808 research outputs found
Bezzia (Bezzia) libanensis Alwin-Kownacka & Szadziewski & Szwedo 2017, sp. nov.
Bezzia (Bezzia) libanensis Alwin-Kownacka & Szadziewski sp. nov. urn:lsid:zoobank.org:act: B1B97EC8-DFA5-4537-BB13-FAC1B73487C6 Fig. 1 Diagnosis A small dark brown species without fore femoral spines. The male genitalia are unique in having a greatly elongate sternite IX, with narrow V-shaped caudomedian cleft, and a Y-shaped aedeagus without ventral spicules and an abruptly tapered, sharply pointed apex. Female unknown. Etymology The specific name refers to the Latin name of Lebanon (Libanus), where the holotype was collected. Material examined Holotype LEBANON: ♂, Fanar, 33º52.680′ N, 35º33.547′ E, 138 m a.s.l, at light, 6 May 2012, P. Dominiak leg. (CEIUG). Description Male HEAD. Uniformly brown. Antenna (Fig. 1C) pale brown; flagellomeres 10–13 predominately dark brown; flagellomere 10 elongate, considerably longer than 11–13; total flagellum length 0.90 mm; antennal ratio 0.70. Palpus slender, brownish; segment 3 slender with few sensilla capitata; palpal ratio 4.00. THORAX (Fig. 1B). Dark brown. Wing (Fig. 1A) veins pale; 1 poorly marked elongate radial cell; wing length 1.30 mm; costal ratio 0.60. Halter brown. Legs (Figs. 1 D–F) with brown coxae; all femora without ventral spines; fore and mid femora yellowish, proximal ½ of venter brown and with subapical, apical dark brown bands; hind femur brown, with apical, subapical yellow bands; tibiae brown, with sub-basal, subapical yellow bands; tarsi yellow, with pale brown apices, tarsomeres 4–5 slightly darker than 1–3; tarsal ratio of fore leg 2.20, of mid leg 2.40, of hind leg 2.20. ABDOMEN (Fig. 1G). Brown. GENITALIA (Fig. 1 G–J). Sternite IX heavily sclerotized, long, apex extending to mid-length of gonocoxites, with narrow, V-shaped caudomedian cleft. Tergite IX short, slender, with elongate cerci. Gonocoxite short, stout; gonostylus moderately long, nearly straight, slightly curved distally, apex tapered, with pointed tip. Parameres (Fig. 1I) fused, rod-like; basal arms short, broad, bifurcate with unequal-length prongs; distal portion slightly vasiform, with heavily sclerotized margins, apical ⅓ narrower, more lightly sclerotized, with rounded tip. Aedeagus (Fig. 1J) with bare ventral membrane; basal arms broad, heavily sclerotized; basal arch curved, extending ⅓ of total aedeagus length; distal portion short, margin heavily sclerotized, with sharply pointed apex. Female Unknown. Distribution Lebanon; known only from the type locality. Remarks The male antennal flagellum of this new species has an elongate flagellomere 10 and a bare aedeagus, characters typical of members of the subgenus Bezzia. This new species differs from all other Palearctic and Afrotropical members of the genus by its greatly elongate sternite IX, which extends to the midlength of the gonocoxites and covers their bases.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2017, Predatory midges of the tribes Palpomyiini and Sphaeromiini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 1-30 in European Journal of Taxonomy 318 on pages 4-6, DOI: 10.5852/ejt.2017.318, http://zenodo.org/record/382745
Bezzia (Homobezzia) sharjahi Alwin-Kownacka & Szadziewski & Szwedo 2017, sp. nov.
Bezzia (Homobezzia) sharjahi Alwin-Kownacka & Szadziewski sp. nov. urn:lsid:zoobank.org:act: 07610F8D-C955-4FD8-851A-395BA06AFB87 Fig. 2 Diagnosis This new species differs from all other Middle East congeners in having the following combination of characters: thorax orange-brown with distinctly paler scutellum; legs uniformly brownish; fore femur with 6–9 ventral spines; male gonostylus long, with pointed apex; parameres long, slender, slightly curved dorsally; female with 2 subspherical seminal capsules with very long necks. Etymology The specific name refers to the Sharjah Desert, where most of the type series was collected. Material examined Holotype UNITED ARAB EMIRATES: ♂, adult, Sharjah Desert Park, 18 Jan. 2005, A. van Harten leg. (CEIUG). Paratypes UNITED ARAB EMIRATES: 3 ♀♀, 3 ♂♂, same collection data as holotype except 18–25 Jan. 2005, light trap; 3 ♀♀, 1 ♂, same collection data except 9 Mar. 2005; 1 ♀, same collection data except 30 Apr. 2005; 1 ♀, same collection data except 30 Jun. 2005; 1 ♀, 1 ♂, same collection data except 21 Jul. 2005; 1 ♀, Wadi Wurayah, light trap, 15 Jan.–22 Feb. 2009, A. van Harten leg. (CEIUG). YEMEN: 2 ♀♀, Al Mukalla, light trap, 1. Feb. 2003, A. van Harten leg. (CEIUG). Description Male HEAD. Uniformly pale brown. Antenna (Fig. 2B) pale brown; flagellar plume poorly developed; flagellomeres 2(3)–13 with paler basal halves; flagellomeres 10–13 short; total flagellum length 0.50– 0.70 mm; antennal ratio 0.50–0.70. Palpus yellow, slender; third palpal segment short, with distinct capitate sensilla; palpal ratio 2.00. THORAX. Orange-brown; scutellum paler. Wing pale, with barely visible veins; 1 poorly marked long radial cell; wing length 0.90–1.30 mm; costal ratio 0.70–0.80. Legs uniformly pale brown, tarsi slightly paler, coxae dark brown; claws small and equal on all legs; fore femur armed with 4–8 short ventral spines; tarsal ratio of fore leg 1.60–1.90, of mid leg 1.80–2.30, of hind leg 2.10–2.20. ABDOMEN. Orange brown. GENITALIA (Fig. 2 D–F). Tergite IX short, reaching apex of gonocoxite; sternite IX short. Gonocoxite short, stout; gonostylus long, narrow, distal half curved, with sharp apex. Parameres (Fig. 2E) fused, tongue-like, slightly curved dorsally, with blunt apex. Aedeagus (Fig. 2F) triangular, with low basal arch and barely visible rounded apex. Female HEAD. Brown. Antennal flagellum (Fig. 2C) with short flagellomeres, brown, proximal portions yellow; total flagellum length 0.50–0.70 mm; antennal ratio 0.80–1.10. Palpus slender, yellow; third palpal segment short, rather stout, without sensory pit; palpal ratio 2.00–2.50. Mandible with 6–8 massive distal teeth and numerous smaller proximal teeth. THORAX. Orange brown; scutellum paler. Wing (Fig. 2A) venation similar to that of male; wing length 1.70–1.80 mm; costal ratio 0.80. Femoral coloration as in male; fore femur armed with 6–9 ventral spines; fore and mid tibiae with slightly dark bases, occasionally with slightly darker central areas, and dark apices; hind tibia with slightly darker base; tarsal ratio of fore leg 1.80–2.10, of mid leg 1.90–2.50, of hind leg 2.00–2.50. ABDOMEN. Two subspherical seminal capsules (Fig. 2G) with long, thick necks; measuring 0.04–0.05 × 0.03–0.04 mm and 0.05–0.06 × 0.04–0.05 mm. Distribution United Arab Emirates and Yemen. Remarks This new species is a typical member of the subgenus Homobezzia. It most closely resembles B. kuhetiensis, which has similar male genitalia and seminal capsules. However, B. kuhetiensis differs from B. sharjahi sp. nov. by its whitish legs with distinct, dark bands, the fore femur has only 2 ventral spines, and the parameres are straight.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2017, Predatory midges of the tribes Palpomyiini and Sphaeromiini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 1-30 in European Journal of Taxonomy 318 on pages 7-10, DOI: 10.5852/ejt.2017.318, http://zenodo.org/record/382745
Ceratopogon azari Dominiak, Alwin & Gilka 2014
Ceratopogon azari Dominiak, Alwin & Giłka, 2014 Ceratopogon azari Dominiak, Alwin & Giłka, 2014: 138 (male, Lebanon). Diagnosis. Males differ from all other species of the genus by the following combination of characters: apicolateral processes of tergite IX elongate, each bearing two setae, parameres fused on proximal 1/3, aedeagus with seed-like medial prongs, lateral prongs lightly sclerotized with setose apices. Material examined. LEBANON, Anti-Lebanon Mts., Maaraboun village near Baalbek (in the locality named Sheaibe), N 33˚ 55.376 E 36˚15.490, altitude 1781 m, stream and helocrene, 3 males, 3 females, 5.V.2012, net, leg. P. Dominiak. Distribution. A montane species living in Anti-Lebanon Mountains. Discussion. Ceratopogon including 41 extant species (Borkent & Grogan 1995) is a relictual Boreal genus which has Holarctic distribution (Szadziewski 2008). The species C. azari extends the southernmost distribution for this genus in the Western Palaearctic. This montane species was collected at altitude 1781 m a.s.l.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2016, Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 551-572 in Zootaxa 4079 (5) on pages 560-561, DOI: 10.11646/zootaxa.4079.5.3, http://zenodo.org/record/105087
Kolenohelea levantica Alwin-Kownacka & Szadziewski & Szwedo 2016, sp. nov.
Kolenohelea levantica Szadziewski & Alwin sp. nov. (Figures 3, 4b) Diagnosis. Males of this species can be easy distinguished by having very long, blunt and singular or bilobed apicolateral processes, each bearing one small seta. Parameres slightly S-shaped, weakly pointed. Female unknown. Description. Male. Head. Uniformly brown. Antenna uniformly brown, all flagellomeres separated. Three distal flagellomeres elongated (Fig. 3a). Total length of flagellum 0.65–0.73 mm, AR 1.03–1.15. Palpus brown, slender, 0.18–0.23 mm long. Third palpal segment slender with distinct sensory pit (Fig. 3b), PR 2.5–3.5. Thorax. Dark brown. Scutum and scutellum with numerous setae. Wing pale, two radial cells well marked. First one narrow and about 1.6–1.9 times shorter than second one. Wing length 1.10–1.32 mm, CR 0.64. Legs brown and stout, tibiae armed with strong posterior bristles. Claws small and equal on all legs. Fore leg. Coxa slightly darker than rest of leg. Tarsus slightly paler than tibia. First tarsomere armed with one apical, one subbasal spines and 1–3 central, more delicate spines. Second and third tarsomeres each with one apical, very delicate spine. Fourth tarsomere cordiform, bearing ventrally at apex a pair of hyaline sensilla. TR(I) 2.0–2.1. Mid leg. Coxa and trochanter darker than rest of leg. First tarsomere armed with two apical, two subbasal and three central spines, all more delicate than on fore leg. Second tarsomere with two apical spines, third and fourth tarsomeres each with one apical spine. Fourth tarsomere with pair of hyaline sensilla at apex. TR(II) 1.9. Hind leg. Coxa and trochanter darker than rest of leg. Femur swollen. Tibial comb with seven large setae. Tarsus slightly paler than tibia. Basitarsus bent, with one complete row of palisade setae, one apical and one subbasal spine present. TR(III) 1.9– 2.0. Abdomen. Dark brown. Genitalia (Figs 3 c-g). Sternite IX short, with indistinct caudal margin. Tergite IX with very long (134 µm), blunt, singular or bilobed apicolateral processes; each bearing one small seta (Figs 3c, g). Gonocoxite 0.33 mm long, broad, slightly expanded at middle, with distinct ventral and dorsal apodemes (Fig. 3c, d). Gonostylus 0.24 mm long, almost straight, with blunt apex. Parameres slightly S-shaped, strongly sclerotized; apex weakly pointed (Figs 3e, 4b, c). Aedeagus triangular with bicorned apex and distinct ribs on lateromedian surfaces; basal arch very low (Fig. 3f). Female. Unknown. Distribution. Israel and Lebanon. Material examined. Holotype: male. ISRAEL. Nahal Qumeran, 22.III.1993, leg. A. Freidberg [TAU]. Paratype: LEBANON. Jezzine, Pont Al Khalass, 7.IX.2013, 1 male, leg. A. Alwin [CEI UG]. Etymology. Specific epithet is derived from historical name of the area of presence of the species—the Levant. Discussion. The holotype from Israel is distinctly darker and bigger than the paratype from Lebanon, moreover the holotype has bilobed apicolateral processes while in Lebanese specimen apicolateral processes are single. We treat these differences as infraspecific variability as noted in Kolenohelea calcarata (Szadziewski 1992). This species is very similar to Kolenohelea calcarata Goetghebuer, 1920 from the Palaearctic and K. leonina de Meillon & Wirth, 1987 and K. dycei de Meillon & Wirth, 1981 from the Afrotropical Region. Parameres in K. calcarata are almost straight to slightly C-shaped with flat sharp tips (Fig. 4a), while in K. levantica parameres are more stout and S-shaped, and their tips are cylindrical and more blunt (Figs 4b, c). K. leonina and K. dycei have S-shaped parameres and aedeagus with bilobed apex like in K. levantica. However, both African species have more slender parameres, especially at the base, and gonostyli armed with distinct apical teeth.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2016, Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 551-572 in Zootaxa 4079 (5) on pages 561-563, DOI: 10.11646/zootaxa.4079.5.3, http://zenodo.org/record/105087
FIGURE 3 in Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species
FIGURE 3. Kolenohelea levantica Szadziewski & Alwin sp. nov., male; a—flagellum, b—palpus, c,d—genitalia, eparameres, f—aedeagus, g—apicolateral process of tergite IX.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2016, Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 551-572 in Zootaxa 4079 (5) on page 562, DOI: 10.11646/zootaxa.4079.5.3, http://zenodo.org/record/105087
Palpomyia freidbergi Alwin-Kownacka & Szadziewski & Szwedo 2017, sp. nov.
Palpomyia freidbergi Alwin-Kownacka & Szadziewski sp. nov. urn:lsid:zoobank.org:act: 740DBF40-E804-477B-A437-E73110DF1287 Fig. 4 Diagnosis This new species is distinguished by the following combination of characters: males with 4–5 fore femoral ventral spines; very short sternite IX; parameres fused, curved ventrally, apex pointed; and aedeagus with very high basal arch and plate-like apex. Females with 6–9 fore femoral spines; claws without basal inner teeth; and 2 large seminal capsules. Etymology The species is named for Amnon Freidberg, Department of Zoology, Tel Aviv University, Israel, who kindly sent us biting midges from Israel. Material examined Holotype ISRAEL: ♂, Enot Zukim, 11 Oct. 1994, A. Freidberg and F. Kaplan leg. (TAU). Paratypes ISRAEL: 4 ♀♀, same collection data as holotype except 22 Apr. 1998, A. Freidberg leg. (TAU); 1 ♀, En Feshkha, 11 Aug. 1986, A. Freidberg leg. (TAU); 9 ♀♀, Roch Zukim, 10 Apr. 1994, A. Freidberg and F. Kaplan leg. (TAU). Description Male HEAD. Uniformly brown. Antenna pale brown, with sparse plume; flagellomeres 9–13 slightly darker than 1–8; flagellomeres 10–13 elongate; total flagellum length of 1.00 mm; antennal ratio 1.40. Palpus (Fig. 4C) yellowish, slender; third segment cylindrical, with several distinct capitate sensilla; palpal ratio 3.00. THORAX (Fig. 4B). Brown with scutum, scutellum, katepisternum and mediotergite slightly darker; scutum, scutellum, anteroanepisternum with fine setae, katepisternum and mediotergite bare; 1 row of 3 bristles posterior to sutura transveralis; 1 bristle on scutum just anterior of scutellum; scutellum with 4 marginal bristles. Wing (Fig. 4A) with distinct veins; first radial cell about 2 × shorter than second; wing length 1.34 mm; costal ratio 0.70. Legs (Fig. 4B) slender; fore coxa yellow, mid and hind coxae brownish; fore and mid femora yellow, hind femur yellow with dark brown apex; fore femur with 4–5 stout, black ventral spines; fore and mid tibiae yellow, slightly smoky on proximal portions; hind tibia uniformly dark brown; tarsi yellow, tarsomeres 4–5 slightly darker than 1–3; tarsal ratio of fore leg 1.90, of mid leg 3.30, of hind leg 2.40. ABDOMEN (Fig. 4B). Pale brown. GENITALIA (Fig. 4 E–G). Sternite IX (Fig. 4G) narrow. Gonocoxite slender, elongate; gonostylus swollen on basal portion, distal half slender, greatly curved, with blunt apex. Parameres (Fig. 4F) fused, base W-shaped, expanded on mid portion, apex pointed, bent ventrally. Aedeagus (Fig. 4G) triangular; basal arch very high; membrane with numerous fine microtrichia; apex plate-like. Female Similar to male with the usual sexual differences. Antenna uniformly pale brown; total flagellum length of 1.40–1.60 mm; antennal ratio 1.40–1.60. Palpal ratio 4.50. Mandible with 6–7 large teeth. THORAX. Coloration and arrangement of bristles as in male. Wing as in male but larger with longer costa (Fig. 4A); wing length 2.10–2.20 mm; costal ratio 0.80. Coloration of legs as in male; fore femur armed with 6–9 stout, black ventral spines (Fig. 4D); tarsal ratio of fore leg 2.10–2.40, of mid leg 3.80–4.50, of hind leg 2.30–2.50. ABDOMEN. Pale brown, with 2 pairs of internal tergal apodemes. Two large, subequal, ovoid seminal capsules (Fig. 4H) with short necks, measuring 0.08–0.10 × 0.12–0.14 mm and 0.06–0.08 × 0.10–0.12 mm. Distribution Israel; known only from the type locality. Remarks Males of this new species have an aedeagus with an unusual plate-like apex that is similar to that of males of P. mahyoubi from Yemen. This newly described species differs mostly in having pale legs with a dark brown hind tibia, only the fore femur armed with ventral spines and a distinctly curved gonostylus. Males of P. mahyoubi have a nearly straight gonostylus, dark brown legs, and all femora have ventral spines.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2017, Predatory midges of the tribes Palpomyiini and Sphaeromiini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 1-30 in European Journal of Taxonomy 318 on pages 18-20, DOI: 10.5852/ejt.2017.318, http://zenodo.org/record/382745
Brachypogon (Brachypogon) afifi Boorman & Harten 2002
Brachypogon (B.) afifi Boorman & Harten, 2002 Brachypogon afifi Boorman & Harten, 2002: 452 (male, Yemen). Diagnosis. Gonostylus in male genitalia short, stout and slightly curved, parameres separate. Female unknown (Boorman & Harten 2002). Distribution. Yemen.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2016, Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 551-572 in Zootaxa 4079 (5) on page 558, DOI: 10.11646/zootaxa.4079.5.3, http://zenodo.org/record/105087
Stilobezzia harurii Boorman & van Harten 2002
Stilobezzia harurii Boorman & van Harten, 2002 Stilobezzia harurii Boorman & van Harten, 2002: 454 (male, Yemen); Szadziewski et al. 2011: 649 (male, female, UAE). Diagnosis. Wings grey, with dark patch over the vein r-m. Scutellum yellow, with six marginal setae; palpus dark brown; legs with brown rings; abdominal tergites with darker margins (Boorman & van Harten 2002, Szadziewski et al. 2011). Distribution. UAE, Yemen.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2016, Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 551-572 in Zootaxa 4079 (5) on page 569, DOI: 10.11646/zootaxa.4079.5.3, http://zenodo.org/record/105087
Serromyia arabica Szadziewski, Gwizdalska-Kentzer & Gilka 2011
Serromyia arabica Szadziewski, Gwizdalska-Kentzer & Giłka, 2011 Serromyia arabica Szadziewski, Gwizdalska-Kentzer & Giłka, 2011: 648 (female, UAE). Diagnosis. The species is characteristic in having spotted body; female with two very large ovoid seminal capsules, claw of hind leg 1.8 times longer than fifth tarsomere. Male unknown (Szadziewski et al. 2011). Distribution. United Arab Emirates.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2016, Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 551-572 in Zootaxa 4079 (5) on page 564, DOI: 10.11646/zootaxa.4079.5.3, http://zenodo.org/record/105087
Brachypogon (Brachypogon) yarimii Boorman & Harten 2002
Brachypogon (B.) yarimii Boorman & Harten, 2002 Brachypogon yarimii Boorman & Harten, 2002: 453 (male, female, Oman, Yemen). Diagnosis. Males of the species are characteristic in having long and slender gonostylus, and Y-shaped aedeagus; female with two subequal pear-shaped seminal capsules (Boorman & Harten 2002). Distribution. Oman, Yemen.Published as part of Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek, 2016, Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species, pp. 551-572 in Zootaxa 4079 (5) on page 559, DOI: 10.11646/zootaxa.4079.5.3, http://zenodo.org/record/105087
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