18, 19. Host plant(s) within Iran. Qurcus sp., Fagus sp. (Fagaceae) (Davatchi et al. 1959); Quercus sp., Alnus sp., Carpinus sp. (Betulaceae) (Adeli 1972); Alder (Alnus sp., Betulaceae), Oak (Quercus sp.) (Farahbakhsh 1960; Modarres Awal 1997). Distribution within Iran. Kuhgiloyeh & Boyerahmad (Borumand 2002), Mazandaran (as aladaghensis) (Bílý 1983 a; Barimani et al. 2009), Northern Iran (Davatchi et al. 1959), Kaspische Wälder (Adeli 1972), Zagros area (Abaei 2014), Iran (no locality cited) (Modarres Awal 1997; Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Albania, Algeria, Armenia, Austria, Azerbaijan, Belarus, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Republic, France,? Georgia, Germany, Greece, Hungary, Iraq, Israel, Italy, Kazakhstan, Macedonia, Montenegro, Morocco, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Slovenia, Spain, Switzerland, Syria, Tunisia, Turkey, Ukraine (Jendek 2006; Bellamy 2008 d). Agrilus hyperici (Creutzer) 1799: 122 (Buprestis); Bílý 1983 a: 84; Jendek 2006: 395; Bellamy 2008 d: 2129; Barimani et al. 2009: 67, 2010: 131. synonym: prasinus Mulsant & Rey 1863 a, b: 17; Jendek 2006: 395 (as species); 2007: 90 (synonym of hyperici). synonym: impressus Rey 1891 b: 20 (preoccupied by impressus Chevrolat 1835). synonym: modestulus Semenov-Tian-Shankij 1895 b: 247. synonym: albaniae Obenberger 1922: 66. synonym: trinacriae Obenberger 1924 c: 52; Jendek 2006: 402, 2007: 90 (synonym of hyperici). synonym: reyellus Obenberger 1936 a: 1000 (nomen novum for impressus Rey 1891). synonym: hofferi Obenberger 1936 b: 111. [subgenus Spiragrilus; hyperici species group]. Immatures: Perris 1875: 406; 1877: 148; Alexeev 1964 b: 24, 1981 b: 75. Biology/Bionomy: Bílý 2003: 19. Distribution within Iran. Golestan (Bílý 1983 a), Mazandaran (Barimani et al. 2009, 2010), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Albania, Armenia, Australia (introduced), Austria, Azerbaijan, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Republic,? Denmark, Estonia, France, Georgia, Germany, Greece, Hungary, Italy, Kazakhstan, Latvia, Lithuania, Macedonia, Moldavia, Montenegro, the Netherlands, Poland, Portugal, Romania, Russia, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine, USA (introduced) (Jendek 2006; Bellamy 2008 d). Agrilus lineola Kiesenwetter 1857: 147; Jendek 2006: 389; Bellamy 2008 d: 2166. synonym: hermineus Abeille de Perrin 1907 b: li; Bílý 1983 a: 84; Abai & Adeli 1984: 26; Modarres Awal 1997: 132; Borumand 2002: 50. synonym: rumanicus Obenberger 1924 c: 48. [subgenus Agrilus; viridis species group]. Immatures: Alexeev 1961: 8, 1960: 1507, 1981 b: 76; Bílý 1999: 34. Biology/Bionomy: Bílý 2003: 21. Host plant(s) within Iran. Hornbeam (Carpinus sp., Betulaceae), sapwood (Abai & Adeli 1984; Modarres Awal 1997, as hermineus). Distribution within Iran. Khorasan (Bílý 1983 a, as hermineus; Borumand 2002), Mazandaran, Qazvin (Borumand 2002), northern provinces (Abai & Adeli 1984; Modarres Awal 1997, as hermineus). Distribution outside Iran. Albania, Austria, Bosnia & Herzegovina, Bulgaria, Belarus, Croatia, Czech Republic, France, Germany, Greece, Hungary, Italy, Kazakhstan, Macedonia, Moldavia, Poland, Romania, Russia, Slovakia, Slovenia, Switzerland, Turkey, Ukraine, former Yugoslavia (Jendek 2006; Bellamy 2008 d). Note. Records for A. hermineus in Iran probably refer to A. lineola shamyl Obenberger, 1922 occurring in adjacent areas of Armenia and Turkmenistan (Volkovitsh & Alexeev 1994). Records for Carpinus as a host plant in Abai & Adeli (1984) and Modarres Awal (1997) are most probably based on misidentification; A. lineola is associated with Salix spp. (Salicaceae). Agrilus lituratus (Klug) 1829: No. 13 (Buprestis); Bílý 1983 a: 83; Jendek 2006: 391; Bellamy 2008 d: 2168; Bílý et al. 2011: 212. synonym: kiesenwetteri Pic 1894: 71. [subgenus Micragrilus]. Immatures: Mateu 1974: 281. Biology/Bionomy: Mateu 1974: 283. Host plant(s) within Iran. (Adult) Acacia flava (Fabaceae) (Bílý 1983 a). Distribution within Iran. Bushehr, Hormozgan, Sistan & Baluchestan (Bílý 1983 a), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d; Bílý et al. 2011). Distribution outside Iran. Algeria, Chad, Egypt, Israel, Jordan, Libya, Mauritania, Morocco, Saudi Arabia, Senegal, Sudan, Tunisia, UAE, Yemen (Jendek 2006; Bellamy 2008 d; Bílý et al. 2011). * Agrilus lopatini Alexeev 1964 a: 11; Bílý 1983 a: 84; Jendek 2006: 395; Bellamy 2008 d: 2170. [subgenus Spiragrilus]. Distribution within Iran. Kuhgiloyeh & Boyerahmad (Bílý 1983 a), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Kazakhstan, Tajikistan (Jendek 2006). Note. The records for Iran probably refer to A. araxenus Iablokoff-Khnzorian, 1960 occurring in adjacent areas of Transcaucasia and Turkmenistan; the records for Georgia and Turkmenistan in Bellamy (2008 d) are also possibly refer to the same species. Agrilus obscuricollis Kieswetter 1857: 144; Jendek 2006: 392; Bellamy 2008 d: 2208; Barimani et al. 2010: 131; Jendek & Grebennikov 2011: 143. synonym: foveola Rey 1891 b: 19. synonym: reyi Bauduer 1878: 83. synonym: savalae Stark 1953: 300. [subgenus Quercuagrilus; sulcicollis species group]. Biology/Bionomy: Bílý 2003: 22. Distribution within Iran. Mazandaran (Barimani et al. 2010), Iran (no locality cited) (Jendek & Grebennikov 2011). Distribution outside Iran. Albania, Algeria, Armenia, Austria, Azerbaijan, Belarus, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Republic, France, Germany, Greece, Hungary, Israel, Italy, Kazakhstan, Malta, Macedonia, Moldavia, the Netherlands, Poland, Romania, Russia, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine, former Yugoslavia (Jendek 2006; Jendek & Grebennikov 2011). Agrilus pecirkai Obenberger 1916: 273; Jendek 2006: 389; Bellamy 2008 d: 2228; Ghahari et al. 2008: 638; Jendek & Grebennikov 2011: 147; Ghahari & Bellamy 2012: 41. [subgenus Agrilus; viridis species group]. Biology/Bionomy: Makhnovskij 1959: 32 (as chrysoderes); Kadyrov 1976: 52 (as pecsikoi). Host plant(s) within Iran. Oak, Quercus rotundifolia (Fagaceae) (Ghahari et al. 2008), Prunus persica Ghahari & Bellamy 2012), Rosa sp. (Rosaceae) (Jendek & Grebennikov 2011). Distribution within Iran. Golestan (Ghahari & Bellamy 2012), Khorasan (Ghahari et al. 2008), Iran (no locality cited) (Jendek & Grebennikov 2011). Distribution outside Iran. Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan (Jendek 2006; Jendek & Grebennikov 2011). Note. Some records for the host plants are probably based on misidentification. This species have never been recorded from Quercus spp. or Prunus spp. outside Iran. Agrilus pratensis (Ratzeburg) 1837: 56 (Buprestis); Jendek 2006: 393; Bellamy 2008 d: 2243; Barimani et al. 2009: 67, 2010: 131; Jendek & Grebennikov 2011: 158. synonym: robertii Chevrolat 1838 a: 89. synonym: pseudocoeruleus Obenberger 1930 a: 112. synonym: praeclarus Krogerus 1925: 75 (variety of robertii); Jendek 2006: 393 (subspecies of pratensis); Bellamy 2008 d: 2245 (subspecies of pratensis); Jendek & Grebennikov 2011: 159 (synonym of pratensis). synonym: djukini Obenberger 1935 c: 169 (subspecies of robertii); Jendek 2006: 393 (subspecies of pratensis); Bellamy 2008 d: 2244 (subspecies of pratensis); Jendek & Grebennikov 2011: 159 (synonym of pratensis). synonym: chankae Obenberger 1935 c: 169 (subspecies of betuleti); Jendek 2006: 393 (synonym of pratensis djukini); Bellamy 2008 d: 2245 (subspecies of pratensis djukini); Jendek & Grebennikov 2011: 159 (synonym of pratensis). synonym: fennicus Obenberger 1936 a: 1029 (unnecessary replacement name for praeclarus); Jendek 2006: 393 (synonym of pratensis praeclarus); Bellamy 2008 d: 2245 (subspecies of pratensis); Jendek & Grebennikov 2011: 159 (synonym of pratensis). [subgenus Robertius; betuleti species group]. Biology/Bionomy: Bílý 2003: 23. Distribution within Iran. Golestan (Jendek & Grebennikov 2011), Mazandaran (Barimani et al. 2009, 2010), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Albania, Austria, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, China, Croatia, Czech Republic, Estonia, Finland, France, Germany, Georgia, Great Britain, Greece, Hungary, Italy, Kazakhstan, Latvia, Lichtenstein, Lithuania, Luxembourg Macedonia, Moldavia, Mongolia, the Netherlands, Norway, Poland, Romania, Russia, Serbia, Slovakia, Spain, Sweden, Switzerland, Turkey, Ukraine (Jendek 2006; Bellamy 2008 d; Jendek & Grebennikov 2011). Agrilus purpuratus (Klug) 1829: No. 12 (Buprestis); Bílý 1983 a: 84; Jendek 2006: 391; Bellamy 2008 d: 2255. [subgenus Duttus]. Immatures: Mateu 1974: 244. Biology/Bionomy: Mateu 1974: 273. Host plant(s) within Iran. Acacia flava (Fabaceae) (larva) (Bílý 1983 a). Distribution within Iran. Hormozgan (Bílý 1983 a), Iran (no locality cited) (Jendek 2006; Bellamy 2008 d). Distribution outside Iran. Algeria, Chad, Democratic Republic of Congo, Egypt, Israel, Jordan, Kenya, Lybia, Morocco, Saudi Arabia, Syria, Tanzania, Tunisia, Yemen (Jendek 2006; Bellamy 2008 d). Agrilus roscidus Kiesenwetter 1857: 127; Bílý 1983 a: 83 (as roscidus sieversi); Modarres Awal 1997: 132; Borumand 2002: 50 (as roscidus sieversi); Jendek 2006: 394; Bellamy 2008 d: 2268; Havaskary et al. 2012: 1; Ghahari & Bellamy 2012: 41. synonym: ascendens Abeille de Perrin 1895 b: cxxv. synonym: alticola Abeille de Perrin 1897: 8 (variety of roscidus). synonym: willcocksi Théry 1914: 129. synonym: graecus Obenberger 1916: 275. synonym: affectans Obenberger 1923 b: 86. synonym: subroscidus Obenberger 1924 c: 54. synonym: satunini Obenberger 1936 b: 110 (variety of roscidus). synonym: schevyrevi Stark 1955: 786. [subgenus Rosagrilus, roscidus species-group]. Immatures: Xambeu 1893 b: 93; Alexeev 1960: 1509, 1961: 7, 1981 b: 81; Bílý 1999: 35. Biology/Bionomy: Bílý 2003: 25. Host plant(s) within Iran. (Adult) Pistac
Chrysobothris Eschscholtz 1829
Genus CHRYSOBOTHRIS Eschscholtz, 1829 synonym: Amblis Gistel 1834. synonym: Odontomus Kirby 1837. synonym: Aglaura Gistel 1848. synonym: Tamina Gistel 1848. synonym: Euocys Gistel 1856. synonym: Knowltonia Fisher 1935. Chrysobothris affinis (Fabricius) 1794: 450, 458 (Buprestis); Richter 1952: 212; Radjabi 1968: 71 (?variety of affinis and var. heterochroa Obenberger 1916, unavailable); Modarres Awal 1997: 135 (with var. heterochroa Obenberger); Borumand 2002: 47; Kubáň 2006: 384; Bellamy 2008 c: 1590; Barimani et al. 2008: 201, 2009: 66; Rajabpour et al. 2012 (author: Ménétriés). synonym: congener (Paykull) 1799: 222 (Buprestis). synonym: tristis Abeille de Perrin 1900: 4 (preoccupied by tristis Deyrolle 1864). synonym: adusta Abeille de Perrin 1901: 8. synonym: cavinithorax Obenberger 1917 d: 52 (variety of affinis; unused original spelling). synonym: carinithorax Obenberger 1917 d: 52 (variety of affinis). synonym: materculae Holynski 1975: 357. [subgenus Chrysobothris]. Immatures: Schiødte 1870: 357, 372; Perris 1875: 381, 1877: 123; Richter 1952: 214; Palm 1962: 75; Bílý 1975 a: 419, 1999 a: 29. Biology/Bionomy: Bílý 2003: 50. Host plant(s) within Iran. Apple (Malus sp.), apricot (Prunus sp.), pear (Pyrus, Rosaceae) white mulberry (Morus sp., Moraceae) (Modarres Awal 1997); Albizia lebbec k, Prosopis cineraria (Fabaceae), Callistemon citrinus (Myrtaceae) (Rajabpour et al. 2012). Distribution within Iran. Chaharmahal & Bakhtiary, Kerman (Borumand 2002), East Azarbaijan (Modarres Awal 1997; Borumand 2002), Guilan, Hamadan, Isfahan, Lorestan, Sistan & Baluchestan, Yazd, Zanjan (Modarres Awal 1997), Khuzestan (Rajabpour et al. 2012), Mazandaran (Barimani et al. 2008, 2009), Qazvin (Radjabi 1968; Borumand 2002), Tehran (Radjabi 1968; Modarres Awal 1997; Borumand 2002), Iran (no locality cited) (Richter 1952; Kubáň 2006; Bellamy 2008 c). Distribution outside Iran. Albania, Algeria, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bosnia- Herzegovina, Bulgaria, Croatia,? Cyprus, Czech Republic, Denmark, Egypt, France, Georgia, Germany, Greece, Hungary, Italy, Kazakhstan, Lichtenstein, Luxembourg Macedonia, Moldavia, Montenegro, Morocco,? the Netherlands, Norway, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Syria, Tajikistan, Tunisia, Turkey, Turkmenistan, Ukraine, Uzbekistan (Kubáň 2006; Bellamy 2008 c). Note. Two parasitoid wasps, Glyptomorpha pectoralis (Brullé, 1832) (Hymenoptera: Braconidae: Braconinae) (2 ♀, 2 ♂) and Varzobia tibialis Nikol'skaya, 1960 (Hymenoptera: Chalcididae) (3 ♀) were collected as the parasitoid of Chrysobothris affinis in Lahijan (Guilan province) and Golestan National Park (Golestan province), respectively. Chrysobothris affinis nevskyi Richter 1944: 72; Farahbakhsh 1960: 74; Modarres Awal 1997: 135; Kubáň 2006: 384; Bellamy 2008 c: 1593. Host plant(s) within Iran. Apricot (Prunus sp., Rosaceae), white mulberry (Morus sp., Moraceae) (Farahbakhsh 1960; Modarres Awal 1997). Distribution within Iran. Northern provinces (Farahbakhsh 1960; Modarres Awal 1997), Iran (no locality cited) (Kubáň 2006; Bellamy 2008 c). Distribution outside Iran. Kazakhstan, Tajikistan, Turkmenistan, Uzbekistan (Kubáň 2006; Bellamy 2008 c). Chrysobothris affinis tetragramma (Ménétriés) 1832: 149 (Buprestis); Théry 1925: 81 (variety of affinis); Richter 1952: 214; Davatchi et al. 1959: 245; Farahbakhsh 1960: 74; Radjabi 1968: 71; Adeli 1972: 11; Modarres Awal 1997: 135; Borumand 2002: 47; Kubáň 2006: 384; Bellamy 2008 c: 1593; Barimani et al. 2010: 131. synonym: reitteri Kerremans 1900: 304 (nomen novum for inaequalis Reitter 1895); Kubáň 2006: 44 (new assignment), 384. synonym: inaequalis Reitter 1895 b: 128 (preoccupied by inaequalis Waterhouse 1889); Kubáň 2006: 44 (new assignment), 384. Host plant(s) within Iran. Ulmus sp., Salix sp., Juglans sp., fruit trees (Adeli 1972), apple (Malus sp., Rosaceae), elm (Ulmus sp., Ulmaceae), walnut (Juglans sp., Juglandaceae), willow (Salix sp., Salicaceae) (Davatchi et al. 1959; Farahbakhsh 1960; Modarres Awal 1997). Distribution within Iran. Golestan (Richter 1952), Hamadan (Théry 1925), Isfahan (Richter 1952, as tetragramma ab. bienerti), Mazandaran (Barimani et al. 2010), Sistan & Baluchestan (Radjabi 1968; Modarres Awal 1997; Borumand 2002), northern provinces (Adeli 1972; Farahbakhsh 1960; Modarres Awal 1997), Iran (no locality cited) (Davatchi et al. 1959; Kubáň 2006; Bellamy 2008 c). Distribution outside Iran. Armenia, Azerbaijan, Georgia, Turkey, Turkmenistan (Kubáň 2006; Bellamy 2008 c). Chrysobothris bicolorata Bílý 2000: 20; Kubáň 2006: 383; Bellamy 2008 c: 1601. [subgenus Abothris]. Distribution within Iran. Khuzestan (Bílý 2000), Iran (no locality cited) (Kubáň 2006; Bellamy 2008 c). Distribution outside Iran. None recorded. Chrysobothris catei Barries 2009: 55. [subgenus Chrysobothris]. Distribution within Iran. Kuhgiloyeh & Boyerahmad (Barries 2009). Distribution outside Iran. None recorded. Chrysobothris hexastigma Mannerheim 1837: 78; Semenov - Tian-Shankij & Richter 1934: 93 (synonym of affinis); Kubáň 2006: 383; Volkovitsh 2006: 58 (ressurected from synonymy); Bellamy 2008 c: 1637. synonym: deserticola Semenov-Tian-Shankij & Richter 1934: 94; Radjabi 1974: 38; Abai & Adeli 1984: 71; Modarres Awal 1997: 135; Borumand 2002: 48; Volkovitsh 2006: 57 (synonym of hexastigma). [subgenus Chrysobothris]. Biology/Bionomy: Kaplin 1981: 48, 52 (as deserticola) (with bibliography). Host plant(s) within Iran. Platanus orientalis L. (Platanaceae), trunk (Abai & Adeli 1984, as deserticola). Distribution within Iran. Khorasan (Radjabi 1974; Borumand 2002, as deserticola), generally distributed (Abai & Adeli 1984, as desertcola). Distribution outside Iran. Kazakhstan, Tajikistan, Turkmenistan, Uzbekistan (Kubáň 2006; Bellamy 2008 c). Note. Records for Platanus as a host plant and general distribution within Iran in Abai & Adeli (1984) are most probably based on misidentification; according to Kaplin (1981) and our observations (MGV) this species is associated with Ammodendron spp. (Fabaceae) in sandy deserts. * Chrysobothris jakovlevi Semenov-Tian-Shankij 1891: 340; Abai & Adeli 1984: 67; Modarres Awal 1997: 135; Kubáň 2006: 384; Bellamy 2008 c: 1643. unused original spelling: jakowlewi Semenov-Tian-Shankij 1891: 340. [subgenus Abothris]. Biology/Bionomy: Kaplin 1981: 48, 52 (with bibliography). Host plant(s) within Iran. Pistacia spp. (Anacardiaceae), branches (Abai & Adeli 1984), wild pistachio (Pistacia sp.) (Modarres Awal 1997). Distribution within Iran. Khorasan (Abai & Adeli 1984; Modarres Awal 1997). Distribution outside Iran. Turkmenistan (Kubáň 2006; Bellamy 2008 c). Note. Probable misidentification; according to Kaplin (1981) and our observations (MGV) this species associates with Ammodendron spp. (Fabaceae) in sandy deserts. Chrysobothris nana Fairmaire 1892 b: clii; Kubáň 2006: 384; Bellamy 2008 c: 1654. synonym: turkestanica Obenberger 1924 c: 29. synonym: partha Obenberger 1924 c: 30. [subgenus Abothris]. Distribution within Iran. Iran (no locality cited) (Kubáň 2006; Bellamy 2008 c). Distribution outside Iran. Afghanistan, Kazakhstan, Tajikistan, Turkmenistan, Uzbekistan (Kubáň 2006; Bellamy 2008 c). Chrysobothris parvipunctata Obenberger 1914 d: 133 (variety of solieri); Richter 1952: 220; Bílý 1983 a: 83; Borumand 2002: 48; Kubáň 2006: 385; Bellamy 2008 c: 1661; Bílý et al. 2011: 211; Abaei 2014: 691 (Chrisobothris). synonym: zarudniana Semenov-Tian-Shankij & Richter 1934: 94. synonym: beesoni Obenberger 1928 c: 308; Bílý 1983 a: 83; Borumand 2002: 48. synonym: kheirii Cobos 1970: 191 (subspecies of beesoni). [subgenus Chrysobothris]. Immatures: Cobos 1970: 189 (as beesoni). Host plant(s) within Iran. In Punica sp. (Lythraceae) wood (Borumand 2002), Pistacia sp. (Anacardiaceae) (Bílý et al. 2011), Platanus sp. (Farabi et al. 2012). Distribution within Iran. Hormozgan (Bílý 1983 a, as beesoni; Bílý et al. 2011), Kerman (Bílý et al. 2011), Kuhgiloyeh & Boyerahmad (Bílý 1983 a, as beesoni; Bílý et al. 2011), Qom (Bílý et al. 2011), Sistan & Baluchestan (Semenov-Tian-Shankij & Richter 1934, type of zarudniana; Richter 1952, as zarudniana; Borumand 2002, as beesoni; Bílý et al. 2011), Tehran (Bílý et al. 2011; Farabi et al. 2012), Yazd (Borumand 2002), Zagros area (Abaei 2014), Iran (no locality cited) (Kubáň 2006; Bellamy 2008 c). Distribution outside Iran. Afghanistan, India, Iraq, Pakistan, Turkmenistan, UAE (Bílý et al. 2011). Chrysobothris solieri Laporte & Gory 1837: 10; Abaei 2006 a: 123; Abaei 2006 b: 235; Kubáň 2006: 386; Bellamy 2008 c: 1681. synonym: pini Klingelhöffer 1845: 349. synonym: multipunctata Campanyo 1863: 641. synonym: quadridens Rey 1890: 172. [subgenus Chrysobothris]. Immatures: Perris 1854: 117; Bílý 1975 a: 423, 1999 a: 29. Biology/Bionomy: Bílý 2003: 52. Host plant(s) within Iran. Pinus eldarica (Pinaceae) (Abaei 2006 a, b). Distribution within Iran. Tehran (Abaei 2006 a, b), Iran (no locality cited) (Kubáň 2006; Bellamy 2008 c). Distribution outside Iran. Algeria,? Armenia,? Austria,? Azerbaijan, Bosnia-Herzegovina, Bulgaria, Croatia,? Czech Republic, Egypt, France, Germany, Greece,? Hungary, Iraq, Israel, Italy (Pantelleria), Lebanon, Libya, Macedonia, Malta, Montenegro, Morocco,? the Netherlands,? Poland, Portugal,? Romania,? Slovakia, Slovenia, Spain, Switzerland, Syria, Tunisia, Turkey,? Ukraine (Kubáň 2006; Bellamy 2008 c). Note. Two parasitoids including, Liotryphon sp. (Hymenoptera: Ichneumonidae) and Hockeria sp. (Hymenoptera: Chalcididae) were collected as the larval parasitoids by Abaei (2006 a, b).Published as part of Ghahari, Hassan, Volkovitsh, Mark G. & Bellamy, Charles L., 2015, An annotated catalogue of the Buprestidae of Iran (Coleoptera: Buprestoidea), pp. 1-141 in Zootaxa 3984 (1) on pages 97-99, DOI: 10.11646/zootaxa.3984.1.1, http://zenodo.org/record/24515
Chlaenius Bonelli 1810
Genus Chlaenius Bonelli, 1810 Subgenus Dinodes Bonelli, 1810 Chlaenius (Dinodes) cruralis Fischer von Waldheim, 1829 = persicus LaFerte-Senectere, 1851 Studied material: (ZMTU): Kordak (Prov. of Qazvin). Citations: Astrabad (Prov. of Gilan; Mandl. 1983); Moghan (Prov. of Ardabil; Mandl, 1983);), Astrabad and Kopt Dagh (Prov. of Golestan; Mandl, 1983); Basmenj Village (Prov. of East Azarbaijan; Atamehr, 2013); Gorgan env. (Prov. of Golestan; Afshari & Khormali, 2014). Chlaenius (Dinodes) decipiens (Dufour, 1820) Citations: Without exact locality (Hieke & Wrase, 1988). Chlaenius (Dinodes) huedepohli Mandl, 1983 Citations: Ziba-Kessar, Rasht (Prov. of Gilan; Mandl, 1983); 5 km West of Samsuni 5 - 6.6.2008 3500 m (Prov. of Chahar Mahal va Bakhtiari; Anichtchenko, 2008). Chlaenius (Dinodes) viridis (Ménétriés, 1832) Citations: Without exact locality (Mandl, 1983). Subgenus Chlaeniellus Reitter, 1908 Chlaenius (Chlaeniellus) extensus Mannerheim, 1825 Citations: Without exact locality (Kirschenhofer, 1985); Shiraz (Prov. of Fars; Mohammadzadehfard, 2005). Chlaenius (Chlaeniellus) flavipes Ménétriés, 1832 Studied material: (ZMTU): Damavand, 2008 (Prov. of Tehran), Dohezar River, 2008 (Prov. of Mazandarn) and Khane Zenian, 2010 (Prov. of Fars). Citations: Pirbakran (Prov. of Esfahan; Mandl, 1963). Chlaenius (Chlaeniellus) intermedius Chaudoir, 1856 = fulvipes Chaudoir, 1842 Citations: Without exact locality (Kirschenhofer, 2003); Basmenj Village (Prov. of East Azarbaijan; Atamehr, 2013). Chlaenius (Chlaeniellus) koenigi Semenov, 1888 Citations: Without exact locality (Kirschenhofer, 2003). Chlaenius (Chlaeniellus) laetiusculus Chaudoir, 1856 = rapylii Morvan 1975 Citations: IranShahr (Prov. of Sistan va Balochestan; Morvan, 1975). Chlaenius (Chlaeniellus) latithorax Mannerheim, 1844 Citations: Province Mazandaran (Chaudoir, 1844). Chlaenius (Chlaeniellus) nigricornis Fabricius, 1787 Citations: Farsistan region (prov. of Fars; Redtenfoacher, 1848). Chlaenius (Chlaeniellus) nitidulus Schrank, 1781 Studied material: (ZMTU): Astaneh, Safid Rud (Prov. of Gilan). Citations: Province Mazandaran (Chaudoir, 1842). Chlaenius (Chlaeniellus) syriacus Chaudoir, 1876 Citations: Without exact locality (Kirschenhofer, 2003). Chlaenius (Chlaeniellus) tenuilimbatus Ballion, 1871 Citations: Lurestan (Prov. of Lolrestan; Mandl, 1983); Elburz Mts. R (Mandl, 1983). Chlaenius (Chlaeniellus) tibialis Dejean, 1826 Studied material: (ZMTU): Astane, 2008 (Prov. of Gilan). Citations: Without exact locality (Kirschenhofer, 2003). Chlaenius (Chlaeniellus) tristis tristis (Schaller, 1783) Citations: Keretsch, West of Tehran (Prov. of Tehran; Mandl, 1967 b); Nowshaht (Prov. of Azerbaijan; Gueorguiev, 2011). Chlaenius (Chlaeniellus) vestitus (Paykull, 1790) Studied material: Khirood forest and Siah bishe (prov,. Mazandaran). Citations: Distributed all over country. Farsistan region (prov. of Fars; Redtenfoacher, 1848); Qualeh-e Asghard (Prov. of Kerman; Mandl, 1963); Pirbakran (Prov. of Esfahan; Mandl, 1963); SultanAbad, Arak (Prov. of Markazi; Mandl, 1991); Province Gilan (Salari et al., 2012); Shiran Village (Prov. of Ardabil; Atamehr, 2013); Gorgan env. (Prov. of Golestan; Afshari & Khormali, 2014); Siah-Bisheh (prov. of Mazandaran; Azadbakhsh et al., 2015). Subgenus Agostenus Fischer von Waldheim, 1829 Chlaenius (Agostenus) alutaceus Gebler, 1829 Citations: Province Mazandaran (Chaudoir, 1844); Shazawar (Prov. of Mazandaran; Mandl, 1983); Dagestan (Prov. of Golestan; Mandl, 1983); Talysh (Prov. of Gilan; Mandl, 1983); Province Giyla (Salari et al., 2012). Subgenus Epomis Bonelli, 1810 Chlaenius (Epomis) circumscriptus Duftschmid, 1812 Citations: Province Mazandaran (Chaudoir, 1842). Subgenus Pseodochlaeniellus Jeannel, 1949 Chlaenius (Pseodochlaeniellus) iranensis Kirschenhofer, 1998 Citations: Minab (Prov. of Hormozgan; Kirschenhofer, 1998). Chlaenius (Pseodochlaeniellus) puncticollis Dejean, 1826 = persicus Jedllicka, 1968 jedlickai Mandl, 1991 Studied material: Geno, 2015 (Prov. of Hormozgan). Citations: Minab (Prov. of Hormozgan; Jedlička, 1968). Subgenus Chlaeniostenodes Basilewsky, 1953 Chlaenius (Chlaeniostenodes) canariensis persicus Redtenfoacher, 1850 = persicus Morvan, 1975 Citations: Farsistan region (prov. of Fars; Redtenfoacher, 1848); Isin (Prov. of Hormozgan; Morvan, 1974). Note: Ghahari et al. (2010) cited the species from Amol (Prov. of Mazandaran), Also Ghahari et al. (2009 b) cited the species from Babol (Prov. of Mazandaran). This subspecies is known only from South of Iran so that this citation very probably belongs to C. flavipes, and it has been another misdiagnosis of this author. Chlaenius (Chlaeniostenodes) canariensis seminitidus Chaudoir, 1856 Studied material: Bandar Abbas, 2014 (Prov. of Hormozgan). Citations: Without exact locality (Kirschenhofer, 2003). Subgenus Chlaenites Motschulsky, 1860 Chlaenius (Chlaenites) spoliatus spoliatus Rossi, 1792 Studied material: (ZMTU): Khane Zenian, 2010 (Prov. of Tehran) and Shahre Kord, 2012 (Prov. of Chahar Mahal va Bakhtiari). Citations: Farsistan region (prov. of Fars; Redtenfoacher, 1848); Babol (Prov. of Mazandaran; Mandl, 1963); Province Gilan (Salari et al., 2012). Subgenus Amblygenius LaFerte-Senectere, 1851 Chlaenius (Amblygenius) dimidiatus Chaudoir, 1842 Studied material: Khirood forest, 2012 (Prov. of Mazandaran). Citations: Province Gilan (Salari et al., 2012); Aras Baran (Prov. of East Azarbaijan; Ghahari et al., 2010); Khirood forest, NowShahr (Prov. of Mazandaran; Azadbakhsh et al., 2015). Subgenus Chlaenius Bonelli, 1810 Chlaenius (Chlaenius) festivus festivus Panzer, 1796 Studied material: Lale Zar. 2014. (prov,. of Kerman) and Khirood Forest, 2012 (Prov. of Mazandaran); (ZMTU): Shahre Kord, 2012 (Prov. of Chahar Mahal va Bakhtiari). Citations: Farsistan region (prov. of Fars; Redtenfoacher, 1848); Sabzawaran and Qualeh-e Asghard (Prov. of Kerman; Mandl, 1963); Pirbakran (Prov. of Esfahan; Mandl, 1963); Zanjan (Prov. of Zanjan; Mandl, 1963); Province Gilan (Salari et al., 2012); Basmenj Village (Prov. of East Azarbaijan; Atamehr, 2013). Subgenus Stenochlaenius Reitter, 1908 Chlaenius (Stenochlaenius) coeruleus (Steven, 1809) Citations: Farsistan region (prov. of Fars; Redtenfoacher, 1848). Chlaenius (Stenochlaenius) dostojevskji Tschitschérine, 1895 Studied material: Koh paye, 2014 (Prov. of Kerman); Haji Abad, 2014- 2015 (Prov. of Hormozgan). Citation: Bandar Abbas (Prov. of Hormozgan; Mandl, 1972 a); Without exact locality (Kirschenhofer, 2003). Chlaenius (Stenochlaenius) lederi Reitter, 1888 = picicornis (Mandl, 1972) Citations: SultanAbad, Arak (Prov. of Markazi; Mandl, 1972 a). Subgenus Trichochlaenius Seidlitz, 1887 Chlaenius (Trichochlaenius) aeneocephalus aeneocephalus Dejean, 1826 Studied material: Khirood forest, 2012 (Prov. of Mazandaran); (ZMTU): Karaj, University of Agriculture (Prov. of Elburz), Kolijak and 40 km S Tavir (Prov. of Mazandaran). Citations: Gorgan env. (Prov. of Golestan; Afshari & Khormali, 2014); Khirood forest, NowShahr (Prov. of Mazandaran; Azadbakhsh et al., 2015). Chlaenius (Trichochlaenius) gotschii Chaudoir, 1846 = angusticollis Mannerheim, 1844 Citations: Astrabad (Prov. of Mazandaran; Chaudoir, 1844). Chlaenius (Trichochlaenius) steveni Quensel, 1806 Citations: Samin & Sakenin (2014) cited the species from Savadkooh (Prov. of Mazandaran), but this record is dubious as in the last Palearctic catalogue. Subgenus Lithochlaenius Kryzhanovskij, 1976 Chlaenius (Lithochlaenius) asara Kirschenhofer, 2014 Citations: Elburz Mts. R, btw Karaj and Chalus, Asara env., 1900 - 2050 m (Prov. of Elburz; Kirschenhofer, 2014).Published as part of Azadbakhsh, Saeed & Nozari, Jamasb, 2015, Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae), pp. 1-108 in Zootaxa 4024 (1) on pages 66-69, DOI: 10.11646/zootaxa.4024.1.1, http://zenodo.org/record/24049
Acrosternum heegeri Fieber 1861
Acrosternum heegeri Fieber, 1861 (Figs. 1–3) Acrosternum heegeri Fieber, 1861: 331. SynTypes: südliches Europa [= souThern Europe]; Muséum NaTional d’HisToire NaTurelle, Paris (Orian 1965). The nomenclaTural hisTory and synonyms of The species were reViewed by Rider (2006). Proposed Hungarian name. Rőtcsápú vándorpoloska Material examined. Hungary, Budapest, Békásmegyer, Királyok útja, 47°35'51.9"N 19°3'59.4"E, 2nd floor balcony, at light, 24.vii.2016, leg. B. Károlyi (six males and eight females deposited in the HNHM). New record to Hungary. Diagnosis. Body rather uniformly green, with pronotum, costal margin of fore wing, and connexivum narrowly pale, without contrasting markings on scutellum; membrane uniformly whitish; integument smooth; head about 1.2 times as broad as its length along midline, clypeus anteriorly free; antenna contrastingly bicolorous, scape and basipedicellite green, distipedicellite and basiflagellum red except of their bases, distiflagellum red; apex of labium generally surpassing middle of abdominal ventrite III (Figs. 1–2). Genitalia of the male were accurately illustrated by Linnavuori (1972), Fuente (1975), Josifov & Heiss (1989), and Ribes & Pagola-Carte (2013), and less accurately by a number of other authors. Intraspecific variability. Overwintering individuals are reddish brown. Individuals with the head and anterior portion of pronotum being pale yellow occur sporadically in all populations in low frequency. Bionomics, economic importance. The species is apparently polyphagous, feeding on both gymnosperms and angiosperms. Published data on its host plants were summarized by Ribes & Pagola-Carte (2013) and Ghahari et al. (2014). Mehrnejad (2010) claimed that it is a significant pest of the pistachio, Pistacia vera L. (Anacardiaceae) in Iran; besides its direct damage, it is also a vector of Eremothecium coryli (Peglion) Kurtzman (Saccharomycetaceae), a plant pathogen fungus causing stigmatomycosis (Ershad & Barkhordary 1974). Distribution. A Holomediterranean-Turanian species (Carapezza 1997, Ribes & Pagola-Carte 2013). Its distribution was summarized by Rider (2006), Aukema et al. (2013b) and Ribes & Pagola-Carte (2013); a rather incomplete distribution map was provided by Tamanini (1981: 135, fig. 45) which is updated now (Fig. 3). In the Mediterraneum the distribution of the species strictly follows the vegetational zone of Mediterranean forests, woodlands and scrub; in the Middle East, however, it extends far into the zone of temperate deciduous and mixed forests eastwards. The species has been recorded from Iraq by Al-Ali (1968, 1977) and a few subsequent authors; all of these records are doubtful, some of them, e.g. the one by Augul et al. (2015: 125, fig. 2a) are evidently based on misidentification and pertain to other species, most probably A. breviceps (Jakovlev, 1889). Records from “Arabien” [= Arabia, referring either to Saudi-Arabia or the Arabian Peninsula] and Socotra (Distant 1903, Stichel 1961) are erroneous and at least partly pertain to A. arabicum Wagner, 1959 (Linnavuori & Al-Safadi 1993); the voucher specimen referred to by Distant could not be found in the Natural History Museum, London, and the species was not collected during a recent thorough faunistic exploration of Socotra (A. Carapezza & P. Kment, pers. comm.). The species has been recorded from Madagascar (Signoret 1861, as Rhaphygaster incerta), but no subsequent author confirmed this record (Cachan 1952), therefore its occurrence in the island is considered as doubtful; Signoret’s record is probably based on mislabelled or misidentified specimens and it potentially pertains to Chinavia bergrothi (Horváth, 1904). Old records from Belgian Congo, now Democratic Republic of the Congo (Schouteden 1909), British and Dutch East Africa, now Kenya ad Tanzania (Jeannel 1913) were considered as questionable by Linnavuori (1972). The figure of the genital capsule drawn after a male from East Africa (Kilimanjaro, Tanzania) (Cachan 1952: 442, fig. 525) is apparently different from the condition found in A. heegeri (posterior margin distinctly concave at midline, not nearly transversely truncate as in A. heegeri) and therefore probably shows another species. Therefore we consider all Afrotropical records, including those listed above as well as those provided by Distant (1901), Schouteden (1905, 1929), Villiers (1950, 1952, 1954, 1956) and Robertson (2009) as in need of confirmation. The records from the Seychelles (Distant 1913, Orian 1965, Roche 1977, Gerlach 2003, Gerlach et al. 2006) are controversial and data published by different authors are conflicting, therefore we consider the presence of the species in the Seychelles and the Mascarene Archipelago as in need of verification. The available information on the distribution of the species is summarized below. For countries for which comprehensive faunistic studies summarizing earlier literature are available, e.g. Italy (Servadei 1967), the former Yugoslav countries (Protić 2001), Iran (Ghahari et al. 2014), Spain and France (Ribes & Pagola-Carte 2013), earlier references are not cited. EUROPE. Spain: numerous localities in Navarra, Aragón, Catalunya, Valencia, Madrid, Andalucia, Baleares: Eivissa Is., Mallorca Is., Menorca Is., Cabrera Is. (Ribes & Pagola-Carte 2013). Portugal: Beja (Oliveira 1895), Alfeite (Seabra 1926), Várzea da Orada, Serra de Monchique (Pissarro 1951). France: numerous localities in the Departments of Pyrénées-Orientales, Aude, Hérault, Gard, Ardèche, Vaucluse, Bouches-du-Rhône, Alpes- Maritimes, Alpes de Haute Provence, and Corse (Ribes & Pagola-Carte 2013). Monaco (Ponel et al. 2013). Italy: Liguria, Emilia Romagna, Toscana, Umbria, Lazio, Puglia, Basilicata, Calabria, Sicilia, Sardegna, Giglio Is., Tremiti Is., Pantelleria Is. (Servadei 1967, Tamanini 1981, Carapezza 1995). Slovenia: Dragonja, Pišine (Gogala 2008). Croatia: Rijeka, Sveta Jelena, Vrana, Kaštel Stari, Sućurac, Split, Janjina, Dubrovnik, Korčula Is., Trstenik Is., Lopud Is. (Protić 2001), Lošinj Is. (Furlan & Gogala 1995). Bosnia and Herzegovina: Domanovići (Protić 2001). Montenegro: Stari Bar, Kotor (Protić 2001). Albania: Lushnjë (Misja 1973). Hungary: Budapest (new record). Romania: Moldova Nouă (Kis 1984). Bulgaria: Sandanski (Göllner-Scheiding & Arnold 1988), Kresna Gorge (Simov 2001). Macedonia: Udovo, Stari Dojran (Protić 2001). Greece: Kalambaka (Günther 1990), Argolis (Ribes & Pagola-Carte 2013), Crete: several localities (Heiss 1983, Heckmann et al. 2015). Ukraine: Yalta, Alushta (Jakovlev 1906); earlier records from Kharkiv are erroneous (Putshkov & Putshkov 1996). Russia: Caucasus (Kerzhner & Jaczewski 1964). Turkey (European part): Karaağaç (Fent & Aktaç 2007). — ASIA. Turkey (Asian part): Adana, Antalya, Diyarbakır, Gaziantep, Hatay, İzmir, Manisa, Mersin, Muğla, Kayseri, Şanlıurfa Provinces (Önder et al. 2006), Manavgat, Posof, Artvin, Şenkaya, Erenköy, Kemaliye, Datça, Trabzon (Külekçi et al. 2009), Osmangazi (Gençer et al. 2004), Tapan, Ekbaz (Hoberlandt 1956), Atabey (Karsavuran et al. 2008), Bursa (Kaya & Kovanci 2004), Mardin (Özgen et al. 2005), Aksu, Kepez, Serik, Karacabey, İspir, Kahramanmaraş (Yazici et al. 2014), Bodrum (Tezcan et al. 2013), Kayseri (Ribes & Pagola-Carte 2013). Cyprus: Akrotiri (Lindberg 1948). Syria (Frey-Gessner 1881). Israel: env. of Jericho (Hoberlandt 1952), Rehovot (Linnavuori 1960). Jordan: several localities around Amman (Katbeh et al. 2000). Georgia: Sukhumi (Kiritshenko 1939). Azerbaijan: Aresch [= Agdash] (Sienkiewicz 1964), Absheron, Agdzhabedin [= Aghjabadi] Shamkhor [= Shamkir] Distr., Kirovabad [= Ganja] (Gidayatov 1982). Iraq ? (Al-Ali 1968, 1977). Iran: Ardabil, Bushehr, Fars, Hormozgan, Khuzestan, Lorestan, East Azarbaijan, Golestan, Guilan, Mazandaran, Kerman, Khorasan, Sistan & Baluchestan, Tehran (Ghahari et al. 2014). Afghanistan: Sarobi [= Surobi] (Hoberlandt 1984). Pakistan: Rahim Yar Khan (Ahmad & Rana 1989). — NORTH AFRICA. Madeira (Ribes & Heiss 2008). Canary Islands: Tenerife (Aukema et al. 2006), La Palma (Aukema et al. 2013a). Morocco: Debdou, Oudja, Berkane, Ras Foughal, Fès [= Fez], Asni, Dar Drius (Vidal 1937, 1949), Marrakech [= Marrakesh] (Ribes & Pagola-Carte 2013). Algeria: Biskra (Lethierry 1889, Deghiche-Diab et al. 2015), Tahount Arak, Mouydir, Imarera, Amguid (Bergevin 1934), Oued R[h]iou, Chellala, Teniet el Haad [= Theniet el Had], Mt. Ouarsenis, Tiaret (Sienkiewicz 1964), Oued Marsa (Wagner 1965), Boufarik (Ribes & Pagola-Carte 2013). Tunisia: Hadjeb el Aioun [= Hajeb El Ayoun], Porto Farina [= Ghar al Milh], Kairouan (Sienkiewicz 1964), Korbous, Téboursouk, Tabarka, Barrage Mellègue, Ousseltia, Thala, Gafsa (Carapezza 1997). Libya: Tripoli, Mellaha, El Giofra (Mancini 1937), Serti [= Sirte], Jebel Akhdar Messa, El Merj [= Marj], Tobruck [= Tobruk] (Eckerlein & Wagner 1970). Egypt: Cairo, Meadi, Siwa (Linnavuori 1964), Zaranik Protectorate (El-Moursy et al. 2001). — AFROTROPICAL REGION. Senegal ? (Villiers 1956). Mali ? (Robertson 2009). Niger ? (Villiers 1950, 1954). Benin ? (Villiers 1952). Nigeria ? (Robertson 2009). Sudan ? (Villiers 1956). Ethiopia ? (Schouteden 1905, 1929). Eritrea ? (Robertson 2009). Gabon ? (Distant 1901). Democratic Republic of the Congo ? (Schouteden 1909). Kenya ? (Jeannel 1913). Tanzania ? (Jeannel 1913). Madagascar ? (Signoret 1861). Rodrigues Is. ? (China 1926). Seychelles ?: Mahé Is.?, Silhouette Is.?, Aride Is.? (Roche 1977, Gerlach 2003, Madl 2006).Published as part of Károlyi, Balázs & Rédei, Dávid, 2017, Acrosternum heegeri Fieber, 1861 (Hemiptera: Heteroptera: Pentatomidae), another Mediterranean bug expanding to the north in Zootaxa 4347 (2), DOI: 10.11646/zootaxa.4347.2.13, http://zenodo.org/record/104567
Leptopus gakalae Khazaei & Polhemus & Tahami 2020, n. sp.
Leptopus gakalae n. sp. (Figs. 2–5) Type material examined. Holotype, male, IRAN, Kohgiluyeh and Boyer-Ahmad Prov., Gakal Cave, 30°18’44”N, 51°9’28”E, 1100 m elev., 18 October 2015, Zohreh Khazaei (BPBM). Paratypes: IRAN, Kohgiluyeh and Boyer- Ahmad Prov.: 4 males, 5 females (one apterous), same data as holotype (1 female BPBM, remainder ZM-CBSU). Description. Winged male: Body length (from head to apical tip of forewings) 2.5–3.5 mm; maximum width (across hemelytra) 1.0 mm (Fig. 1). General coloration pale whitish-tan, with thorax and broad maculae on hemelytra yellowish-brown, appendages pale (Fig. 1); entire dorsal surface bearing moderately long, erect, stout, pale setae. Structural characters: Head length 0.60, width across eyes 0.90, brown; frons declivent, nearly vertical, bearing three long, stout, gold setae basally at junction with vertex; width of vertex 0.50, with semi-spherical protuberance medially bearing 2 ocelli; a pair (1 + 1) of whitish setae set at a short distance from each other immediately behind ocelli; two whitish setae anteriorly on midline of vertex; suture present at midline of vertex next to mid-projection of vertex; clypeus yellowish; posteroventral margins dark; three long, whitish spines ventrally to either side of longitudinal midline on post-genal sclerites. Eyes red, prominent, semi-circular, bearing short, pale ocular setae, inner margins bi-concave. Rostrum attaining fore coxae, segment II yellowish, longest, with two pairs (2+2) of long spines located on tubercles (Fig. 2); segment III yellowish, with two pairs of long tuberculate spines; segment IV shortest, dark brown at apex. Antennae with segment I short, 0.4× as long as segment II, thickened in the middle and narrowed proximally and apically; segment II 1.25× length of segment III, yellowish, clavate, and thickened apically; segments III and IV dark brown, segment IV 0.28× as long as segment III, with recumbent dark brown setae; lengths of segments I–IV = 0.15, 0.40, 0.80, 0.45. Pronotum length 0.60, maximum width 0.75, yellowish-brown, punctate, with scattered spines; pronotum partitioned into anterior and posterior parts near middle by prominent transverse furrow; anterior lobe with longitudinal furrow along midline, separating calli, each callus with two small angular processes; lateral margins very narrowly explanate, with scattered erect, spine-like pale brown setae. Scutellum length 0.25, width 0.35, brown with a pale tip, bearing a pair (1+1) of erect, spine-like pale brown setae on either side of midline; central section depressed. Elytra coleopteriform, completely sclerotized and lacking membrane; clavus and corium well defined, punctate, set with scattered erect, spine-like, pale brown setae, ground color whitish-tan, with prominent X-shaped, yellowish-brown fascia traversing center of corium (Fig. 2); hind wings reduced, half as long as forewings; basal section of hemelytra inside of medial fracture raised; hemelytral margin narrowly explanate, embolium only very weakly expanded. Abdomen brown dorsally, with 7 visible segments including medio- and laterotergites. Venter with mesosternum and metasternum brown, tumescent; abdomen yellowish-brown, bearing sparse inconspicuous pale setae. Legs yellowish-white with a preapical dark spot on femora; foretibia with three spines dorsally originating from tubercles (Fig. 3), apical 3 rd spine shorter, plus three light brown spines ventrally, the middle one longer; fore femur yellowish-white with a preapical dark spot, four fine tuberculate spines dorsally, the proximal one shorter, five tuberculate spines ventrally, the proximal 2 nd one and the apical 5 th one shorter; middle and hind femora yellowishwhite with a preapical dark spot; tarsi yellowish-white, middle and hind tarsi brown apically, claws also brown; apices of fore tibia and fore tarsus brown, fore tarsus also bearing long pale setae. Lengths of leg segments as follows: fore femur–tibia–tarsomere 1– tarsomere 2–tarsomere 3 = 0.85/0.85/0.05/0.10/0.10; middle femur–tibia–tarsomere 1– tarsomere 2–tarsomere 3 = 0.75/1.00/0.05/0.10/0.15; hind femur–tibia– tarsomere 1– tarsomere 2– tarsomere 3 = 0.95/1.45/0.05/0.15/ 0.20. Male genitalia with parameres symmetrical, elongate, apices broadly rounded (Fig. 4). Winged female: Similar to winged male in general structure and coloration (Fig. 2); length 3.20, maximum width 1.20. Etymology. The specific name “gakalae” refers to the Gakal Cave, where the type series was collected. Remarks. Our new species clearly belongs to the clade containing Leptopus, Leptopoides J. Polhemus & D. Polhemus, 1991, and Patapius Horvath, 1912, due to the presence of stout spines on both of the first two visible rostral segments (J. Polhemus & D. Polhemus 1991, Linnavuori & van Harten 2002). It runs to the genus Leptopus in the key of J. Polhemus & D. Polhemus (1991), based on the presence of short, slender ocular setae, and the presence of two divergent rows of stout spines on the fore tibia (Fig. 3). Seven species of Leptopodidae have been previously recorded from Iran (Hoberlandt 1983, Schuh et al. 1987, Ghahari et al. 2013): Erianotus lanosus Dufour, 1834; Patapius sentus Drake & Hoberlandt, 1951; Patapius spinosus (Rossi, 1790); Valleriola assouanensis (Costa, 1875); Leptopus decus Drake, 1955; Leptopus hispanus Rambur, 1840; and Leptopus marmoratus (Goeze, 1778). The Iranian records for L. marmoratus provided by Sakenin et al. (2010) and Samin et al. (2011) have, however, been called into question (see subsequent discussion under that species). Most of the above species records also lack information on ecological context, although Patapius spinosus was noted as having been taken from the margins of a small stream near Ramhormoz in Khuzestan (Linnavuori 2009). In addition to the two widespread species L. hispanus Rambur (Fig. 7) and L. marmoratus (Fig. 10), which occur primarily to the west of Iran from southern Europe through the Balkans to trans-Caucasia, three additional species, L. travancorensis Distant, L. scitulus Drake (Fig. 8) and L. decus Drake, have been described from the western areas of India; the latter taxon was described from Punjab, and its range extends westward to the Sistan & Baluchestan Province of Iran. Given that no overall revision of Leptopodidae has been undertaken since that of Horváth (1911), which is now taxonomically incomplete, it has been necessary to accurately determine that our new species from Iranian caves is indeed discrete from those previously described. The second author examined specimens of all the taxa noted above during a visit to the collections of the Smithsonian Institution in Washington, DC, and was able to verify that L. gakalae is distinct from all of them (see key). Another species of Leptopus, L. markusiki Vinakurov, 2012, has been described from moderate elevation in the Himalayan foothills of Himanchal Pradesh state (Vinokurov 2012) but has not been examined by the authors. However based on the figures provided by Vinokurov (2012), this species is easily separated from L. gakalae by its blotched rather than fasciate color pattern on the hemelytra, and the distinctive, club-like structure of the male paramere. Habitat notes. Gakal Cave is located in the Zagros Mountains woodland, where Persian oak is the dominant tree (Fig. 1). In the dark zone of the cave, there is a puddle of water supplied with a spring and fed by water dripping down into the puddle from the cave’s ceiling (Fig. 6). The Zohreh River is the nearest surface water feature, lying about 10 km east of the cave. The specimens were collected in the entrance zone, where they could be seen clearly with the amount of sunlight coming in. The leptopodids were observed adjacent to a population of Collembola, which may provide a food source.Published as part of Khazaei, Zohreh, Polhemus, Dan A. & Tahami, Mohadeseh S., 2020, A new species of Leptopus (Heteroptera: Leptopodidae) from caves in Iran, with notes on other cavernicolous Iranian Heteroptera, pp. 246-258 in Zootaxa 4763 (2) on pages 248-251, DOI: 10.11646/zootaxa.4763.2.7, http://zenodo.org/record/375830
