238 research outputs found

    Cretaceomma RASNITSYN & MAALOUF & MAALOUF & AZAR 2022, gen. nov.

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    Genus Cretaceomma Rasnitsyn & Azar gen. nov. Type species. Cretaceomma libanensis sp. nov.; by present designation. Other species. Galloromma turolensis Ortega et al., 2011a, transferred herein to Cretaceomma gen. nov. Etymology. The genus name combines parts of Cretaceous and Galloromma. Diagnosis. Female antenna 14-segmented with 7- segmented funicle and 5-segmented loose (movably interconnected) clava. Pronotum reaching wing base or nearly so. Remarks. New genus differs from the type genus of the family Galloromma Schlüter, 1978 in more polymerous antenna (13-segmented with 4-segmented club in Galloromma) and pronotum apparently reaching wing bases (shortened posterolateral and far not reaching wing bases in Galloromma). Galloromma turolensis Ortega et al., 2011 from the Albian San Just amber in Spain is similar to the type species in above characters and that is why it is transferred to the new genus.Published as part of RASNITSYN, ALEXANDR P., MAALOUF, MOUNIR, MAALOUF, RAMY & AZAR, DANY, 2022, New Serphitidae and Gallorommatidae (Insecta: Hymenoptera: Microprocta) in the Early Cretaceous Lebanese amber, pp. 120-136 in Palaeoentomology 5 (2) on pages 128-129, DOI: 10.11646/palaeoentomology.5.2.4, http://zenodo.org/record/653041

    Aenigmaraphidophora AZAR & MAALOUF & NEL 2022, gen. nov.

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    Genus Aenigmaraphidophora gen. nov. urn:lsid:zoobank.org:act: 7EFAE62D-2D6E-4DB7- B2EC-F6C6579455A5 Type species. Aenigmaraphidophora mouniri sp. nov. Etymology. Named after the Latin ‘aenigma’ for its enigmatic family position, and the genus name Raphidophora, Gender feminine. Diagnosis. Very long maxillary palps, longer than fore legs; upper margin of first segment of hind tarsi without any spines; fore coxa with a lateral tubercle; mid femora with one inner and one outer relatively large movable apical spines; male subgenital plate with styles; longest apical spur of hind tibiae slightly shorter than first and second segments of hind tarsi combined; no process on dorsal surface of abdominal tergites; epiproct without process; hind 2 nd and 3 rd tarsomeres with a widened pulvillus.Published as part of AZAR, DANY, MAALOUF, RAMY & NEL, ANDRÉ, 2022, An enigmatic Tettigoniidea from the Lower Cretaceous amber of Bqaatouta, Lebanon (Orthoptera, Ensifera), pp. 233-239 in Palaeoentomology 5 (3) on page 234, DOI: 10.11646/palaeoentomology.5.3.5, http://zenodo.org/record/682083

    Leptoserphites iriae RASNITSYN & MAALOUF & MAALOUF & AZAR 2022, sp. nov.

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    Leptoserphites iriae Rasnitsyn & Azar, sp. nov. (Figs 6–8) Type material. Holotype, female, specimen no QBC- 16A (Fig. 6A–C), Maalouf collection, complete insect partially distorted postmortem (possibly diagenetically due to plastical deformation of enclosing amber under uneven pressure of deposits); paratypes, QBC-16B (Fig. 6D) and QBC-16C (Fig. 6E, F), Maalouf collection, synincluded with holotype in the same piece of amber and still more suffered from deformation (QBC-16B is particularly poor and included mainly being synincluded and having similar wing venation). All type material is from Qanat Bakish (Baskinta) amber outcrop, Mouhafazet Jabal Loubnan (Governorate Mount Lebanon), Caza (= District) El-Maten, Central Lebanon; lower Barremian. Type material is deposited in the Natural History Museum of the Lebanese University, Faculty of Sciences II, Fanar, Lebanon. Etymology. Specific name is after Iria Maalouf Alonso, daughter of one of us (RM). Diagnosis. Differs from the type species (L. pabloi sp. nov.) in having antenna 11-segmented with no clear traces of fusion of two basal flagellomeres into one, flagellomeres longer, propodeum transridged, pterostigma wider, 2r-rs shorter, no tubular structures present but R and pterostigma, hind wing with no r-m, petiolar segments longer. Locality and horizon. Baskinta, in the locality of Qanat Bakish (33°57′3″N, 35°47′24″E, elev. 1,488 m), Caza (= District) El-Maten, Mouhafazet (= Governorate) Mount Lebanon, in Central Lebanon; lower Barremian. Description. (Based on holotype otherwise stated) Female. Color dark fuscous, legs paler. Body with no surface sculpture and prominent vestiture discernible. Head elongate as preserved (Figs 7A, 8A, both on paratype QBC-16C) (possibly due to deformation), with eyes small, not much convex, placed before middle of head sides (more rostrally). Antenna 11-segmented [Figs 8A (paratype QBC-16C), 8D (holotype)], with scape long and wide, about as long as next three or four antennomeres combined, pedicel shorter than some apical segments, flagellum wide, somewhat flattened, slightly narrowing toward base, most segments longer than wide, basal flagellomere slightly longer than next one. Distinct subequal mandibles, right with three teeth and left with two ones. Of maxillary (Fig. 8B) and labial palps (Fig. 8C), only their elongate apical segments visible. Mesosoma with dorsal pronotal part apparently short, mesoscutellum in side view with hind part somewhat elevated,metascutum forming high step above wide trough of metapostnotum, propodeum short, gently arching toward metasomal base in side view, transridged. Forewing (Fig. 8E) with C, R behind pterostigma, RS before and behind 2r-rs (except for short section forming “ T ” pattern together with 2r-rs), M+Cu and M lost as pigmented veins or nearly so; R and pterostigma tubular; RS &M, 2r-rs and 1-Cu nebulous; pterostigma high (distinctly less than twice as long as wide), together with 2r-rs enclosed in lightly pigmented area; 2r-rs half as long as pterostigma wide; all membrane covered with short setae (margin included). Hind wing (Fig. 8F) with only R pigmented (tubular), R apex with three hamuli, hind margin with fringe of long setae. Legs ordinary except that fore tibial spur (calcar) straight except bent subapical with bifid apex (Fig. 8G), and fore basitarsus excised almost for all its length and with single row of setae along excision; tibial spurs 1: 1: 2; tibiae and tarsomeres 1–4 with stiff apical setae, mid and hind tibiae also with thick setae ventrally in distal half or so, basitarsi shorter than or (fore basitarsus) subequal to tarsomeres 2–4 combined, tarsomeres 3–4 almost equal in length, elongate, claw simple, arolium about as long as claw. Metasoma with petiolar segments very long, with no rough sculpture apparent, first segment slightly longer (in holotype) or shorter (in QBC-16C) than second [Figs 7B, 8I (in QBC-16C), 8H (holotype)], petiolar attachment to gaster frontal. Ovipositor straight, partially external (longest visible in QBC-16C, Fig. 8I). Measurements. Holotype: body length, 1.4 mm, head length, 0.30 mm, antenna length, 0.51 mm, mesosoma length, 0.45 mm, forewing length, 0.80 mm, width, 0.36 mm, first petiolar segment length 0.12 mm, second petiolar segment length 0.14 mm, gaster length (ovipositor excluded) 0.37 mm; paratype QBC-16C: body length, 1.4 mm, head length 0.30 mm, antenna length, 0.60 mm, mesosoma length, 0.45 mm, forewing length, 0.65 mm, width, 0.30 mm, first petiolar segment length 0.15 mm, second petiolar segment length 0.10 mm, gaster length (ovipositor excluded) 0.37 mm; paratype QBC- 1B: body length, 1.4 mm, antenna (deformed) length 0.49 mm, forewing length 0.80 mm, width 0.33mm. Remarks. Holotype and two paratypes are incompletely preserved and so with many features obscure. Yet they are considered as conspecific based on their burial in one and the same piece of amber, their similarity in features less affected with postmortem deformation (wing venation, to an extent also antenna and legs, and, to our wonder, most measurements) and apparent absence of counter-evidence of their conspecifity.Published as part of RASNITSYN, ALEXANDR P., MAALOUF, MOUNIR, MAALOUF, RAMY & AZAR, DANY, 2022, New Serphitidae and Gallorommatidae (Insecta: Hymenoptera: Microprocta) in the Early Cretaceous Lebanese amber, pp. 120-136 in Palaeoentomology 5 (2) on pages 126-128, DOI: 10.11646/palaeoentomology.5.2.4, http://zenodo.org/record/653041

    Leptoserphites pabloi RASNITSYN & MAALOUF & MAALOUF & AZAR 2022, gen. nov.

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    Leptoserphites pabloi Rasnitsyn & Azar, gen. nov. (Figs 3–5) Type material. Holotype, female, specimen no QBC-9A (complete insect suffered from sidewise compression and partially enclosed in a bubble obscuring some body parts, particularly on head and mesosoma), Maalouf collection; from Qanat Bakish (Baskinta) amber outcrop, Mouhafazet Jabal Loubnan (Governorate Mount Lebanon), Caza (= District) El-Maten, Central Lebanon; lower Barremian. Type material is deposited in the Natural History Museum of the Lebanese University, Faculty of Sciences II, Fanar, Lebanon. Etymology. Specific name is after Pablo Maalouf Alonso, son of one of us (RM), discoverer of the amber site. Diagnosis. Differs from the only congener in having antenna either distinctly 12-segmented or with clear traces of two basal flagellomeres fused secondarily into one, flagellomeres shorter, propodeum smooth, pterostigma narrower, 2r-rs longer, tubular (very dark) dot at M+Cu apex developed, hind wing r-m present, long, petiolar segments shorter. Locality and horizon. Baskinta, in the locality of Qanat Bakish (33°57′3″N, 35°47′24″E, elev. 1,488 m), Caza (= District) El-Maten, Mouhafazet (= Governorate) Mount Lebanon, in Central Lebanon; lower Barremian. Description. Female. Colour dark fuscous as preserved, metasoma and legs slightly paler. Body with no surface sculpture and prominent vestiture discernible. Head elongate as preserved (possibly in part due to deformation), with eyes small, not much convex, placed near middle of head sides (Figs 4A, 5B). Antenna 12- segmented (Fig. 5A), as preserved with scape long and wide, pedicel longer than any flagellar segment, flagellum wide, slightly narrowing toward base and apex, all segments wider than long except longest apical one, basal flagellomeres particularly short (width of segments could be exaggerated by depression due to diagenetic deformation, unless antennae are naturally flattened). Distinct large subequal mandibles, right with three strong and sharp teeth (visible in an aspect difficult to illustrate) and left with two strong and sharp ones. Maxillary palpi at least 4-segmented; labial palpi apparently 3-segmented, all segments thin. Mesosoma with dorsal surface deformed and poorly visible, dorsal pronotal part apparently moderately long (longer than in other Serphitidae), in side view metascutum forming high step above wide trough of metapostnotum, propodeum short, smoothly arching toward metasomal base. Forewing (Figs 4B, 5C) with C, R behind pterostigma, RS before and behind 2r-rs (except for short section forming T together with 2r-rs) and M lost as pigmented veins or nearly so; R, pterostigma and dot at junction of M+Cu and RS&M tubular; RS&M, M+Cu and 1-Cu nebulous; pterostigma low (twice as long as wide); 2r-rs as long as pterostigma wide, together with pterostigma enclosed in lightly pigmented area; all membrane covered with short setae (margin included). Hind wing (Figs 4B, 5D) with only R and r-m pigmented (tubular), R apex with three hamuli, r-m reaching wing hind margin (posterior wing area lost); membrane setose (much as in forewing), hind margin with fringe of long setae. Legs ordinary except that fore tibial spur (calcar) massive, strongly bent subapically and bifid, and fore basitarsus excised almost for all its length and with single row of setae along excision (so the antenna cleaning apparatus more characteristic of Mymarommatoidea than of Serphitidae); tibial spurs 1: 1?: 1?; basitarsus shorter than tarsomeres 2–4 combined, tarsomeres 3–4 almost equal in length, elongate, claw simple, arolium about as long as claw. Metasoma with petiolar segments elongate, with no rough sculpture apparent, first segment slightly longer and narrower than second, petiolar attachment to gaster frontal, metasomal terga ratio as preserved 1: 0.95: 1.4: 1.0: 0.9: 0.6 (gaster possibly extended longitudinally, secondarily). Ovipositor extruded (Figs 4D, 5E), stylets broken basally and bent down but not torn off, 1.5× as long as petiolar segments combined, very narrow supposed sheaths 0.25× as long as stylets, moderately wide, thin, convex transversally. Measurements. Length of body (as preserved, up to apex of metasomal tergum 7) 1.5 mm, head 0.28 mm, mesosoma 0.49 mm, forewing 0.80 mm, metasoma 0.78 mm, petiole 1 0.13 mm, petiole 2 0.10 mm.Published as part of RASNITSYN, ALEXANDR P., MAALOUF, MOUNIR, MAALOUF, RAMY & AZAR, DANY, 2022, New Serphitidae and Gallorommatidae (Insecta: Hymenoptera: Microprocta) in the Early Cretaceous Lebanese amber, pp. 120-136 in Palaeoentomology 5 (2) on pages 124-126, DOI: 10.11646/palaeoentomology.5.2.4, http://zenodo.org/record/653041

    Cretaceomma libanensis RASNITSYN & MAALOUF & MAALOUF & AZAR 2022, sp. nov.

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    Cretaceomma libanensis Rasnitsyn & Azar, sp. nov. (Figs 9, 10) Type material. Holotype, specimen no 1922, Azar collection, well preserved female; from Mdeirij-Hammana amber outcrop; Mouhafazet Jabal Loubnan (Governorate Mount Lebanon), Caza (= District) Baabda, Central Lebanon; lower Barremian. Type material is deposited in the Natural History Museum of the Lebanese University, Faculty of Sciences II, Fanar, Lebanon. Holotype. Etymology. Specific name is after the country of origin. Diagnosis. Same as for the genus and head elongate with eyes small, claval segments all elongate, probasitarsus with row of long setae, and first petiolar segment three times longer than high. Locality and horizon. Mdeirij-Hammana outcrop (33°48′13″N, 35°43′45″E, elev. 1,340 m), Caza (= District) Baabda, Central Lebanon, lower Barremian. Description. Colour dark, integument with weak metallic shine. Head and most of mesosoma mainly with transverse sculpture (finely distantly costulate to use terms by Harris, 1979), shining between costulae, with propodeal dorsum smooth, shining, legs and gaster dull. Prominent vestiture on antennae, legs, wings and metasomal apex. Head some 1.5× as long as wide (Fig. 10A), eye some half as long as head, weakly convex, distant from head posterolateral end for about its length, not reaching dorsal head surface, with some 6×10 rows of ommatidia; ocelli not distinct; antennal attachments well above upper eye margin, less than their diameters apart; frons very high (possibly with clypeus which base not distinct). Antenna 14-segmented (Fig. 10B), with segment length rates (scape without radix) = 1: 0.77: 0.51: 0.51: 0.60: 0.60: 0.70: 0.77: 0.71: 0.70: 0.75: 0.73: 0.66: 0.77: 0.86, scape and pedicel some 1.5× as wide as funicular segments (except somewhat wide ultimate one) and some 0.7 as wide as claval segments, all segments longer than wide including claval ones; claval segments with trichoid sensilla and thicker, bent, semiprocumbent ones. Mouthparts not visible. Mesosoma (Fig. 10C) with pronotum visible from above as narrow strip, apparently reaching wing bases, with lateral lobes longitudinally costulate, with ventral angles short,wide.Propleura not forming neck,prosternum external, both smooth. Mesonotum convex, smooth with sparse transverse costulae, lacking notauli and apparently parapsids; mesoscutellum subquadrate, with two basal foveae, metanotum half as long as mesoscutellum, entire, apparently smooth; metapostnotum weakly differentiated at propodeal base, propodeum transversely costate, evenly bent toward metasomal base. Mesopleura convex laterally, with longitudinal impression through its entire length, apparently smooth. Forewing (Fig. 10D) stalked, with only tubular R reaching basal wing quarter only, with membrane surface not distinctly modified, densely setose except for stalk, non-stalk margin entirely setose, with no single individualized seta, with distal and posterior setae very long and deeply rooted in membrane except those less long and short rooted in basal half of hind margin, with fore margin bearing both long and short setation, short setae being basal to long ones. Hind wing short, with stalk only (no membrane), with details obscured by attached tiny segmented object of obscure nature. Legs elongate (femora slightly inflated), ordinary except that fore coxa transverse costate; femora (dorsal), tibiae and tarsi setose, each tibia with one spur, fore spur straight, bent at distal third, bifid (Fig. 10E); all tarsomeres much longer than wide, basitarsi barely or not at all as long as two next segments combined, fore basitarsus concave over all ventral surface, with row of 12 or so long setae, claw small, simple, arolium long, narrow; fore tarsomere length ratio = 1: 0.50: 0.51: 0.51: 0.63, mid one = 1: 0.61: 0.55: 0.54: 0.63. Metasoma with narrow petiolar segments (Fig. 10F), first 2.9× as long as high, slightly narrowing to both ends, second 1.5× as long as high, slightly narrowing anterior, 0.4× as long as first, attaching to gaster from below; gaster (Fig. 10G) with first tergum slightly longer than petiolar segments combined, taking half gaster length, twice as long as second one, following terga short externally. Ovipositor scarcely visible externally, cerci with long setae. Measurements. Body length 0.69 mm, head length 0.14 mm, antenna length 0.47 mm, mesosoma length 0.24 mm, forewing length, 0.53 mm, width, 0.18 mm, petiolar segments 1, 2, 0.16 mm and 0.10 mm, respectively. Remarks. New species differs from the only congener Cretaceomma turolensis (Ortega-Blanco et al., 2011a) in body longer (0.7 vs. 0.5 mm) and more elongate, with head elongate (vs. semiglobose) and eyes small (vs. taking most of head sides), claval segments all elongate (vs. quadrate to transverse), probasitarsus with row of long (vs. short) setae, and first petiolar segment three times (vs. scarcely twice) longer than high.Published as part of RASNITSYN, ALEXANDR P., MAALOUF, MOUNIR, MAALOUF, RAMY & AZAR, DANY, 2022, New Serphitidae and Gallorommatidae (Insecta: Hymenoptera: Microprocta) in the Early Cretaceous Lebanese amber, pp. 120-136 in Palaeoentomology 5 (2) on page 130, DOI: 10.11646/palaeoentomology.5.2.4, http://zenodo.org/record/653041

    Microserphites libanensis RASNITSYN & MAALOUF & MAALOUF & AZAR 2022, sp. nov.

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    Microserphites libanensis Rasnitsyn & Azar, sp. nov. (Figs 1, 2) Type material. Holotype, specimen no 1819V, Azar collection; from Mdeirij-Hammana amber outcrop; Mouhafazet Jabal Loubnan (Governorate Mount Lebanon), Caza (= District) Baabda, Central Lebanon; lower Barremian. Type material is deposited in the Natural History Museum of the Lebanese University, Faculty of Sciences II, Fanar, Lebanon. Etymology. The specific name is derived from Lebanon in Greek, Λίβανος = Libanos. Diagnosis. New species differs from both its congeners in further reduced venation (all veins other than R nebulous or lost, pterostigma reduced up to a colored strip, hind wing with only R preserved and with membrane almost lost basally, and with R not reaching wing midlength) as well as in long, helmet-like head with flat eyes, in smaller size (forewing length under 0.5 mm), and from M. soplaensis also in flagellum much less widened apically. Locality and horizon. Mdeirij-Hammana outcrop (33°48′13″N, 35°43′45″E, elev. 1,340 m), Caza (= District) Baabda, Central Lebanon, lower Barremian. Description. Head, meso- and metasoma brownish, antennae and legs pale, petiole pale brown. Integuments mainly smooth, mesonotum between notauli rugose, propodeum carinate. Vestiture not apparent except wings with abundant longer marginal and shorter surface setae. Head dorsally long trapezoid with straight sides and deeply concave hind margin (Fig. 2A), eyes almost perfectly flat, taking most part of head sides, ocelli large, in almost equilateral triangle, with lateral ocelli close to but not touching eyes, occipital carina not apparent. Antenna 9-segmented (Fig. 2A, B), scape hardly visible, pedicel longer than wide, wider that flagellomeres except apical ones, flagellum gradually widening apicad, with flagellomeres changing from distinctly transverse basal ones to distinctly elongate apical one. Mesosoma with mesonotum strongly convex, pronotum forming wide belt on its fore declivity, posterolaterally not reaching wing base (tegula not identifiable), mesonotum with notauli very wide, not reaching rather long and transcarinate prescutellar fossa, mesoscutellum long with distinct posterior declivity, mesopleuron with complete oblique suture, metanotum following mesoscutellar declivity, with short medial upward tooth, propodeum short, similarly declivitous, with several short medial upward teeth (three, one apicad and two in distal third) and with oblique lateral carina. Legs weak, with coxae small, trochantelli developed, femora and tibiae narrow, femora not much longer than tibiae, tarsi not much longer than tibiae, tarsal segments all elongate, basitarsi much shorter than respective 3 middle tarsal segments but clearly longer than each of them, segment 4 scarcely shorter than segment 3, tibial spurs 1: 1: 2; claw simple, arolium present. Fore wing short, wide, with narrow base embracing all venation (nebulous M+Cu, basal vein [= RS&M] and Cu) except the only tubular vein R and weakly colored strip-like rudiment of pterostigma not reaching basal 0.4 of wing length; pre-radial (costal) membrane not visible. Hind wing with the only vein R present (Figs 1C, 2C), bearing 2 hamuli apical and hardly reaching wing midlength, with wing membrane very narrow before 0.4 wing length and parallel-sided with rounded apex further apical. Metasoma with petiolar segments tubular, of subequal height, first one some twice as long as high, second subquadrate; gaster with petiolar attachment ventral (not frontal), gastral terga 2–3 subequal in length and distinctly shorter than tergum 1; form of metasomal apex and sex unknown. Measurements. Body length 0.65 mm, fore wing 0.47 mm, hind wing 0.33 mm. Remarks. The new species is ascribed to Microserphites based on pronotum not reaching wing base and much reduced wing venation (pterostigma disintegrated, no tubular veins but R, etc.). It is described as new because of deep differences in wing venation and head form (cf. Diagnosis). Worth mentioning is that this oldest member of the genus is also the most modified venationally.Published as part of RASNITSYN, ALEXANDR P., MAALOUF, MOUNIR, MAALOUF, RAMY & AZAR, DANY, 2022, New Serphitidae and Gallorommatidae (Insecta: Hymenoptera: Microprocta) in the Early Cretaceous Lebanese amber, pp. 120-136 in Palaeoentomology 5 (2) on pages 121-123, DOI: 10.11646/palaeoentomology.5.2.4, http://zenodo.org/record/653041

    Aenigmaraphidophora mouniri AZAR & MAALOUF & NEL 2022, sp. nov.

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    Aenigmaraphidophora mouniri sp. nov. (Figs 1–3) urn:lsid:zoobank.org:act: DF023468-4A2E-45B3-8D0B- 13826D4F2F13 Material. Holotype of Aenigmaraphidophora mouniri gen. et sp. nov. (a nearly complete fossil, either a larva or a male), specimen number BKT-3A, Maalouf Collection, is stored at the Natural History Museum of the Lebanese University, Faculty of Sciences II, Fanar, Lebanon. Etymology. Named after Dr Mounir Maalouf, father of RM and discoverer of the amber site and studied material. Diagnosis. As for the genus, vide supra. Type locality and horizon. Bqaatouta amber outcrop (33°58′0″N, 35°47′13″E, elev. 1,177 m), Caza (= District) Kesserouan, Central Lebanon, lower Barremian. Description. Body 2.61 mm long (from head tip to abdomen); head 1.43 mm long; clypeus divided into an ante- and a postclypeus; antenna with 55 segments, 8.46 mm long; maxillary palp 2.15 mm long; apical sensory zone of maxillary palp elongate (Fig. 2B), much longer than palp apical width; thorax 0.97 mm long, 0.92 mm high; four tarsomeres on all legs; fore leg with tibia 0.95 mm long, tarsus 0.4 mm long, distal articulation zone of fore coxa displaced laterally; tympan present on fore tibia (Fig. 2C); mid femur 2.53 long, with one inner and one outer relatively large movable apical spines, tibia 0.91 mm long, tarsus 0.55 mm long; hind femur 2.06 mm long, with outer side without any chevron ridges, tibia 1.76 mm long, with longest apical spur slightly shorter than first and second segments of hind tarsi combined, hind tarsi laterally compressed, 1 st tarsomere with one flat pulvillus, 2 nd and 3 rd tarsomeres with a widened pulvilllus; abdomen 1.25 mm long, 1.3 mm high; no femoro-abdominal stridulatory apparatus observable; abdominal tergites without obvious process on dorsal surface; epiproct without posterior process; cerci 0.95 mm long, straight, bearing very long setae, but without any bulbous-like sensillae; subgenital plate with styles (Fig. 3C).Published as part of AZAR, DANY, MAALOUF, RAMY & NEL, ANDRÉ, 2022, An enigmatic Tettigoniidea from the Lower Cretaceous amber of Bqaatouta, Lebanon (Orthoptera, Ensifera), pp. 233-239 in Palaeoentomology 5 (3) on page 234, DOI: 10.11646/palaeoentomology.5.3.5, http://zenodo.org/record/682083

    New material of Paleopsychoda jarzembowskii Azar & Maksoud, 2022 from Bqaatouta amber outcrop, showing the importance of insect fossils in biostratigraphy

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    HOSSNY, TAMARA EL, MAALOUF, MOUNIR, MAALOUF, RAMY, AZAR, DANY (2023): New material of Paleopsychoda jarzembowskii Azar & Maksoud, 2022 from Bqaatouta amber outcrop, showing the importance of insect fossils in biostratigraphy. Palaeoentomology 6 (1): 8-12, DOI: 10.11646/palaeoentomology.6.1.3, URL: http://dx.doi.org/10.11646/palaeoentomology.6.1.

    Libanophlebotomites ramyi AZAR & MAALOUF & MAKSOUD 2022, sp. nov.

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    Libanophlebotomites ramyi sp. nov. (Figs 1–6) Etymology. The species epithet is in honor of Ramy Maalouf, son of one of us (MM). Diagnosis. As for the genus (vide supra). Locality and horizon. Qanat Bakish, Baskinta; Caza (= District) El-Maten; Mouhafazet (= Governorate) Mount Lebanon; central Lebanon (pictures of the site and geological map are given in Maksoud et al., 2021: 368, fig. 1), lower Barremian (Granier et al., 2016; Maksoud et al., 2017, 2021). Type material. Female holotype specimen QBC 42A (Maalouf collection) from Qanat Bakish, Baskinta outcrop (central Lebanon), deposited in the Museum of Natural History of the Lebanese University, Faculty of Sciences II, Fanar, Lebanon. Description. Head 0.280 mm long (without mouthparts) (Fig. 2A); mouthparts well developed (0.15 mm long), with sharp and elongate mandibles and laciniae (Fig. 2B), palp 0.325 mm long, with five palpomeres (Fig. 3A, B), with palpomeres II and III longest, palpomeres I–V lengths respectively 0.038-0.091-0.088- 0.047- 0.061 mm; eyes rounded, 0.18 mm of diameter; antenna 1.070 mm long, with 15 flagellomeres (Fig. 4A, B); first flagellomere comparatively short, almost as long as following flagellomere; last flagellomere (apiculus) short and drop like (Fig. 4A, B); flagellomere I–XV lengths respectively:0.150-0.093-0.093-0.087-0.081-0.075-0.075- 0.068-0.068-0.059-0.056-0.050-0.050-0.050-0.015; scape nearly cylindrical, 0.050 mm long. Pedicel ellipsoid, 0.093 mm long; all flagellomeres bearing curved setae, ascoids are not discernable. Thorax 0.820 mm long, 0.500 mm high, not well preserved; proscutum and scutum not strongly gibbous; legs slightly shorter than entire body; tibiae of forelegs each with at least three strong and long setae aligned along the inner side. Wing 1.420 mm long, 0.450 mm wide, hyaline (Fig. 5A, B); humeral vein reaching costal margin; subcostal vein (Sc) distally fused with R1, 0.450 mm from wing base, and with crossvein reaching costal margin; R1 reaching costal margin 0.960 mm from wing base; Rs four-branched, with all its branches extending to wing margin; R2 and R3 separating 0.930 mm distally; R4 and R5 almost straight, reaching wing margin respectively at 1.400 and 1.410 mm apically; M1 and M2 bifurcating 0.900 mm distally, reaching wing margin respectively at 1.400 and 1.320 mm; M1 distally nearly straight; M2 slightly shorter than M1; M3 reaching wing margin at 1.150 mm from wing base; M4 [CuA 1] reaching wing margin 1.010 mm distad wing base. CuA [CuA 2] (Cu after Byers et al., 1989; Krzemiński & Krzemińska, 2003) short and week reaching posterior margin of wing. All main veins and wing margin bearing long macrotrichiae. Halteres 0.300 mm long; knob fusiform 0.125 mm long. Abdomen 0.930 mm long including genital appendages, dorsal surfaces of all abdominal segments bearing few setae. Female genital appendages (Fig. 6A, B) with cercus elliptic (0.097 mm long); male unknown.Published as part of AZAR, DANY, MAALOUF, MOUNIR & MAKSOUD, SIBELLE, 2022, Libanophlebotomites ramyi, a new genus and species of phlebotomine sandflies from Lower Cretaceous Lebanese amber (Diptera: Psychodidae: Phlebotominae), pp. 340-346 in Palaeoentomology 5 (4) on pages 342-343, DOI: 10.11646/palaeoentomology.5.4.6, http://zenodo.org/record/733355

    Paleopsychoda jarzembowskii Azar & Maksoud 2022

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    <i>Paleopsychoda jarzembowskii</i> Azar & Maksoud, 2022 <p> <b>Type material.</b> Holotype 1615A, from Mdeyrij-Hammana, Central Lebanon, Azar collection, deposited in the Natural History Museum of the Lebanese University, Faculty of Sciences II, Fanar, Lebanon.</p> <p> <b>New material.</b> Specimen BKT-9A, Maalouf collection, from Bqaatouta, deposited in the Natural History Museum of the Lebanese University, Faculty of Sciences II, Fanar, Lebanon.</p> <p> <b>Locality and horizon.</b> Bqaatouta, Caza (= District) Kesserouan (Central Lebanon), lower Barremian (Granier <i>et al</i>., 2016; Maksoud <i>et al</i>., 2017, 2022).</p> <p> <b>Description.</b> The specimen of Bqaatouta is a complete sample but somehow with incomplete antennae and it carries all the diagnostic features of <i>Paleopsychoda jarzembowskii</i>, <i>i</i>. <i>e</i>., eye-bridge incomplete, mouthparts toothed, welldeveloped and of phlebotomine-type, with sharp toothed mandibles and laciniae; maxillary palp four-segmented, with last palpomere twice as long as others and presenting a constriction in its basal third; anterior sub-basal part of costal vein inflated, from either side of end of humeral vein, allowing as such larger surface, with insertions of a basal wing tuft of setae (specific male character); R1 reaching costa beyond level of radial fork; M3 and M4 [CuA1] meeting basally for a very small distance; CuA [CuA2] well-developed, reaching wing margin beyond half of wing; CuP [A] well developed; male genitalia with stylus bearing two apical spines; a specialized long seta strongly curved in its middle part arising between gonocoxite and gonostylus; parameres paired with apically series of four strong setae with incurved apices; surstyli joined basally, bearing apically two strong setae on each. Measurements of the different body parts are very similar to the type specimen, the holotype number 1615A.</p>Published as part of <i>HOSSNY, TAMARA EL, MAALOUF, MOUNIR, MAALOUF, RAMY & AZAR, DANY, 2023, New material of Paleopsychoda jarzembowskii Azar & Maksoud, 2022 from Bqaatouta amber outcrop, showing the importance of insect fossils in biostratigraphy, pp. 8-12 in Palaeoentomology 6 (1)</i> on page 9, DOI: 10.11646/palaeoentomology.6.1.3, <a href="http://zenodo.org/record/7754898">http://zenodo.org/record/7754898</a&gt
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