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    Figs 93–100. Live specimens. 93–96 in New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae)

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    Figs 93–100. Live specimens. 93–96. Pholcus tambunan Huber sp. nov., Crocker Range, ♂, penultimate ♂, and ♀ with egg-sac. 97–100. P. bario Huber sp. nov., Bario, ♂ and ♀ with egg-sac.Published as part of Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200 on page 26, DOI: 10.5852/ejt.2016.200, http://zenodo.org/record/89721

    Figs 59–66. Live specimens. 59–62 in New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae)

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    Figs 59–66. Live specimens. 59–62. Pholcus bukittimah Huber sp. nov., Dairy Farm, ♂, penultimate ♂, and ♀ with partly parasitized egg-sac. 63–66. P. barisan Huber sp. nov., Bukit Barisan, ♂ and ♀♀ with variably expanded abdomens.Published as part of Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200 on page 18, DOI: 10.5852/ejt.2016.200, http://zenodo.org/record/89721

    Figs 2–9. Live specimens. 2–3. Pholcus phui Huber, 2011 in New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae)

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    Figs 2–9. Live specimens. 2–3. Pholcus phui Huber, 2011, Hala Bala, ♂ and ♀ with egg-sac. 4–7. P. tanahrata Huber sp. nov., Cameron Highlands, ♂, penultimate ♂, and ♀. 8–9. P. uludong Huber sp. nov., Ulu Dong, ♂ and ♀.Published as part of Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200 on page 6, DOI: 10.5852/ejt.2016.200, http://zenodo.org/record/89721

    Figs 32–38. Live specimens. 32–35. Pholcus gombak Huber, 2011 in New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae)

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    Figs 32–38. Live specimens. 32–35. Pholcus gombak Huber, 2011, Kemensah (32) and Gunung Liang (33–35), ♂, ♀ with parasitized egg-sac seven days before eclosion of wasps (33), one day before eclosion (34), and at eclosion (35). 36–38. P. ledang Huber, 2011, Gunung Ledang, ♂ and ♀ with egg-sac.Published as part of Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200 on page 14, DOI: 10.5852/ejt.2016.200, http://zenodo.org/record/89721

    Figs 39–47. Pholcus gombak Huber, 2011 in New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae)

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    Figs 39–47. Pholcus gombak Huber, 2011, SEM micrographs (ZFMK, Ar 15701–02). 39–40. Male and female prosomata, frontal views. 41. Left procursus; asterisk marks large prolatero-dorsal process. 42– 43. Right appendix and embolus, prolateral and prolatero-distal views. 44. Female ALS. 45. Cuticular processes on whitish retrolateral area of left procursus. 46. Male gonopore. 47. Epigynum, ventral view. Scale bars: 39–40 = 300 µm; 41, 47 = 200 µm; 42, 43 = 100 µm; 44 = 20 µm; 45 = 10 µm; 46 = 30 µm.Published as part of Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200 on page 15, DOI: 10.5852/ejt.2016.200, http://zenodo.org/record/89721

    Figs 54–58. Pholcus ledang Huber, 2011 in New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae)

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    Figs 54–58. Pholcus ledang Huber, 2011, SEM micrographs (ZFMK, Ar 15704–05). 54. Left procursus, retrolateral view. 55. Right procursus, distal view. 56. Cuticular processes on whitish retrolateral area of left procursus. 57. Male gonopore. 58. Epigynum, ventral view. Scale bars: 54, 58 = 200 µm; 55 = 100 µm; 56 = 10 µm; 57 = 30 µm.Published as part of Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200 on page 17, DOI: 10.5852/ejt.2016.200, http://zenodo.org/record/89721

    Pholcus domingo

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    Pholcus domingo species group Diagnosis The two species included in this group are medium-sized, long-legged spiders (body length ~4.5, male leg 1 length:~35–40), distinguished from similar P. kerinci group by bipartite appendix (sclerotized retrolateral and whitish prolateral part; Figs 140, 150, 157), by shape of procursus (presence of dorsal process; absence of distal spine-like process; Figs 141, 158), and by dorsal trichobothrium of male palpal tibia in ‘usual’ position (i.e., at ~45% of tibia length, versus 55–60% in P. kerinci group); from other species groups in Pholcus by combination of the following characters: elongate abdomen angular dorso-posteriorly (Fig. 139); eight eyes; male chelicerae almost unmodified (indistinct lateral humps proximally; Fig. 142); male genital bulb without uncus; epigynum weakly sclerotized, with numerous transversal folds (Figs 143, 156, 159). Description See individual descriptions of the two very similar species below. Composition Only the two species described below. Natural history Both species were found on the undersides of live leaves where they built very fine webs tightly attached to the leaf surface. For further information, see description of P. domingo Huber sp. nov. below. Distribution The P. domingo group is restricted to Mindanao (Fig. 1).Published as part of Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200 on page 34, DOI: 10.5852/ejt.2016.200, http://zenodo.org/record/89721

    Pholcus phui Huber 2011

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    <i>Pholcus phui</i> Huber, 2011 <p>Figs 2–3</p> <p> <i>Pholcus phui</i> Huber, 2011: 178 –180, figs 749–752, 779–780, 809–813 (♂♀).</p> <p> <b>Diagnosis</b></p> <p> Easily distinguished from most similar known relatives (other species of the <i>P. ethagala</i> group on the Malay Peninsula) by morphology of male palps (figs 809–810 in Huber 2011; procursus with two distinctive dorsal processes and curved distal process; large appendix distally with whitish element and with sclerotized element set with small scales), and by details of female internal genitalia (figs 812–813 in Huber 2011; distinctive pair of anterior lateral structures; roundish pore plates close together).</p> New material examined <p>THAILAND: 2 ♂♂, 1 ♀, 1 juv., ZFMK (Ar 15691), and 1 ♂, 1 ♀, PSUZC, Narathiwat Province, Hala Bala Wildlife Sanctuary, ‘site 3’ (5.808°N, 101.823°E), 220 m a.s.l., forest near road, leaf litter, 2 Mar. 2015 (B.A. Huber, B. Petcharad); 4 ♀♀, 3 juvs, in absolute ethanol, ZFMK (Mal 309), same data.</p> Description – amendments <p>Tibia 1 in three newly collected males: 9.3, 9.6, 9.7; tibia 2/tibia 4 length: 1.08.</p> Natural history <p> All specimens were collected within a small swampy area in the forest that was characterized by numerous large dead leaves on the ground. Almost each of the largest leaves had an adult <i>Pholcus phui</i> on its lower surface, while the more numerous smaller leaves were mostly either unoccupied or occupied by juveniles. No further specimens were found during three days of collecting in various parts of the forest.</p>Published as part of <i>Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200</i> on pages 5-6, DOI: 10.5852/ejt.2016.200, <a href="http://zenodo.org/record/897219">http://zenodo.org/record/897219</a&gt

    Pholcus bario Huber, sp. nov.

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    Pholcus bario Huber sp. nov. urn:lsid:zoobank.org:act:D680CB67-1525-4263-B8D4-8E44E587C51E Figs 97–100, 121–124, 128–130 Diagnosis Easily distinguished from most similar known relative (P. tambunan Huber sp. nov.) by morphology of male palps (Figs 121–122; longer trochanter apophysis; more slender femur; distinctive structures on tip of procursus), by different shapes of frontal cheliceral apophyses (Fig. 120), and by longer epigynal scape (Fig. 123). Etymology The species name is derived from the type locality; noun in apposition. Material examined Holotype MALAYSIA-BORNEO: ♂, ZFMK (Ar 15716), Sarawak, Bario, forest along river W of town (3.736°N, 115.437– 115.443°E), 1150–1250 m a.s.l., on underside of leaf, 30 Jul. 2014 (B.A. Huber, S.B. Huber). Other material MALAYSIA-BORNEO: 2 ♂♂, 2 ♀♀, ZFMK (Ar 15717–18), same data as holotype; 2 ♀♀, in absolute ethanol, ZFMK (Bor 232), same data. Description Male (holotype) MEASUREMENTS. Total body length 3.9, carapace width 0.9. Leg 1: 34.5 (8.1 + 0.4 + 8.2 + 15.5 + 2.3), tibia 2: 5.3, tibia 3: 3.2, tibia 4: 4.7; tibia 1 L/d: 104. Distance PME-PME 370 µm, diameter PME 95 µm, distance PME-ALE ~35 µm; no trace of AME. COLOR. Carapace whitish with dark median mark widening anteriorly and including ocular area and clypeus in upper part and laterally; sternum whitish; legs pale ochre-yellow with dark brown patellae and tibia-metatarsus joints; abdomen pale ochre-gray with black marks dorsally and laterally, monochromous ventrally. BODY. Habitus as in Figs 97–99; ocular area slightly raised, each triad on additional short stalk directed towards lateral (Fig. 120); carapace without median furrow; clypeus unmodified; sternum wider than long (0.64/0.52), unmodified. CHELICERAE. As in Fig. 120, with distinctive pair of frontal apophyses (large flat curved plates) and small rounded lateral humps; without modified hairs; without stridulatory ridges. PALPS. As in Figs 121–122; coxa unmodified; trochanter with strong ventral apophysis; femur cylindrical, with two small retrolateral humps proximally and small ventral hump at half length; procursus with strong ventral ‘knee’, with distinctive small prolateral sclerotized process at 1/3 length (arrow in Fig. 121), distal half with large whitish area retrolaterally, tip complex with distinctive sclerites; bulb elongated, long appendix with distinctive prolateral process provided with small teeth, further teeth along ventral rim, proximal processes of appendix ‘guiding’ embolus between them; embolus long and weakly sclerotized. LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments not seen in dissecting microscope. Male (variation) Tibia 1 in 2 other males: 7.8, 8.0. Female In general similar to male (Fig. 100) but carapace mark only in posterior half; ocular area dark as in male but clypeus light. Eye triads closer together than in male (PME-PME distance: 230 µm). Dark marks on abdomen less distinct. Tibia 1 in 2 females: 6.6 (both). Epigynum weakly sclerotized (Fig. 128), wide transversal plate with long scape directed towards anterior (Fig. 123); dark internal arch and lateral posterior structures visible through cuticle; internal genitalia as in Figs 124, 129, 130, with pair of highly distinctive three-layered telescopic tubes. Natural history All specimens were found in one small section of forest along a small river. No further specimens were found in two days of intensive search. They built barely visible webs on the undersides of live leaves, and often shared webs with large numbers of Diptera (Cecidomyiidae, in ZFMK). Distribution Known from type locality in northern Borneo (Sarawak) only (Fig. 1).Published as part of Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petcharad, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), pp. 1-45 in European Journal of Taxonomy 200 on pages 31-33, DOI: 10.5852/ejt.2016.200, http://zenodo.org/record/89721
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